Itemirus Temporal range: Turonian, | |
---|---|
Reconstruction | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Dromaeosauridae |
Clade: | † Eudromaeosauria |
Subfamily: | † Dromaeosaurinae |
Genus: | † Itemirus Kurzanov, 1976 |
Species: | †I. medullaris |
Binomial name | |
†Itemirus medullaris Kurzanov, 1976 | |
Itemirus is a genus of dromaeosaurid theropod dinosaur from the Turonian age of the Late Cretaceous period of Uzbekistan.
Itemirus is known from a single small, damaged fossil braincase or neurocranium, in 1958 found near the village of Itemir at the Dzharakuduk escarpment in layers of the Bissekty Formation. This holotype has accession number PIN 327/699. The type species, Itemirus medullaris, was named and described by Sergei Kurzanov in 1976. The generic name refers to Itemir. The specific name refers to the medulla oblongata, the part of the brain encased by the partial braincase. [1]
Kurzanov noted anatomical similarities to the Tyrannosauridae and the Dromaeosauridae; he assigned Itemirus to a separate Itemiridae. In 2004, Thomas Holtz suggested it was a member of the Tyrannosauroidea. [2] Nicholas Longrich and Philip J. Currie in 2009 included Itemirus in a cladistic analysis of internal dromaeosaurid relationships and found it to be a velociraptorine. [3] In 2014, during a study assigning more material to Itemirus, it was found that the genus could be placed in Dromaeosaurinae in a phylogeny. [4]
Saurornitholestes is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States.
Bambiraptor is a Late Cretaceous, 72-million-year-old, bird-like dromaeosaurid theropod dinosaur described by scientists at the University of Kansas, Yale University, and the University of New Orleans.
Avimimus, meaning "bird mimic", is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 to 70 million years ago.
Adasaurus is a genus of dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous period about 70 million years ago. The genus is known from two partial specimens found in the Nemegt Formation of Mongolia that were partially described in 1983 by the paleontologist Rinchen Barsbold.
Caenagnathasia is a small caenagnathid oviraptorosaurian theropod from the Late Cretaceous of Uzbekistan.
The Bayan Shireh Formation is a geological formation in Mongolia, that dates to the Cretaceous period. It was first described and established by Vasiliev et al. 1959.
Bissektipelta is a genus of ankylosaurine thyreophoran dinosaurs that lived in Asia during the Late Cretaceous in what is now the Bissekty Formation of Uzbekistan. Bissektipelta is a monospecific genus, containing only the type species B. archibaldi.
Pectinodon is a genus of troodontid theropod dinosaurs from the end of the Maastrichtian age of the Late Cretaceous period (66 mya). It currently contains a single valid species, Pectinodon bakkeri, known only from teeth.
Hulsanpes is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. Hulsanpes represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae.
Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. Definitive fossils attributed to the subfamily have only been found in the Late Cretaceous deposits of Asia, with Kansaignathus as the basalmost member. While numerous taxa from North America are previously assigned to the velociraptorines, they are now reclassified within different lineages of Eudromaeosauria. Several teeth that may belong to indeterminate velociraptorines have also been discovered in Germany, dating to the Kimmeridgian stage of the Late Jurassic.
Austroraptor is a genus of dromaeosaurid theropod dinosaur that lived during the Campanian and Maastrichtian ages of the Late Cretaceous period in what is now Argentina.
Hesperonychus is a genus of small paravian theropod dinosaur. It may be a dromaeosaurid or an avialan. There is one described species, Hesperonychus elizabethae. The type species was named in honor of Dr. Elizabeth Nicholls of the Royal Tyrrell Museum of Palaeontology who collected it as a student in 1982. It is known from fossils recovered from the Dinosaur Park Formation and possibly from the uppermost strata of the Oldman Formation of Alberta, dating to the Campanian stage of the Late Cretaceous around 75 million years ago.
Saurornitholestinae is a subfamily of the theropod group Dromaeosauridae. The saurornitholestines currently include monotypic genera Atrociraptor marshalli, Saurornitholestes langstoni, and possibly Bambiraptor feinbergi and Acheroraptor temertyorum. All are medium-sized dromaeosaurs from the Late Cretaceous of western North America. The group was originally recognized by Longrich and Currie as the sister taxon to a clade formed by the Dromaeosaurinae and Velociraptorinae. However, not all phylogenetic analyses recover this group and/or with the same proposed genera.
The Bissekty Formation is a geologic formation and Lagerstätte which crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period. Laid down in the mid to late Turonian, it is dated to about 92 to 90 Ma.
Eudromaeosauria is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were small to large-sized predators that flourished during the Cretaceous Period. Eudromaeosaur fossils are known almost exclusively from the northern hemisphere.
Acheroraptor is an extinct genus of dromaeosaurid theropod dinosaur known from the latest Maastrichtian Hell Creek Formation of Montana, United States. It contains a single species, Acheroraptor temertyorum. A. temertyorum is one of the two geologically youngest known species of dromaeosaurids, the other being Dakotaraptor steini, which is also known from Hell Creek. A basal cousin of Velociraptor, Acheroraptor is known from upper and lower jaw material.
This timeline of dromaeosaurid research is a chronological listing of events in the history of paleontology focused on the dromaeosaurids, a group of sickle-clawed, bird-like theropod dinosaurs including animals like Velociraptor. Since the Native Americans of Montana used the sediments of the Cloverly Formation to produce pigments, they may have encountered remains of the dromaeosaurid Deinonychus hundreds of years before these fossils came to the attention of formally trained scientists.
Timurlengia is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is Timurlengia euotica.
Ulughbegsaurus is a dubious genus of theropod dinosaur from the Late Cretaceous aged Bissekty Formation, Uzbekistan. The type species is Ulughbegsaurus uzbekistanensis.
Dzharaonyx is a genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Bissekty Formation of Uzbekistan. The type species is Dzharaonyx eski; eski being an Uzbek word for "old".