Hadrosauroids Temporal range: Early-Late Cretaceous, | |
---|---|
Probactrosaurus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ornithopoda |
Clade: | † Hadrosauriformes |
Superfamily: | † Hadrosauroidea Cope, 1869 |
Type species | |
† Hadrosaurus foulkii Leidy, 1858 | |
Subgroups | |
| |
Synonyms | |
Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or Hadrosauridae, and all dinosaurs more closely related to them than to Iguanodon . Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian Probactrosaurus and Altirhinus , have traditionally been included in a paraphyletic (unnatural grouping) "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only Iguanodon and perhaps its closest relatives.
The cladogram below follows an analysis by Andrew McDonald, 2012, and shows the position of Hadrosauroidea within Styracosterna: [1]
The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012): [2]
Cladogram after Prieto-Marquez and Norell (2010): [3]
Hadrosauroidea | |
A phylogenetic analysis performed by Ramírez-Velasco et al. (2012) found a large polytomy between all hadrosauroids that are more derived than Probactrosaurus but less derived than Hadrosauridae. The exclusion of Claosaurus , Jeyawati , Levnesovia , Nanyangosaurus , Shuangmiaosaurus and Telmatosaurus from the analysis resulted in a more resolved topology: [4]
Hadrosauroidea | |||||||||||||
Hadrosaurus is a genus of hadrosaurid ornithopod dinosaurs that lived in North America during the Late Cretaceous Period in what is now the Woodbury Formation in New Jersey about 78-80 Ma. The holotype specimen was found in fluvial marine sedimentation, meaning that the corpse of the animal was transported by a river and washed out to sea.
Ornithopoda is a clade of ornithischian dinosaurs, called ornithopods. They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively small-sized, while advanced members of the subgroup Iguanodontia became quadrupedal and developed large body size. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids, before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known worldwide.
Eolambia is a genus of herbivorous hadrosauroid dinosaur from the early Late Cretaceous of the United States. It contains a single species, E. caroljonesa, named by paleontologist James Kirkland in 1998. The type specimen of Eolambia was discovered by Carole and Ramal Jones in 1993; the species name honors Carole. Since then, hundreds of bones have been discovered from both adults and juveniles, representing nearly every element of the skeleton. All of the specimens have thus far been found in Emery County, Utah, in a layer of rock known as the Mussentuchit Member of the Cedar Mountain Formation.
Iguanodontidae is a family of iguanodontians belonging to Styracosterna, a derived clade within Ankylopollexia.
Barsboldia is a genus of large hadrosaurid dinosaur from the early Maastrichtian Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, partial pelvis, and some ribs.
Anasazisaurus is a genus of saurolophine hadrosaurid ("duckbill") ornithopod dinosaur that lived about 74 million years ago, in the Late Cretaceous Period. It was found in the Farmington Member of the Kirtland Formation, in the San Juan Basin of New Mexico, United States. Only a partial skull has been found to date. It was first described as a specimen of Kritosaurus by Jack Horner, and has been intertwined with Kritosaurus since its description. It is known for its short nasal crest, which stuck out above and between its eyes for a short distance.
Fukuisaurus is a genus of herbivorous ornithopod dinosaur that lived during the Early Cretaceous in what is now Japan. The type species is F. tetoriensis, which was named and described in 2003.
Gilmoreosaurus is the name given to a genus of dinosaur from the Cretaceous of Asia. The type species is Gilmoreosaurus mongoliensis. It is believed to be a hadrosaur or iguanodont from the Iren Dabasu Formation of Inner Mongolia, dating to 96 Ma ago. Additional specimens have been described as distinct species, including G. atavus from the Khodzhakul Formation of Uzbekistan and G. arkhangelskyi from the Bissekty Formation. However, these are based on very fragmentary remains, and their classification is dubious. An additional species, G. kysylkumense is sometimes included, though it has also been referred to the related genus Bactrosaurus.
Probactrosaurus is an early herbivorous hadrosauroid iguanodont dinosaur. It lived in China during the Early Cretaceous period.
Lambeosaurinae is an extinct group of crested hadrosaurid dinosaurs.
Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadrosaurinae is based on the genus Hadrosaurus which was found in more recent studies to be more primitive than either lambeosaurines or other traditional "hadrosaurines", like Edmontosaurus and Saurolophus. As a result of this, the name Hadrosaurinae was dropped or restricted to Hadrosaurus alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae. Recent phylogenetic work by Hai Xing indicates that Hadrosaurus is placed within the monophyletic group containing all non-lambeosaurine hadrosaurids. Under this view, the traditional Hadrosaurinae is resurrected, with the Hadrosauridae being divided into two clades: Hadrosaurinae and Lambeosaurinae.
Tethyshadros is a genus of hadrosauroid dinosaur from the Calcare di Aurisina of Trieste, Italy. The type and only species is T. insularis.
Willinakaqe is a dubious genus of saurolophine hadrosaurid dinosaur described based on fossils from the late Cretaceous of the Río Negro Province of southern Argentina.
Huehuecanauhtlus is a genus of hadrosauroid dinosaur known from the Late Cretaceous of Michoacán, western Mexico. It contains a single species, Huehuecanauhtlus tiquichensis.
Latirhinus is an extinct genus of lambeosaurine hadrosaurid dinosaur from the Late Cretaceous of Mexico. The type species, Latirhinus uitstlani, was named in 2012 on the basis of a partial skeleton from the Campanian-age Cerro del Pueblo Formation. The specific name uitstlani means "southern" in the Náhuatl language of Mexico, a reference to the species' southern occurrence in the Cretaceous landmass Laramidia.
The year 2012 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2012 in paleontology included various significant developments regarding archosaurs.
Gongpoquansaurus is an extinct genus of basal hadrosauroid dinosaur that was not formally named until 2014, while the name was a nomen nudum for many years previously. It is known from IVPP V.11333, a partial skull and postcranial skeleton. It was collected in 1992 at locality IVPP 9208–21, from the Albian Zhonggou Formation, in Mazongshan, Gansu Province, China. The specimen was first described and named by Lü Junchang in 1997 as the third species of Probactrosaurus, Probactrosaurus mazongshanensis. Following its description, several studies found it to be less derived than the type species of Probactrosaurus in relation to Hadrosauridae. Therefore, "Gongpoquansaurus" had been suggested, yet informally, as a replacement generic name. In 2014, the species was formally redescribed, and the describers erected Gongpoquansaurus.
This timeline of hadrosaur research is a chronological listing of events in the history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosaurs. Scientific research on hadrosaurs began in the 1850s, when Joseph Leidy described the genera Thespesius and Trachodon based on scrappy fossils discovered in the western United States. Just two years later he published a description of the much better-preserved remains of an animal from New Jersey that he named Hadrosaurus.
Hadrosauromorpha is a clade of iguanodontian ornithopods, defined in 2014 by David B. Norman to divide Hadrosauroidea into the basal taxa with compressed manual bones and a pollex, and the derived taxa that lack them. The clade is defined as all the taxa closer to Edmontosaurus regalis than Probactrosaurus gobiensis. This results in different taxon inclusion depending on the analysis.