Gasparinisaura Temporal range: Early Campanian, | |
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Skeleton mount, Copiapó, Chile | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ornithopoda |
Clade: | † Elasmaria |
Genus: | † Gasparinisaura Coria & Salgado, 1996 |
Species: | †G. cincosaltensis |
Binomial name | |
†Gasparinisaura cincosaltensis Coria & Salgado, 1996 | |
Gasparinisaura (meaning "Gasparini's lizard") is a genus of herbivorous ornithopod dinosaur from the Late Cretaceous.
The first fossils of Gasparinisaura were found in 1992 near Cinco Saltos in Río Negro Province, Argentina. The type species, Gasparinisaura cincosaltensis, was named and described in 1996 by Rodolfo Coria and Leonardo Salgado. The generic name honors Argentine palaeontologist Zulma Brandoni de Gasparini. The specific name refers to Cinco Saltos. [1]
The holotype, MUCPv-208, was uncovered in 1992 in a layer of the Anacleto Formation, dating from the early Campanian, about 83 million years old. It consists of a partial skeleton with skull, lacking much of the vertebral column. The paratype is MUCPv-212, a tail with lower hindlimb elements. [1] In 1997, three additional specimens were described: MUCPv 213, a partial skeleton with skull; MCSPv 111, a postcranial skeleton; and MCSPv 112, a skull with hindlimbs and pelvis. The three later specimens represent juvenile individuals. [2] In 2008, gastroliths present with all three juvenile specimens were described. [3]
Gasparinisaura was a small bipedal herbivore. In 2010, Gregory S. Paul estimated the length at 1.7 metres (5.6 ft) and the weight at 13 kilograms (29 lb). [4]
Gasparinisaura had a rounded head that was moderately elongated. The eye sockets were very large and placed in a high position. The rather long back of the head was pendant. The jugal had a front branch that was thin and wedged between the maxilla and the lacrimal; the back of the jugal was high and broad. The quadratojugal had a long ascending process touching the squamosal, a basal trait. The jaw joints were low. There were about twelve, rather large, maxillary or dentary teeth in each upper or lower jaw. The forelimbs were lightly built. The ilium was short and low, the pubis thin. The hindlimbs were relatively long and powerful, with a short thighbone and a long lower leg. The foot was long and the first metatarsal was reduced to a splint, a derived trait. The tail featured low plate-like triangular chevrons that were expanded towards the back, which is unique in the Euornithopoda. [1]
The gastroliths discovered formed clusters of up to hundred-forty rounded and polished stones in the abdomen with an average stone diameter of about eight millimetres. Representing about 0.3% of total body weight, they were voluminous enough to function as a gastric mill, a function sometimes denied for sauropod gastroliths because of their insufficient relative mass, being an order of magnitude lower. [3]
Gasparinisaurus was originally considered by the describers to have been a basal hypsilophodont, the most basal member of the clade ornithopoda. [1] A phylogenetic analysis included in the description of the Antarctic ornithopod Morrosaurus shows that the latter and Gasparinisaura are part of Elasmaria, a clade of hypsilophodonts known from South America, Antarctica and maybe Australia. Gasparinisaura would be the basalmost member of this group. [5]
Cladogram based in the phylogenetic analysis of Rozadilla et al., 2016:
Ornithopoda is a clade of ornithischian dinosaurs, called ornithopods. They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively small-sized, while advanced members of the subgroup Iguanodontia became quadrupedal and developed large body size. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids, before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known worldwide.
Hypsilophodontidae is a traditionally used family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, and spanning from the Middle Jurassic until the Late Cretaceous. This inclusive status was supported by some phylogenetic analyses from the 1990s and mid 2000s, although there have also been many finding that the family is an unnatural grouping which should only include the type genus, Hypsilophodon, with the other genera being within clades like Thescelosauridae and Elasmaria. A 2014 analysis by Norman recovered a grouping of Hypsilophodon, Rhabdodontidae and Tenontosaurus, which he referred to as Hypsilophodontia. All other analyses from around the same time have instead found these latter taxa to be within Iguanodontia.
Thescelosaurus was a genus of neornithischian dinosaur that appeared at the very end of the Late Cretaceous period in North America. It was a member of the last dinosaurian fauna before the Cretaceous–Paleogene extinction event around 66 million years ago. The preservation and completeness of many of its specimens indicate that it may have preferred to live near streams.
Notohypsilophodon is a genus of ornithopod dinosaur from the Late Cretaceous of Argentina. It was described as the only "hypsilophodont" known from South America, although this assessment is not universally supported, and Gasparinisaura is now believed to have been a basal euornithopod as well.
Parksosaurus is a genus of neornithischian dinosaur from the early Maastrichtian-age Upper Cretaceous Horseshoe Canyon Formation of Alberta, Canada. It is based on most of a partially articulated skeleton and partial skull, showing it to have been a small, bipedal, herbivorous dinosaur. It is one of the few described non-hadrosaurid ornithopods from the end of the Cretaceous in North America, existing around 70 million years ago.
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Loncosaurus is an extinct genus of ornithopod dinosaur from the Upper Cretaceous of Provincia de Santa Cruz, Argentina. The type species is Loncosaurus argentinus, described by the famous Argentinian paleontologist Florentino Ameghino, but is considered a dubious name. Details on this animal are often contradictory, befitting a genus that was long confused for a theropod.
Nanosaurus is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-western United States. The type and only species, Nanosaurus agilis, was described and named by Othniel Charles Marsh in 1877. The taxon has a complicated taxonomic history, largely the work of Marsh and Peter M. Galton, involving the genera Laosaurus, Hallopus, Drinker, Othnielia, and Othnielosaurus, the latter three now being considered to be synonyms of Nanosaurus. It had historically been classified as a hypsilophodont or fabrosaur, types of generalized small bipedal herbivore, but more recent research has abandoned these groupings as paraphyletic and Nanosaurus is today considered a basal member of Neornithischia.
Antarctopelta is a genus of ankylosaurian dinosaur, a group of large, quadrupedal herbivores, that lived during the Maastrichtian stage of the Late Cretaceous period on what is now James Ross Island, Antarctica. Antarctopelta is the only known ankylosaur from Antarctica and a member of Parankylosauria. The only described specimen was found in 1986, the first dinosaur to be found on the continent, by Argentine geologists Eduardo Olivero and Robert Scasso. The fossils were later described in 2006 by paleontologists Leonardo Salgado and Zulma Gasparini, who named the type species A. oliveroi after Olivero.
The Anacleto Formation is a geologic formation with outcrops in the Argentine Patagonian provinces of Mendoza, Río Negro, and Neuquén. It is the youngest formation within the Neuquén Group and belongs to the Río Colorado Subgroup. Formerly that subgroup was treated as a formation, and the Anacleto Formation was known as the Anacleto Member.
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Trinisaura is a genus of ornithopod dinosaur that lived during the late Campanian stage of the Upper Cretaceous, around 73 to 72 million years ago in what is now James Ross Island off the coast of northern Antarctica near Patagonia. It is known from a single, incomplete postcranial skeleton that includes several vertebrae, a partial pelvis, and nearly complete right hindlimb. The fossils were collected in 2008 by paleontologists Juan Moly and Rodolfo Coria from the sandstone of the Snow Hill Island Formation. It remained undescribed in the collections of the Museo de La Plata until its description by Coria and colleagues in 2013, being the basis of the novel genus and species Trinisaura santamartaensis. The genus name is to commemorate the efforts of Argentine geologist Trinidad "Trini" Diaz and the Latin root -sauros, meaning "lizard". The species name is after Santa Marta Cove, where the fossils were collected.
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