Dryosauridae Temporal range: Middle Jurassic – Early Cretaceous, | |
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Dysalotosaurus skeletal mount in Berlin | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ornithopoda |
Clade: | † Iguanodontia |
Clade: | † Dryomorpha |
Superfamily: | † Dryosauroidea Milner & Norman, 1984 |
Family: | † Dryosauridae Milner & Norman, 1984 |
Subgroups | |
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Dryosauridae was a family of primitive iguanodonts, first proposed by Milner & Norman in 1984. They are known from Middle Jurassic to Early Cretaceous rocks of Africa, Europe, and North America. [1] [2] [3] [4]
Until recently, many dryosaurids have been regarded as dubious ( Callovosaurus and Kangnasaurus ) or as species of the type member, Dryosaurus ( Dysalotosaurus , Elrhazosaurus and Valdosaurus ). However, more recent studies redescribe these genera as valid. [3] [4] [5] [6] [7] Under the Phylocode, Madzia et al. (2021) formally defined Dryosauridae as "the largest clade containing Dryosaurus altus but not Iguanodon bernissartensis ." [8] The cladogram below follows their chosen reference phylogeny, taken from the description of Orthomerus dolloi . [9]
Iguanodon, named in 1825, is a genus of iguanodontian dinosaur. While many species found worldwide have been classified in the genus Iguanodon, dating from the Late Jurassic to Early Cretaceous, taxonomic revision in the early 21st century has defined Iguanodon to be based on one well-substantiated species: I. bernissartensis, which lived during the Barremian to early Aptian ages of the Early Cretaceous in Belgium, Germany, England, and Spain, between about 126 and 122 million years ago. Iguanodon was a large, bulky herbivore, measuring up to 9–11 metres (30–36 ft) in length and 4.5 metric tons in body mass. Distinctive features include large thumb spikes, which were possibly used for defense against predators, combined with long prehensile fifth fingers able to forage for food.
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.
Ornithopoda is a clade of ornithischian dinosaurs, called ornithopods. They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively small-sized, while advanced members of the subgroup Iguanodontia became quadrupedal and developed large body size. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids, before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known worldwide.
Valdosaurus is a genus of bipedal herbivorous iguanodont ornithopod dinosaur found on the Isle of Wight and elsewhere in England, Spain and possibly also Romania. It lived during the Early Cretaceous.
Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard'.
Hypsilophodontidae is a traditionally used family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, and spanning from the Middle Jurassic until the Late Cretaceous. This inclusive status was supported by some phylogenetic analyses from the 1990s and mid 2000s, although there have also been many finding that the family is an unnatural grouping which should only include the type genus, Hypsilophodon, with the other genera being within clades like Thescelosauridae and Elasmaria. A 2014 analysis by Norman recovered a grouping of Hypsilophodon, Rhabdodontidae and Tenontosaurus, which he referred to as Hypsilophodontia. All other analyses from around the same time have instead found these latter taxa to be within Iguanodontia.
Callovosaurus is a genus of iguanodontian dinosaur known from most of a left thigh bone discovered in Middle Jurassic-age rocks of England. At times, it has been considered dubious or a valid genus of basal iguanodontian, perhaps a dryosaurid.
Cumnoria is a genus of herbivorous iguanodontian dinosaur. It was a basal iguanodontian that lived during the Late Jurassic period in what is now Oxfordshire, United Kingdom.
Kangnasaurus is a genus of iguanodontian ornithopod dinosaur found in supposedly Early Cretaceous rocks of South Africa. It is known from a tooth and possibly some postcranial remains found in the early-Aptian Kalahari Deposits Formation. It was probably similar to Dryosaurus.
Orthomerus is a genus of dubious hadrosaurid dinosaur from the Late Cretaceous of the Netherlands. It is today an obscure genus, but in the past was conflated with the much better known Telmatosaurus.
Theiophytalia is a genus of herbivorous iguanodontian dinosaur from the lower Cretaceous period of Colorado, USA. It contains a single species, T. kerri.
The palpebral bone is a small dermal bone found in the region of the eye socket in a variety of animals, including crocodilians and ornithischian dinosaurs. It is also known as the adlacrimal or supraorbital, although the latter term may not be confused with the supraorbital in osteichthyan fishes. In ornithischians, the palpebral can form a prong that projects from the front upper corner of the orbit. It is large in heterodontosaurids, basal ornithopods such as Thescelosaurus and Dryosaurus, and basal ceratopsians such as Archaeoceratops; in these animals, the prong is elongate and would have stuck out and over the eye like a bony eyebrow. As paleoartist Gregory S. Paul has noted, elongate palpebrals would have given their owners fierce-looking "eagle eyes". In such cases, the expanded palpebral may have functioned to shade the eye.
Dakotadon is a genus of iguanodont dinosaur from the Barremian-age Lower Cretaceous Lakota Formation of South Dakota, USA, known from a partial skull. It was first described in 1989 as Iguanodon lakotaensis, by David B. Weishampel and Philip R. Bjork. Its assignment has been controversial. Some researchers suggest that "I." lakotaensis was more basal than I. bernissartensis, and related to Theiophytalia, but David Norman has suggested that it was a synonym of I. bernissartensis. Gregory S. Paul, working on a revision of iguanodont species, gave "I." lakotaensis its own genus (Dakotadon) in 2008. He measured its length at 6 metres (20 ft) and body mass at 1 metric ton.
Thescelosauridae is a clade of neornithischians from the Cretaceous of East Asia and North America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was used by Brown and colleagues as the group uniting Thescelosaurus and Orodromeus, based on their phylogenetic results. During a phylogenetic revision of neornithischians by Clint Boyd in 2015, the authorship of Thescelosauridae was given to Brown and colleagues, which meant that the similar name Parksosauridae, informally defined in 2002 by Buchholz, would have had priority over Thescelosauridae. The two clades had slightly different definitions, with Parksosauridae referring to all animals closer to Parksosaurus than Hypsilophodon, but they contained the same taxa so Boyd used Parksosauridae under the assumption it had priority. However, in formalizing the clade following the regulations of the PhyloCode, Madzia, Boyd, and colleagues identified in 2021 that Sternberg was the proper authority for Thescelosauridae, giving it priority over Parksosauridae. As well, they gave Thescelosauridae the definition of the largest clade containing Thescelosaurus neglectus but not Iguanodon bernissartensis, as long as Hypsilophodon foxii was not in the group, modifying previous definitions for Thescelosauridae in order to maintain its modern use, so that the clade was not applied if Thescelosaurus fell within Hypsilophodontidae, a family that has not been recently used but may be revived if the systematic position of Hypsilophodon was solidified at some point in the future. Madzia et al. identified the analysis of Madzia et al. in 2018 as the reference analysis for the name Thescelosauridae, an analysis based on a revised version of the 2015 Boyd analysis.
Owenodon is a genus of iguanodontian dinosaur known from a partial lower jaw discovered in Early Cretaceous-age rocks of Durlston Bay, Dorset, United Kingdom, and possibly also Romania and Spain. The specimen, NHM R2998, comes from the Purbeck Limestone, dating to the middle Berriasian stage. It was first described by Richard Owen, who in 1874 assigned it to Iguanodon as the type specimen of the new species I. hoggii, the specific name honouring naturalist A.J. Hogg who had originally collected the fossil in 1860. The bone was damaged during initial preparation but was freed from the surrounding rock matrix by an acid bath between 1975 and 1977. David Norman and Paul Barrett subsequently transferred the species to Camptosaurus in 2002, but this was challenged, and in 2009 Peter Galton assigned the species to the new genus Owenodon, meaning "Owen's tooth", named after Sir Richard Owen. Galton interpreted the genus as an iguanodontoid more derived than Camptosaurus but less derived than Lurdusaurus.
The Kalahari Deposits is an Early Cretaceous (Aptian) geologic formation in South Africa. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation. The depositional environment is described as a crater lake where poorly lithified, concretionary conglomerate and volcaniclastic, intraclastic, calcareous mudstone were deposited under quiet subaqueous conditions, probably a "crater-fill succession above an olivine-melilitie intrusion".
Hypselospinus is a genus of iguanodontian dinosaur which was first described as a species of Iguanodon by Richard Lydekker in 1889, the specific name honouring William Henry Fitton.
Hadrosauromorpha is a clade of iguanodontian ornithopods, defined in 2014 by David B. Norman to divide Hadrosauroidea into the basal taxa with compressed manual bones and a pollex, and the derived taxa that lack them. The clade is defined as all the taxa closer to Edmontosaurus regalis than Probactrosaurus gobiensis. This results in different taxon inclusion depending on the analysis.