Eousdryosaurus Temporal range: Late Jurassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ornithopoda |
Family: | † Dryosauridae |
Genus: | † Eousdryosaurus Escaso et al., 2014 |
Species: | †E. nanohallucis |
Binomial name | |
†Eousdryosaurus nanohallucis Escaso et al., 2014 | |
Eousdryosaurus ("eastern Dryosaurus ") is a genus of basal iguanodontian dinosaur known from a partial skeleton discovered in Upper Jurassic rocks in western Portugal. The type, and only species, is Eousdryosaurus nanohallucis, named and described in 2014.
Eousdryosaurus is based on SHN(JJS)-170, kept in the Sociedade de História Natural in Torres Vedras, Portugal. The articulated and well-preserved specimen includes a hip vertebra and eight vertebrae from the proximal part of the tail, their chevrons, the left ilium, the entire left leg, and the right thigh bone (femur). These bones were discovered in sandstone of the late Kimmeridgian-age (late Jurassic, approximately 152 million years old) Praia da Amoreira-Porto Novo Member of the Lourinhã Formation in Porto das Barcas, Lourinhã, Portugal. The genus name is a reference to Eousdryosaurus being a relative of Dryosaurus on the east side of the Atlantic Ocean. The type and only species is E. nanohallucis, meaning "small hallux", in reference to the small size of that toe. [1] Eousdryosaurus was named in 2014 by Fernando Escaso and colleagues, [1] after having been described briefly in 2000. [2]
The only known individual of Eousdryosaurus was a small dryosaurid, estimated at 1.6 metres (5.2 ft) in length, comparable to immature individuals of Dryosaurus and Dysalotosaurus . The right femur measures 188.5 millimetres (7.42 in) long and the left shin bone (tibia) measures 200 millimetres (7.9 in) long. [1]
It is differentiated from other dryosaurids by various details of the vertebrae, hip, and hind limb. The foot is unique among ornithopods in that the first digit (the hallux or big toe) includes a single phalanx bone; most basal ornithopods had two phalanges, and most derived ornithopods, including hadrosaurs ("duckbills"), lost this digit. [1]
Escaso et al. performed a phylogenetic analysis and found Eousdryosaurus to be a basal dryosaurid within the Dryomorpha, a type of small bipedal herbivore near the base of Iguanodontia. [1]
On the other hand, Dieudonné et al. (2020) found Eousdryosaurus to be a sister taxon to the Elasmaria. [3]
In their 2024 description of the contemporary Hesperonyx , Rotatori et al. found support for a dryosaurid position of Eousdryosaurus in both their parsimony and Bayesian analyses, contrasting with the suggestions of Dieudonné et al. (2020). Their results are displayed in the cladogram below: [4]
Iguanodontia |
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Ornithopoda is a clade of ornithischian dinosaurs, called ornithopods. They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively small-sized, while advanced members of the subgroup Iguanodontia became quadrupedal and developed large body size. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids, before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known worldwide.
Valdosaurus is a genus of bipedal herbivorous iguanodont ornithopod dinosaur found on the Isle of Wight and elsewhere in England, Spain and possibly also Romania. It lived during the Early Cretaceous.
Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard'.
Hypsilophodontidae is a traditionally used family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, and spanning from the Middle Jurassic until the Late Cretaceous. This inclusive status was supported by some phylogenetic analyses from the 1990s and mid 2000s, although there have also been many finding that the family is an unnatural grouping which should only include the type genus, Hypsilophodon, with the other genera being within clades like Thescelosauridae and Elasmaria. A 2014 analysis by Norman recovered a grouping of Hypsilophodon, Rhabdodontidae and Tenontosaurus, which he referred to as Hypsilophodontia. All other analyses from around the same time have instead found these latter taxa to be within Iguanodontia.
Callovosaurus is a genus of iguanodontian dinosaur known from most of a left thigh bone discovered in Middle Jurassic-age rocks of England. At times, it has been considered dubious or a valid genus of basal iguanodontian, perhaps a dryosaurid.
Alocodon is a genus of ornithischian dinosaur known from multiple teeth from the Middle or Late Jurassic Cabaços Formation of Portugal, and also the Forest Marble and Chipping Norton Formations of England. A single species is known, A. kuehnei.
Trimucrodon is a genus of ornithischian dinosaur from the Late Jurassic Lourinhã Formation of Portugal. The type, and currently only, species is T. cuneatus.
Dysalotosaurus is a genus of herbivorous iguanodontian dinosaur. It was a dryosaurid iguanodontian, and its fossils have been found in late Kimmeridgian-age rocks of the Tendaguru Formation of Lindi Region in Tanzania. The type and only species of the genus is D. lettowvorbecki. This species was named by Hans Virchow in 1919 in honor of the Imperial German Army Officer, Paul von Lettow-Vorbeck. For much of the 20th century the species was referred to the related and approximately contemporary genus Dryosaurus, but newer studies reject this synonymy.
Dracopelta is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic in what is now the Lourinhã Formation. The type and only species is Dracopelta zbyszewskii, which is represented by a partial skeleton including unpublished material.
Kangnasaurus is a genus of iguanodontian ornithopod dinosaur found in supposedly Early Cretaceous rocks of South Africa. It is known from a tooth and possibly some postcranial remains found in the early-Aptian Kalahari Deposits Formation. It was probably similar to Dryosaurus.
Draconyx is a genus of dinosaur from the Late Jurassic. It was an ornithopod which lived in what is now Portugal and was a herbivore. It was found in the Lourinhã Formation in 1991, and described by Octávio Mateus and Miguel Telles Antunes in 2001.
Dinheirosaurus is a genus of diplodocid sauropod dinosaur that is known from fossils uncovered in modern-day Portugal. It may represent a species of Supersaurus. The only species is Dinheirosaurus lourinhanensis, first described by José Bonaparte and Octávio Mateus in 1999 for vertebrae and some other material from the Lourinhã Formation. Although the precise age of the formation is not known, it can be dated around the early Tithonian of the Late Jurassic.
Dryosauridae was a family of primitive iguanodonts, first proposed by Milner & Norman in 1984. They are known from Middle Jurassic to Early Cretaceous rocks of Africa, Europe, and North America.
The Lourinhã Formation is a fossil rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the earliest Cretaceous (Berriasian). It is notable for containing a fauna especially similar to that of the Morrison Formation in the United States and a lesser extent to the Tendaguru Formation in Tanzania. There are also similarities to the nearby Villar del Arzobispo Formation and Alcobaça Formation. The stratigraphy of the formation and the basin in general is complex and controversial, with the constituent member beds belonging to the formation varying between different authors.
Ankylopollexia is an extinct clade of ornithischian dinosaurs that lived from the Late Jurassic to the Late Cretaceous. It is a derived clade of iguanodontian ornithopods and contains the subgroup Styracosterna. The name stems from the Greek word, “ankylos”, mistakenly taken to mean stiff, fused, and the Latin word, “pollex”, meaning thumb. Originally described in 1986 by Sereno, a most likely synapomorphic feature of a conical thumb spine defines the clade.
Laquintasaura is a genus of Venezuelan ornithischian dinosaur containing only the type species Laquintasaura venezuelae. The species was the first dinosaur to have been identified from Venezuela. It is known from extensive remains, all from a single bonebed locality which has been sampled for specimen blocks over the course of several expeditions, largely led by Marcelo R Sánchez-Villagra. A small animal, it is known for its distinct dental anatomy and for being one of the earliest and most primitive ornithischians in the fossil record. Taxonomic uncertainty has led to conflicting hypotheses that it is either at the base of Ornithischia or at the base of the subgroup Thyreophora. In either model, its dating to around 200 million years ago at the start of the Jurassic, existence in equatorial latitudes, and primitive nature make it a key insight into early ornithischian evolution. It is thought that Laquintasaura would have lived in groups and had a possible omnivorous diet, living on a seasonal alluvial plain and being preyed upon by the contemporary Tachiraptor.
Lusovenator is a genus of carcharodontosaurian theropod dinosaur, from the Late Jurassic (Kimmeridgian) Praia de Amoreira Porto-Novo Member and the Late Jurassic-Early Cretaceous (Tithonian-Berriasian) Assenta Member of the Lourinhã Formation in present-day Portugal. It includes one species, Lusovenator santosi.
Hesperonyx is an extinct genus of dryomorphan ornithopod dinosaur from the Late Jurassic Lourinhã Formation of Portugal. The genus contains a single species, Hesperonyx martinhotomasorum, known from bones of the fore- and hindlimbs.