Penelopognathus Temporal range: Early Cretaceous, | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Order: | † Ornithischia |
Clade: | † Ornithopoda |
Superfamily: | † Hadrosauroidea |
Genus: | † Penelopognathus |
Species: | †P. weishampeli |
Binomial name | |
†Penelopognathus weishampeli Godefroit, Li, and Shang, 2005 | |
Penelopognathus (meaning "wild duck jaw") is a genus of dinosaur which lived during the Early Cretaceous. It was an iguanodont ancestral to hadrosaurids, which also contributes to the hypothesis that hadrosauroids originated in Asia. [1] Fossils have been found in the Bayin-Gobi Formation in what is now China. The type species, Penelopognathus weishampeli, named after David Weishampel, was described by Godefroit, Li, and Shang in 2005, based on fragmentary jaw fossils. [1]
Its jaw is similar to that of Altirhinus and Probactrosaurus , suggesting that Penelopognathus was related to the two genera. [1] Prieto-Márquez and Carrera Farias (2021) found that Telmatosaurus was the sister taxon to Penelopognathus, which was also found to be closely related to Lophorhothon , as opposed to Tethyshadros . [2] Penelopognathus grew up to around 5 metres (16 ft) long and around 2 metres (6.6 ft) tall when fully grown.
Shantungosaurus is a genus of very large saurolophine hadrosaurid dinosaur found in the Late Cretaceous Wangshi Group of the Shandong Peninsula in China, containing a single species, Shantungosaurus giganteus. The stratigraphic interval of Shantungosaurus ranges from the top of the Xingezhuang Formation to the middle of the Hongtuya Formation, middle to late Campanian in age. Shantungosaurus is so far the largest hadrosauroid taxon in the world, reaching between 15 metres (49 ft) to 16.6 metres (54 ft) in length and 13 metric tons to 16 metric tons in body mass.
Aralosaurus was a genus of hadrosaurid dinosaur that lived during the Late Cretaceous in what is now Kazakhstan. It is known only by a posterior half of a skull and some post-cranial bones found in the Bostobe Formation in rocks dated from the Upper Santonian-Lower Campanian boundary, at about 83.6 Ma. Only one species is known, Aralosaurus tuberiferus, described by Anatoly Konstantinovich Rozhdestvensky in 1968. The genus name means Aral Sea lizard, because it was found to the northeast of the Aral Sea. The specific epithet tuberiferus means bearing a tuber because the posterior part of the nasal bone rises sharply in front of the orbits like an outgrowth. Aralosaurus was originally reconstituted with a nasal arch similar to that of North American Kritosaurus. For many years, Aralosaurus was thus placed in the clade of the Hadrosaurinae. This classification was invalidated in 2004, following the re-examination of the skull of the animal which allowed to identify in Aralosaurus many typical characters of Lambeosaurinae. In particular, this study revealed that Aralosaurus had a hollow bony structure located far in front of the orbits, which communicated with the respiratory tract. This structure being broken at its base, its shape and size remains undetermined. More recently, Aralosaurus has been identified as the most basal Lambeosaurinae, and placed with its close relative Canardia from the upper Maastrichtian of France in the new clade of Aralosaurini.
Bactrosaurus is a genus of herbivorous hadrosauroid dinosaur that lived in Asia during the Late Cretaceous, about 96-85 million years ago. The position Bactrosaurus occupies in the Cretaceous makes it one of the earliest known hadrosauroids, and although it is not known from a full skeleton, Bactrosaurus is one of the best known of these early hadrosauroids, making its discovery a significant finding.
Amurosaurus is a genus of lambeosaurine hadrosaurid dinosaur found in the latest Cretaceous period of eastern Asia. Fossil bones of adults are rare, but an adult would most likely have been at least 6 metres (20 ft) long. According to Gregory S. Paul, it was about 8 metres (26 ft) long and weighed about 3,000 kilograms (6,600 lb).
Anasazisaurus is a genus of saurolophine hadrosaurid ("duckbill") ornithopod dinosaur that lived about 74 million years ago, in the Late Cretaceous Period. It was found in the Farmington Member of the Kirtland Formation, in the San Juan Basin of New Mexico, United States. Only a partial skull has been found to date. It was first described as a specimen of Kritosaurus by Jack Horner, and has been intertwined with Kritosaurus since its description. It is known for its short nasal crest, which stuck out above and between its eyes for a short distance.
Pararhabdodon is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Group of Spain. The first remains were discovered from the Sant Romà d’Abella fossil locality and assigned to the genus Rhabdodon, and later named as the distinct species Pararhabdodon isonensis in 1993. Known material includes assorted postcranial remains, mostly vertebrae, as well as maxillae from the skull. Specimens from other sites, including remains from France, a maxilla previously considered the distinct taxon Koutalisaurus kohlerorum, an additional maxilla from another locality, the material assigned to the genera Blasisaurus and Arenysaurus, and the extensive Basturs Poble bonebed have been considered at different times to belong to the species, but all of these assignments have more recently been questioned. It was one of the last non-avian dinosaurs known from the fossil record that went extinct during the Cretaceous-Paleogene extinction event.
Equijubus, is a genus of herbivorous hadrosauroid dinosaur from the Early Cretaceous of northwestern China.
Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to Iguanodon. Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian Probactrosaurus and Altirhinus, have traditionally been included in a paraphyletic "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only Iguanodon and perhaps its closest relatives.
Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadrosaurinae is based on the genus Hadrosaurus which was found in more recent studies to be more primitive than either lambeosaurines or other traditional "hadrosaurines", like Edmontosaurus and Saurolophus. As a result of this, the name Hadrosaurinae was dropped or restricted to Hadrosaurus alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae. Recent phylogenetic work by Hai Xing indicates that Hadrosaurus is placed within the monophyletic group containing all non-lambeosaurine hadrosaurids. Under this view, the traditional Hadrosaurinae is resurrected, with the Hadrosauridae being divided into two clades: Hadrosaurinae and Lambeosaurinae.
Koutalisaurus is a potentially dubious genus of extinct hadrosaurid dinosaur from the Arenysaurini. It is based on a mostly complete dentary from the Maastrichtian-age Upper Cretaceous Tremp Formation near the town of Abella de la Conca, Lleida, Spain.
Wulagasaurus is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China. Its remains were found in a bonebed in the middle Maastrichtian-age Yuliangze Formation, dated to 69 million years ago. This bonebed is otherwise dominated by fossils of the lambeosaurine hadrosaurid Sahaliyania. Wulagasaurus was named by Pascal Godefroit and colleagues in 2008. Only partial remains are known at this time. It is one of several hadrosaurids from the Amur River region named since 2000. The type and only species to date is W. dongi, named in honor of Chinese paleontologist Dong Zhiming. In 2010 Gregory S. Paul estimated its size at 9 meters and 3 tonnes.
The Bayin-Gobi Formation is a geological formation in Inner Mongolia, north China, whose strata date back to Albian of the Early Cretaceous period. Dinosaur remains are among the fossils that have been recovered from the formation.
Tethyshadros is a genus of hadrosauroid dinosaur from Trieste, Italy. The type and only species is T. insularis.
Glishades is a genus of hadrosauroid dinosaur that lived in the Late Cretaceous in North America. It is based on AMNH 27414, two partial premaxillae discovered in the Upper Cretaceous rocks of the upper Two Medicine Formation in Montana, dated to about 74.5 million years ago. Cladistic analysis conducted by Prieto-Márquez suggests that Glishades is a non-hadrosaurid hadrosauroid, probably a sister taxon to Bactrosaurus johnsoni. The type species is Glishades ericksoni.
Willinakaqe is a dubious genus of saurolophine hadrosaurid dinosaur described based on fossils from the late Cretaceous of the Río Negro Province of southern Argentina.
Kundurosaurus is an extinct genus of saurolophine hadrosaurid dinosaur known from the Latest Cretaceous of Amur Region, Far Eastern Russia. It contains a single species, Kundurosaurus nagornyi.
This timeline of hadrosaur research is a chronological listing of events in the history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosaurs. Scientific research on hadrosaurs began in the 1850s, when Joseph Leidy described the genera Thespesius and Trachodon based on scrappy fossils discovered in the western United States. Just two years later he published a description of the much better-preserved remains of an animal from New Jersey that he named Hadrosaurus.
The Basturs Poble bonebed is a mega-bonebed of hadrosaur dinosaur fossils, discovered in Catalonia, Spain. Hundreds of hadrosaur fossils have been found at the site, which would have been on a large island during the Late Cretaceous when the animals preserved were alive. Despite the enormous amount of specimens, taxonomically informative material has been scarce at the site, leading to extensive debate as to its nature. The number of species present, age of the individuals present in the sample, and taxonomic identity of the remains have been the primary matters of debate. Previous considered to represent Koutalisaurus, Pararhabdodon, or multiple, perhaps dwarf species, it is currently thought that a single, indeterminate species of lambeosaurine was present at the site, and that individuals of many different ages were present.
Fylax is a genus of hadrosauroid ornithopod from the Late Cretaceous Figuerola Formation of Spain. The genus contains a single species, Fylax thyrakolasus, known from a nearly complete left dentary.