Trachodon Temporal range: Late Cretaceous, | |
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Illustration of the isolated teeth | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Ornithopoda |
Family: | † Hadrosauridae |
Genus: | † Trachodon Leidy, 1856 |
Species: | †T. mirabilis |
Binomial name | |
†Trachodon mirabilis Leidy, 1856 | |
Synonyms | |
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Trachodon (meaning "rough tooth") is a dubious genus of hadrosaurid dinosaur based on teeth from the Campanian-age Upper Cretaceous Judith River Formation of Montana, U.S. [1] It is a historically important genus with a convoluted taxonomy that has been all but abandoned by modern dinosaur paleontologists. [2]
Despite being used for decades as the iconic duckbill dinosaur, the material it is based on is composed of teeth from both duckbills and ceratopsids (their teeth have a distinctive double root [3] ), and its describer, Joseph Leidy, came to recognize the difference and suggested limiting the genus to what would now be seen as ceratopsid teeth. [2] Restricted to the duckbill teeth, it may have been a lambeosaurine. [4]
In 1856, Joseph Leidy received fragmentary remains from the Judith River Formation, collected by Ferdinand Vandeveer Hayden. From these bones, he provided the first names for North American dinosaurs: Deinodon , Palaeoscincus , Trachodon, and Troodon (then spelled Troödon). [1] [5] The type species of Trachodon is T. mirabilis. The generic name is derived from Greek τραχυς, trakhys, "rough", and όδον, odon, "tooth", referring to the granulate inner surface of one of the teeth. The specific name means "marvelous" in Latin.
Trachodon was based on ANSP 9260, seven unassociated teeth, one of which had double roots. With better remains from Hadrosaurus , he began to reconsider his taxonomy, and suggested, at least informally, that Trachodon should refer to the double-rooted tooth, and the other teeth should be referred to Hadrosaurus. [6] In the Bone Wars that followed, and their wake, the taxonomy of Trachodon and its relatives became increasingly confusing, [2] with one author going so far as to sink all known hadrosaur species into Trachodon except for Claosaurus agilis , [7] but as new material was described from the Rocky Mountain region, Alberta, and Saskatchewan, later authors began progressively restricting the reach of this genus. [4] [8]
By 1942, and the publication of the influential Lull-Wright monograph on duckbills, its holotype was regarded as "typical of all the genera of hadrosaurian dinosaur", except for the roughened margin that gave it its name, and that they regarded as due to the tooth having not been used (p. 149). [9] The name is no longer in use, except in historical discussions, and is considered a nomen dubium. [10] [11] [12]
In 1936, paleontologist Charles Sternberg compared the holotype teeth of Trachodon mirabilis to those of more completely known hadrosaurids and noted that they were most similar to those of lambeosaurines. [4] It has been reported that paleontologist John R. Horner also found that Trachodon teeth compare well with the teeth of lambeosaurines, specifically Corythosaurus , though they also share similarities with the genus Prosaurolophus . [13]
Numerous species have been referred to this genus, mostly before World War I. Only those originally named as a species of Trachodon are considered here.
Type species: T. mirabilis Leidy, 1856 [1]
Other species:
As a hadrosaurid, Trachodon would have been a large, bipedal/quadrupedal herbivore. [12]
Hadrosaurids, or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as Edmontosaurus and Parasaurolophus, was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, and South America.
Edmontosaurus, often colloquially and historically known as Anatosaurus, is a genus of hadrosaurid (duck-billed) dinosaur It contains two known species: Edmontosaurus regalis and Edmontosaurus annectens. Fossils of E. regalis have been found in rocks of western North America that date from the late Campanian age of the Cretaceous period 73 million years ago, while those of E. annectens were found in the same geographic region from rocks dated to the end of the Maastrichtian age, 66 million years ago. Edmontosaurus was one of the last non-avian dinosaurs to ever exist, and lived alongside dinosaurs like Triceratops, Tyrannosaurus, Ankylosaurus, and Pachycephalosaurus shortly before the Cretaceous–Paleogene extinction event.
Lambeosaurus is a genus of hadrosaurid dinosaur that lived about 75 million years ago, in the Late Cretaceous period of North America. This bipedal/quadrupedal, herbivorous dinosaur is known for its distinctive hollow cranial crest, which in the best-known species resembled a mitten. Several possible species have been named, from Canada, the United States, and Mexico, but only the two Canadian species are currently recognized as valid.
Hypacrosaurus was a genus of duckbill dinosaur similar in appearance to Corythosaurus. Like Corythosaurus, it had a tall, hollow rounded crest, although not as large and straight. It is known from the remains of two species that spanned 75 to 67 million years ago, in the Late Cretaceous of Alberta, Canada, and Montana, United States, and is the latest hollow-crested duckbill known from good remains in North America. It was an obscure genus until the discovery in the 1990s of nests, eggs, and hatchlings belonging to H. stebingeri.
Gryposaurus was a genus of duckbilled dinosaur that lived about 80 to 75 million years ago, in the Late Cretaceous of North America. Named species of Gryposaurus are known from the Dinosaur Park Formation in Alberta, Canada, and two formations in the United States: the Lower Two Medicine Formation in Montana and the Kaiparowits Formation of Utah. A possible additional species from the Javelina Formation in Texas may extend the temporal range of the genus to 66 million years ago.
Kritosaurus is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard", but is often mistranslated as "noble lizard" in reference to the presumed "Roman nose".
Stephanosaurus is a dubious genus of hadrosaurid dinosaur with a complicated taxonomic history.
Thespesius is a dubious genus of hadrosaurid dinosaur from the late Maastrichtian-age Upper Cretaceous Lance Formation of South Dakota.
Dysganus (dis-GANN-us) is a dubious genus of ceratopsian dinosaur from the Campanian stage of the Late Cretaceous. The fossil teeth referred to Dysganus were first collected by Charles Sternberg from the Cretaceous Judith River Formation of Montana and later described by Edward Drinker Cope. All of the species are now seen as dubious Ceratopsians, though referred material from tyrannosaurids and hadrosaurids were found in New Mexico.
The Mandschurosaurus are an extinct genus of hadrosaurids based on material from the Late Cretaceous of China and possibly also the Early Cretaceous of Laos. It was the first dinosaur genus named from China, and a mounted skeleton based on the holotype of M. amurensis is on display at the Central Geological and Prospecting Museum in St. Petersburg; however, much of the skeleton is plaster.
Nipponosaurus is a lambeosaurine hadrosaur from sediments of the Yezo Group, in Sinegorsk on the island of Sakhalin, which was part of Japan at the time of the species' classification. The type and only species is N. sachalinensis, known only from a single juvenile specimen discovered in 1934 and named in 1936, by Takumi Nagao, with further material of the same individual found in 1937. Since then, the taxon has been largely ignored, and its validity has been doubted, with synonymy with other Asian hadrosaurs or status as a nomen dubium being suggested. Redescriptions from 2004 and 2017, however, have supported recognition as a distinct species. Dating the only specimen has been difficult, but based on associated mollusc taxa, the species likely lived sometime in the upper Santonian or lower Campanian, around 80 million years ago.
Prosaurolophus is a genus of hadrosaurid dinosaur from the Late Cretaceous of North America. It is known from the remains of at least 25 individuals belonging to two species, including skulls and skeletons, but it remains obscure. Its fossils have been found in the late Campanian-age Upper Cretaceous Dinosaur Park Formation in Alberta, and the roughly contemporaneous Two Medicine Formation in Montana, dating to around 75.5-74.0 million years ago. Its most recognizable feature is a small solid crest formed by the nasal bones, sticking up in front of the eyes.
Diclonius is a genus of dinosaur from the Late Cretaceous. It was a hadrosaur based solely on teeth. Its fossils were found in the Judith River Formation of Montana, northern US. The name is in reference to the method of tooth replacement, in which newly erupting replacement teeth could be in functional use at the same time as older, more worn teeth. Thus, the number of "sprouting" teeth was doubled in comparison to Monoclonius, which used only one set of teeth at a time and which Cope named in the same paper.
Orthomerus is a genus of dubious hadrosaurid dinosaur from the Late Cretaceous of the Netherlands. It is today an obscure genus, but in the past was conflated with the much better known Telmatosaurus.
Koutalisaurus is a potentially dubious genus of extinct hadrosaurid dinosaur from the Arenysaurini. It is based on a mostly complete dentary from the Maastrichtian-age Upper Cretaceous Tremp Formation near the town of Abella de la Conca, Lleida, Spain.
Edmontosaurus regalis is a species of comb-crested hadrosaurid dinosaur. Fossils of E. regalis have been found in rocks of western North America that date from the late Campanian age of the Cretaceous Period 73 million years ago, but it may have possibly lived into the early Maastrichtian.
Edmontosaurus annectens, often colloquially and historically known as the Anatosaurus, is a species of flat-headed saurolophine hadrosaurid dinosaur from the late Maastrichtian age at the very end of the Cretaceous period, in what is now western North America. Remains of E. annectens have been preserved in the Frenchman, Hell Creek, and Lance Formations. All of these formations are dated to the late Maastrichtian age of the Late Cretaceous period, which represents the last three million years before the extinction of the dinosaurs. E. annectens is also found in the Laramie Formation and magnetostratigraphy suggests an age of 69-68 Ma for the Laramie Formation. Edmontosaurus annectens is known from numerous specimens, including at least twenty partial to complete skulls, discovered in the U.S. states of Montana, South Dakota, North Dakota, Wyoming, and Colorado, as well as the Canadian province of Saskatchewan. It had an extremely long and low skull and was quite a large animal, growing up to approximately 12 metres (39 ft) in length and 5.6 metric tons in average asymptotic body mass, although it could have been even larger. E. annectens exhibits one of the most striking examples of the "duckbill" snout that is common to hadrosaurs. It has a long taxonomic history and specimens have at times been classified as Diclonius, Trachodon, Hadrosaurus, Claosaurus, Thespesius, Anatosaurus, and Anatotitan before all being grouped together in Edmontosaurus
Hadrosaurids, also commonly referred to as duck-billed dinosaurs or hadrosaurs, were large terrestrial herbivores. The diet of hadrosaurid dinosaurs remains a subject of debate among paleontologists, especially regarding whether hadrosaurids were grazers who fed on vegetation close to the ground, or browsers who ate higher-growing leaves and twigs. Preserved stomach content findings have indicated they may have been browsers, whereas other studies into jaw movements indicate they may have been grazers.
Kritosaurini is a tribe of saurolophine hadrosaurid dinosaurs from the Late Cretaceous.
This timeline of hadrosaur research is a chronological listing of events in the history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosaurs. Scientific research on hadrosaurs began in the 1850s, when Joseph Leidy described the genera Thespesius and Trachodon based on scrappy fossils discovered in the western United States. Just two years later he published a description of the much better-preserved remains of an animal from New Jersey that he named Hadrosaurus.