Calvarius Temporal range: Late Cretaceous, | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ornithopoda |
Clade: | † Styracosterna |
Genus: | † Calvarius Prieto-Márquez & Sellés, 2023 |
Species: | †C. rapidus |
Binomial name | |
†Calvarius rapidus Prieto-Márquez & Sellés, 2023 | |
Calvarius (meaning "suffering") is a genus of styracosternan ornithopod from the Late Cretaceous Talarn Formation of Spain. The genus contains a single species, Calvarius rapidus, known from a single metatarsal. [1]
The holotype specimen, MCD-8734, is a single fourth metatarsal discovered in 2019 at the Pallars Jussà locality of the Talarn Formation of the Tremp Group, located in Catalonia, Spain.
It was named as the holotype of a new genus of styracosternan dinosaur in 2023 by Albert Prieto-Márquez and Albert Sellés. The genus name, "Calvarius", is Latin for "suffering" (cf. Calvary ), and refers to the type locality, Serrat del Calvari, as well as the genus' proximity to the K-Pg extinction event. The specific name, "rapidus", means "rapid", and refers to its likely cursorial habits. [1]
Calvarius was in 2023 assigned by its describers to the Styracosterna. A more precise position could not be determined because a phylogenetic analysis only places it in a large polytomy of styracosternans. [1]
The highly modified metatarsal of Calvarius has no known equivalents among other ornithopods. It was convergently more similar to those of more basal ornithischians such as Hypsilophodon and Dysalotosaurus than to other styracosternans, and may have filled their niche on its island habitat. [1]
The Talarn Formation also produced the remains of the troodontid Tamarro insperatus . [2]
Hadrosaurus is a genus of hadrosaurid ornithopod dinosaurs that lived in North America during the Late Cretaceous Period in what is now the Woodbury Formation In Pennsylvania about 78-80 Ma. The holotype specimen was found in fluvial marine sedimentation, meaning that the corpse of the animal was transported by a river and washed out to sea.
Secernosaurus is a genus of herbivorous dinosaur. Secernosaurus was a hadrosaur, a "duck-billed" dinosaur which lived during the Late Cretaceous.
Pararhabdodon is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Group of Spain. The first remains were discovered from the Sant Romà d’Abella fossil locality and assigned to the genus Rhabdodon, and later named as the distinct species Pararhabdodon isonensis in 1993. Known material includes assorted postcranial remains, mostly vertebrae, as well as maxillae from the skull. Specimens from other sites, including remains from France, a maxilla previously considered the distinct taxon Koutalisaurus kohlerorum, an additional maxilla from another locality, the material assigned to the genera Blasisaurus and Arenysaurus, and the extensive Basturs Poble bonebed have been considered at different times to belong to the species, but all of these assignments have more recently been questioned. It was one of the last non-avian dinosaurs known from the fossil record that went extinct during the Cretaceous-Paleogene extinction event.
Lambeosaurinae is an extinct group of crested hadrosaurid dinosaurs.
Koutalisaurus is a potentially dubious genus of extinct hadrosaurid dinosaur from the Arenysaurini. It is based on a mostly complete dentary from the Maastrichtian-age Upper Cretaceous Tremp Formation near the town of Abella de la Conca, Lleida, Spain.
Pamparaptor is an extinct genus of paravian theropod dinosaur from the Late Cretaceous Portezuelo Formation of the Neuquén province in Argentine Patagonia. Its precise classification is uncertain, but the authors who described this taxon have argued that it is a dromaeosaurid. The genus contains a single species, P. micros, which is known from a single specimen consisting of a mostly complete and fully-articulated left foot, which preserves the iconic dromaeosaur-like "killing claw".
Latirhinus is an extinct genus of lambeosaurine hadrosaurid dinosaur from the Late Cretaceous of Mexico. The type species, Latirhinus uitstlani, was named in 2012 on the basis of a partial skeleton from the Campanian-age Cerro del Pueblo Formation. The specific name uitstlani means "southern" in the Náhuatl language of Mexico, a reference to the species' southern occurrence in the Cretaceous landmass Laramidia.
Albertadromeus is an extinct genus of orodromine thescelosaurid dinosaur known from the upper part of the Late Cretaceous Oldman Formation of Alberta, Canada. It contains a single species, Albertadromeus syntarsus.
Canardia is an extinct genus of lambeosaurine dinosaur known from the Late Cretaceous Marnes d'Auzas Formation of Haute-Garonne department, in Occitanie region, southwestern France. The type species Canardia garonnensis was first described and named by Albert Prieto-Márquez, Fabio M. Dalla Vecchia, Rodrigo Gaete and Àngel Galobart in 2013. It is only known from juvenile specimens. The name of the genus comes from “canard”, the French word for “duck”, an allusion to the fact that this animal belongs to the hadrosaurids which are also known as duck-billed dinosaurs. The specific epithet garonnensis refers to the Haute-Garonne department where this dinosaur has been found. Although universally recognized as a lambeosaurine, its precise position within them is debated. Some authors consider it as a close relative of the genus Aralosaurus from Central Asia with which it would form the tribe Aralosaurini, while others include it in a more derived clade, the Arenysaurini in which all lambeosaurines from Europe and North Africa are placed. Canardia was one of the last non-avian dinosaurs and lived between 67,5 and 66 my on the former Ibero-Armorican Island, which included much of France and Spain.
Bonapartesaurus is an extinct genus of herbivorous ornithopod dinosaur belonging to Hadrosauridae, which lived in the area of modern Argentina during the Campanian and Maastrichtian stages of the Late Cretaceous.
Adynomosaurus is a genus of lambeosaurine dinosaur from the Late Cretaceous of what is now Catalonia, Spain. First discovered in 2012, it was named in 2019 with the type and only species being Adynomosaurus arcanus. It is only known from scant material, but is distinguished from other hadrosaurs by its weakly developed shoulder blade which would have had underdeveloped musculature, which lends it its scientific name, partially from the Greek word for "weak". Its exact relationships with other hadrosaurs remain unresolved, with it not consistently being recovered as a relative of any other specific genera, though some studies have allied it with Tsintaosaurini or even found it outside of Hadrosauridae. It would have lived as part of a diverse coastal estuary ecosystem, made up of meandering rivers and mud flats. The discovery of Adynomosaurus adds to the very incomplete fossil record of hadrosaurid dinosaurs in the Late Cretaceous of Europe, and it fits into a picture of major ecological turnover that was occurring during the Maastrichtian stage in the region.
Pareisactus is a genus of rhabdodontid ornithopod dinosaur from the Late Cretaceous Conquès Member of the Tremp Formation in the Southern Pyrenees of Spain. The type and only species is P. evrostos, known only from a single scapula.
Aquilarhinus is a genus of hadrosaurid ornithopod dinosaur from the Aguja Formation from Texas in the United States. The type and only species is Aquilarhinus palimentus. Due to its unusual dentary, it has been inferred to have had shovel-like beak morphology, different from the beaks of other hadrosaurs. It was originally classified as a Kritosaurus sp. before being reclassified as a new genus in 2019.
Tamarro is a genus of troodontid theropod from the Late Cretaceous Talarn Formation of Spain. The genus contains a single species, Tamarro insperatus, known from a partial metatarsal described in 2021.
Fylax is a genus of hadrosauroid ornithopod from the Late Cretaceous Figuerola Formation of Spain. The genus contains a single species, Fylax thyrakolasus, known from a nearly complete left dentary.
Garumbatitan is an extinct genus of somphospondylan sauropod dinosaur from the Cretaceous Arcillas de Morella Formation of Spain. The genus contains a single species, G. morellensis, known from multiple partial skeletons.
Farlowichnus is an ichnogenus of small theropod dinosaur footprint. It includes a single species, F. rapidus, known from prints found in the Early Cretaceous Botucatu Formation of Brazil. Farlowichnus is known from several fossil trackways that indicate that it was likely a cursorial animal that was well-adapted to desert environments.