2022 in paleomalacology

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This list of fossil molluscs described in 2022 is a list of new taxa of fossil molluscs that were described during the year 2022, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2022.

Ammonites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acanthohoplites albedocostatus [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Acantholytoceras costaevariavilis [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Aulatornoceras beyrichi [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Buedesheimites [2]	Gen. et sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Genus includes new species B. housei. Announced in 2022; the final article version was published in 2023.
Cadoceras (Catacadoceras) imlayi [3]	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)	Chinitna Formation	United States (  Alaska)
Cadoceras kniazevi [3]	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)		Russia
Cadoceras (Paracadoceras) pseudofrearsi [3]	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)		Russia
Cadomites zelenchukensis [4]	Sp. nov	Valid	Mitta	Middle Jurassic (Bathonian)		Russia (  Karachay-Cherkessia)
Collignoniceras rhodanicum [5]	Sp. nov	Valid	Amédro, Baudouin & Delanoy in Amédro et al.	Late Cretaceous (Turonian)		France
Crassotornoceras tenue [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Crioceratites tepehuaensis [6]	Sp. nov	Valid	Esquivel-Macías et al.	Early Cretaceous		Mexico
Cryptomorphiceras [7]	Gen. et comb. et 2 sp. nov	Valid	Marchesi, Balini & Jenks	Late Triassic (Carnian)		United States (  Nevada)	A member of Trachyceratinae. The type species is C. compressum (Johnston, 1941); genus also includes new species C. crassum, and C. levicostatum. Announced in 2022; the final article version was published in 2023.
Dactylioceras kopiki [8]	Sp. nov	Valid	Rogov in Rogov & Lutikov	Early Jurassic (Toarcian)		Canada (  Northwest Territories)   Norway   United States (  Alaska)   Russia?	A member of the family Dactylioceratidae.
Diplesioceras fischeri [9]	Sp. nov	Valid	Dietze, Schweigert & Chandler	Middle Jurassic (Bajocian)	Oolithe de Bayeux Formation	France
Egoianites jauberti [10]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)		France	A member of the family Heteroceratidae.
Epitornoceras transmediterraneum [11]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)		Morocco	A member of the family Tornoceratidae.
Forresteria (Harleites) seidli [12]	Sp. nov	Valid	Summesberger et al.	Late Cretaceous (Santonian)	Gosau Group (Streiteck Formation)	Austria
Heteroceras arnaudi [10]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)		France	A member of the family Heteroceratidae.
Heteroceras munierchalmasi [10]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)		France	A member of the family Heteroceratidae.
Heteroceras penagei [10]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)		France	A member of the family Heteroceratidae.
Hildoceras appenninicus [13]	Sp. nov	Valid	Ridente	Early Jurassic (Toarcian)		Italy
Kepplerites (Gowericeras) copernici [14]	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)		Germany
Kepplerites (Gowericeras) halleyi [14]	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)		France   Germany   Russia (  Kostroma Oblast)
Kepplerites (Toricellites) besseli [14]	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)		Germany
Kutatissites sudai [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Lentitornoceras [11]	Gen. et sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Red Ironstone Formation	Germany	A member of the family Tornoceratidae. The type species is L. materni.
Linguatornoceras acutilobatum [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Linguatornoceras sandense [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Lissoceras kubanense [15]	Sp. nov	Valid	Mitta	Middle Jurassic (Bajocian)		Russia (  Karachay-Cherkessia)
Lissoceras paviai [15]	Sp. nov	Valid	Mitta	Middle Jurassic (Bajocian)		Russia (  Karachay-Cherkessia)
Megatyloceras leteilensis [16]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Aptian)		France	A member of the family Douvilleiceratidae.
Nejdia arabica [17]	Sp. nov	Valid	Gelin	Early Jurassic (Toarcian)	Marrat Formation	Saudi Arabia
Paracrochordiceras watanabei [18]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of the family Acrochordiceratidae.
Paradanubites ozashiense [18]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Danubitoidea belonging to the family Danubitidae.
Parahedenstroemia? tatjanae [19]	Sp. nov	Valid	Đaković, Krystyn & Sudar	Early Triassic		Montenegro   Russia?	A member of the family Hedenstroemiidae.
Parapopanoceras involutum [18]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Ceratitida belonging to the group Megaphyllitoidea and the family Parapopanoceratidae.
Paucitornoceras [11]	Gen. et comb. nov	Valid	Korn & Bockwinkel	Devonian	Red Ironstone Formation	Germany   United States (  New York)	A member of the family Tornoceratidae. The type species is "Goniatites" paucistriatus d`Archiac & de Verneuil (1842); genus also includes "Tornoceras" rhysum Clarke (1898).
Ponticeras materni [20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Red Ironstone Formation	Germany	A member of Agoniatitida belonging to the group Gephuroceratina and the family Ponticeratidae.
Procheloniceras yajimai [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Pseudocrioceras mazierei [21]	Sp. nov	Valid	Frau & Delanoy	Early Cretaceous (Barremian/Aptian transition)		France
Pseudosaynella pacifica [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Psilokhvalynites [18]	Gen. et sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Dinaritoidea belonging to the family Khvalynitidae. The type species is P. takaizumii.
Shasticrioceras nakazawai [1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Sigaloceras (Gulielmina) fabricius [14]	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)		Germany
Tornoceras incisum [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Tornoceras ventrovaricatum [2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation	Germany	A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Witchellia companyi [22]	Sp. nov	Valid	Sandoval	Middle Jurassic (Bajocian)		Spain	A member of the family Sonniniidae.
Witchellia striata [22]	Sp. nov	Valid	Sandoval	Middle Jurassic (Bajocian)		Spain	A member of the family Sonniniidae.

Ammonite research

• Mironenko & Naugolnykh (2022) describe a collection of cephalopod jaws from the Permian (Artinskian) Divjinskian Formation (Sverdlovsk Oblast, Russia), interpreted as likely fossil material of goniatitid ammonites belonging to the genus Uraloceras , and infer from the anatomy of the studied jaws that representatives of the genus Uraloceras were likely active predators. ^[23]
• A study on the morphological change through ontogeny of beyrichitine ammonites from the Anisian of Nevada is published by Bischof, Schlüter & Lehmann (2022). ^[24]
• A study aiming to determine whether the environmental changes around the Triassic–Jurassic boundary influenced development of septal thickness in ammonites, as well aiming to determine the relation between septal thickness and paleolatitudinal occurrences, and whether ontogenetic septal thickness-trajectories reflected paleogeographic origin or phylogenetic relationships of ammonites, is published by Weber et al. (2022). ^[25]
• A study on the hydrodynamic properties of shells of Late Triassic and Early Jurassic ammonites is published by Hebdon et al. (2022). ^[26]
• A study on the functional morphology of ammonites from the Middle Triassic to the Early Jurassic, reevaluating possible causes of an overall shift of ammonite shell forms during the studied periods, is published by Hebdon et al. (2022), who interpret their findings as indicating that the trade-offs between coasting efficiency, volume accretion per unit surface area accreted and diameter accretion per unit of surface area (three factors proposed as explanations of the shift of shell forms) were identical in the Triassic and in the Jurassic. ^[27]
• Jaws of ammonites belonging to the Boreal family Polyptychitidae, with lower jaws closely resembling jaws of craspeditids, cardioceratids and hoplitoids, are described for the first time from the Valanginian of Russia by Mironenko & Mitta (2022). ^[28]
• A study on changes of septal spacing between succeeding chambers of the shell of Gaudryceras tenuiliratum during its ontogeny is published by Kawakami, Uchiyama & Wani (2022). ^[29]
• A study on the impact of the conch shape on ammonite locomotion, based on data obtained from tests of 3D-printed, biomimetic robots representing the oxycone morphology (based on Sphenodiscus lobatus and S. lenticularis), the serpenticone morphology (based on Dactylioceras commune), the sphaerocone morphology (based on Goniatites crenistria) and the morphospace center, is published by Peterman & Ritterbush (2022). ^[30]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aphelaeceras azzelmattiense [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian–Viséan boundary interval)		Algeria	A member of the family Trigonoceratidae.
Arctoteuthis angusta [32]	Sp. nov	Valid	Efremenko	Early Cretaceous		Russia	A belemnite belonging to the family Cylindroteuthididae.
Binoleniceras [33]	Gen. et sp. nov	Valid	Afhüppe & Becker	Devonian (Givetian)	Hagen-Balve Formation	Germany	A member of Discosorida belonging to the family Entimoceratidae. The type species is B. stichlingi.
Boreioteuthis mirifica [32]	Sp. nov	Valid	Efremenko	Early Cretaceous		Russia	A belemnite belonging to the family Cylindroteuthididae.
Chisiloceras maocaopuense [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)		China	A member of the family Endoceratidae.
Domatoceras tuckeri [35]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States (  Texas)	A member of the family Grypoceratidae.
Endolobus sturgeoni [35]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States (  Texas)	A member of the family Koninckioceratidae.
Enoploceras rieberi [36]	Sp. nov	Valid	Pieroni	Middle Triassic (Anisian)	Besano Formation	Switzerland
Eximioceras [37]	Gen. et sp. nov	Valid	Shchedukhin	Early Permian		Russia (  Bashkortostan)	A member of the family Rhiphaeoceratidae. The type species is E. venustum.
Holmiceras havelense [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany
Kotoceras minosiphonatum [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)		China	A member of the family Endoceratidae.
Lispoceras orbis [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Lituites baculus [38]	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician		Sweden
Lituites bottkei [38]	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician		Poland   Sweden
Lituites clavis [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites dewitzi [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites kruegeri [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites nebeni [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Maccoyoceras concavum [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian–Viséan boundary interval)		Algeria	A member of the family Trigonoceratidae.
Maccoyoceras habadraense [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Maccoyoceras pentagonum [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Maccoyoceras saharensis [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Metacoceras quadratum [35]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States (  Texas)	A member of the family Tainoceratidae.
Michelinoceras minutum [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian and Sandbian)		China
Michelinoceras multiseptum [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)		China
Pararhiphaeoceras parvum [37]	Sp. nov	Valid	Shchedukhin	Early Permian		Russia (  Bashkortostan)	A member of the family Rhiphaeoceratidae.
Planetoceras destrictum [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Planetoceras transforme [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Pleuronautilus chisatoi [40]	Sp. nov	Valid	Ehiro	Permian (Wordian)	Takakurayama Formation	Japan	A member of the family Tainoceratidae.
Proterovaginoceras rapidum [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)		China	A member of the family Endoceratidae.
Rhynchorthoceras kranepuhlense [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany   Poland
Rhynchorthoceras rugium [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany
Rineceras multituberculatum [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Rineceras rectangulatum [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Rineceras tenerum [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Solenochilus jackense [35]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States (  Texas)	A member of the family Solenochilidae.
Stroboceras ancilis [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Viséan)		Algeria	A member of the family Trigonoceratidae.
Stroboceras mane [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Stroborineceras [31]	Gen. et 2 sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae. The type species is S. insalahensis; genus also includes new species S. felis and possibly also "Nautilus (Trematodiscus)" meekianus Winchell (1862) and "Nautilus (Trematodiscus)" strigatus Winchell (1862).
Syllipsimopodi [41]	Gen. et sp. nov	Disputed	Whalen & Landman	Carboniferous (Serpukhovian)	Bear Gulch Limestone	United States (  Montana)	A member of Coleoidea of uncertain affinities. Originally assigned to the clade Vampyropoda; Klug et al. (2023) considered it to be a probable junior synonym of Gordoniconus beargulchensis . [42] [43] The type species is S. bideni, named for Joe Biden. [44]
Thrincoceras devolvere [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Tohokubelus [45]	Gen. et sp. nov	Valid	Niko & Ehiro	Early Triassic (Olenekian)	Osawa Formation	Japan	A sinobelemnitid belemnite. Genus includes new species T. takaizumii.
Trilacinoceras filix [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany	A member of the family Lituitidae.
Trilacinoceras knoefleri [38]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany	A member of the family Lituitidae.
Trilobitoceras [31]	Gen. et sp. et comb. nov	Valid	Korn & Bockwinkel	Carboniferous		Algeria   United States (  Michigan)	A member of the family Trigonoceratidae. The type species is T. peculiaris; genus also includes "Nautilus (Trematodiscus)" planidorsalis Winchell (1862).
Vestinautilus angulatus [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Vestinautilus bicristatus [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Vestinautilus concinnus [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Vestinautilus inflexus [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Vestinautilus padus [39]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Vestinautilus papilio [31]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)		Algeria	A member of the family Trigonoceratidae.
Xainzanoceras densum [34]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Katian)	Pagoda Formation	China	Possibly a member of the family Diestoceratidae.

Other cephalopod research

• A study on the phylogenetic relationships of Cambrian and Ordovician cephalopods is published by Pohle et al. (2022). ^[46]
• Putative tentaculitoid Iwakiella ichiroi is reinterpreted as a sphaerorthoceratid orthocerid cephalopod by Niko (2022). ^[47]
• A study on the identity of enigmatic black amorphous fossils often associated with belemnoid remains, based on the analysis of fossil material from the Upper Triassic (Carnian) Polzberg Lagerstätte (Austria), is published by Lukeneder & Lukeneder (2022), who interpret the enigmatic fossils as likely representing the preservation of a mineralized and secondarily carbonized cephalic-ocular-arm-cartilage complex of the belemnoid Phragmoteuthis bisinuata. ^[48]
• A study on structures interpreted by Eduard Suess as beaks of Phragmoteuthis bisinuata is published by Doguzhaeva et al. (2022), who reinterpret the studied structures as cartilaginous remains of a prey, presumably juvenile fish; the authors also report a gladius of a previously unknown Carnian teuthid from the Cave del Predil (Italy), possibly similar to the Permian taxon Glochinomorpha stifeli, as well as an upper beak of a coleoid from the Anisian Buchenstein Formation (Italy), demonstrating typical coleoid upper beak morphology and composition, and interpreted by the authors as likely indicative of similar upper beak structure in P. bisinuata. ^[49] Their interpretation of purported beaks of P. bisinuata as remains of a prey is subsequently contested by Lukeneder & Lukeneder (2022). ^{[50] [51]}
• Belemnite species Liobelus acrei, otherwise known from the latest Jurassic and Early Cretaceous of the Boreal Realm, is reported from the Valanginian of the Vocontian Basin (France) by Mutterlose, Picollier & Dzyuba (2022), who interpret this finding as evidence of an isolated immigration of belemnites from the north in the early Valanginian, and establish two taxonomically different faunas of Tethyan belemnites. ^[52]
• A study on the age of belemnites from the lower Santonian "Sponge horizon" of the Mozhzhelloovrazhnaya Formation in the Lower Volga region between Saratov and Volgograd (Russia), some of which are interpreted as redeposited from middle and upper Coniacian deposits on the basis of strontium, carbon and oxygen isotope data (with the first representatives of the genus Belemnitella interpreted as late Coniacian in age), is published by Zakharov et al. (2022). ^[53]
• Revision of the type specimens of Eothinoceras americanum is published by Evans & Cichowolski (2022), who restrict the genus Eothinoceras to the type species, and propose a new scheme for classification of the genera belonging to the order Cyrtocerinida. ^[54]
• A study on the distribution of nautiloids throughout the Cenozoic is published by Kiel, Goedert & Tsai (2022), who interpret their findings as indicating that from the Oligocene onward nautiloids became extinct in areas where pinnipeds appeared, while cetaceans (with possible exception of simocetids and agorophiids) were not found to significantly affect the demise of nautiloids. ^[55]
• Fossil material of "Kummelonautilus" taiwanum is described from the Miocene Houdongkeng Formation (Taiwan) by Goedert, Kiel & Tsai (2022), who provisionally transfer this species to the genus Nautilus , and review the fossil record of the genus Nautilus in the Indo-Pacific region. ^[56]
• A study on the external and internal morphology, affinities and ecology of Vampyronassa rhodanica, based on data on soft tissue of specimens from the La Voulte-sur-Rhône (France), is published by Rowe et al. (2022). ^[57]
• First cephalopod statoliths from the Lower Cretaceous of Poland and United Kingdom are described by Pindakiewicz et al. (2022), who compare Mesozoic statoliths to those of extant cephalopods, and find the closest resemblance between Mesozoic forms and statoliths of extant idiosepiids. ^[58]
• A study on the fossil material of Belosaepia from the Egemkapel Clay Member of the Tielt Formation and the Roubaix Clay Member of the Kortrijk Formation (Belgium), applying micro-CT imaging to the studied fossils, is published by Goolaerts et al. (2022), who identify growth lines in the studied fossils, and interpret their findings as indicating that the studied material belongs to representatives of a single species (B. tricarinata). ^[59]

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acila (Truncacila) interferencia [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A species of Acila .
Anniedhondtella [61]	Gen. et sp. nov	Valid	Jagt, Cooper & Jagt-Yazykova	Late Cretaceous (Maastrichtian)	Maastricht Formation		A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species A. rieui.
Aphanaia sobopolensis [62]	Sp. nov	Valid	Biakov & Kutygin	Early Permian		Russia (  Sakha)
Arcopagia esteponensis [63]	Sp. nov	Valid	Brunetti & Della Bella	Pliocene		Spain	A member of the family Tellinidae.
Arcopagia ligustica [63]	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene		Italy	A member of the family Tellinidae.
Arcopella crestii [63]	Sp. nov	Valid	Brunetti & Della Bella	Late Pliocene		Italy	A member of the family Tellinidae.
Arcopella pseudomista [63]	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene		Italy	A member of the family Tellinidae.
Bathymodiolus labayugensis [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A species of Bathymodiolus .
Choniocardia olivieri [64]	Nom. nov	Valid	Pacaud	Eocene		France	A member of the family Carditidae; a replacement name for Cardita (Miodon) bezanconi Cossmann (1887).
Conchocele pangasinanensis [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A member of the family Thyasiridae.
Crassatella (Bathytormus) ignava [65]	Sp. nov	Valid	Berezovsky	Eocene	Malinovskaya Formation	Ukraine	A member of the family Crassatellidae.
Crassatella lassa [66]	Sp. nov	Valid	Berezovsky	Late Eocene		Ukraine
Crassostrea sophiae [64]	Nom. nov	Valid	Pacaud	Eocene		France	A species of Crassostrea ; a replacement name for Ostrea elegans Deshayes (1832).
Cyclotellina hugoi [64]	Nom. nov	Valid	Pacaud	Eocene		France	A member of the family Tellinidae; a replacement name for Tellina (Cyclotellina) bezanconi Cossmann (1906).
Gryphaea (Gryphaea) lavernockensis [67]	Sp. nov	Valid	Hodges	Early Jurassic		United Kingdom
Kasimlara ? antiqua [68]	Sp. nov	Valid	Fang & Carter	Permian		Russia	A member of the family Kalenteridae belonging to the subfamily Healeyinae.
Liaoningoconcha [69]	Nom. nov		Fang & Carter	Late Triassic	Yangcaogou Formation	China	A member of Cardiida belonging to the family Ferganoconchidae; a replacement name for Liaoningia Yu & Dong (1993). Ceccolini & Cianferoni (2021) proposed a replacement name Sinoliaoningia for the same genus in an earlier publication. [70]
Megaxinus gorrospei [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A member of the family Lucinidae.
Meleagrinella golberti [71]	Sp. nov	Valid	Lutikov & Arp	Early Jurassic (Toarcian)		Canada   Germany   Russia   United Kingdom	A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022. [71]
Meleagrinella prima [71]	Sp. nov	Valid	Lutikov in Lutikov & Arp	Early Jurassic (Toarcian)		Russia	A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022. [71]
Monginaia [72]	Gen. et comb. nov	In press	Delvene et al.	Early Cretaceous (Barremian)	Camarillas Formation	Spain	A member of Unionida belonging to the new family Monginaiidae. Genus includes "Elliptio" galvensis Mongin (1966).
Myonia komiensiformis [73]	Sp. nov	Valid	Biakov	Permian		Russia	A member of Pholadomyida belonging to the family Sanguinolitidae.
Myonia kutygini [73]	Sp. nov	Valid	Biakov	Permian		Russia	A member of Pholadomyida belonging to the family Sanguinolitidae.
Oudardia pliorecens [74]	Sp. nov	Valid	Brunetti & Della Bella	Plio-Pleistocene		Italy	A member of the family Tellinidae.
Palaeomutela (Palaeanodonta) berrutii [75]	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Collio Formation	Italy	A member of Actinodontida belonging to the family Palaeomutelidae.
Palaeomutela (Palaeanodonta) guncinaensis [75]	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Guncina Formation	Italy	A member of Actinodontida belonging to the family Palaeomutelidae.
Peronidia iberica [74]	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene		Spain	A member of the family Tellinidae.
Peronidia intermedia [74]	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene		Italy	A member of the family Tellinidae.
Pliocardia ballesterosi [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A species of Pliocardia .
Praebarrettia magnifica [76]	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation	Jamaica	A rudist bivalve.
Psammotreta santacatalinae [63]	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene		Spain	A member of the family Tellinidae.
Sisonia [60]	Gen. et sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A bivalve of uncertain affinities. The type species is S. frijellanae.
Squiresica [77]	Gen. et comb. nov	Valid	Hybertsen, Goedert & Kiel	Oligocene	Lincoln Creek Formation	United States (  Alaska   Washington)	A member of the family Vesicomyidae. The type species is "Archivesica" knapptonensis Amano & Kiel (2007); genus also includes "Archivesica" marincovichi Kiel & Amano (2010).
Verneuilnodon [68]	Gen. et comb. nov	Valid	Fang & Carter	Permian		Russia	A member of the family Kalenteridae belonging to the subfamily Healeyinae; a new genus for "Mytilus (Modiola)" pallasi de Verneuil (1845).
Whitfieldiella catherinei [76]	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation	Jamaica	A rudist bivalve.
Whitfieldiella gracilis [76]	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation	Jamaica	A rudist bivalve.
Wolfgangella [61]	Gen. et 2 sp. nov	Valid	Jagt, Cooper & Jagt-Yazykova	Late Cretaceous (Maastrichtian)	Maastricht Formation		A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species W. neilpearti and W. ignota.

Bivalve research

• Late Triassic bivalves are identified for the first time in olistostromes from the Sêwa Formation in the Riganpeicuo area (Tibet, China) by Xiao et al. (2022). ^[78]
• A study on the impact of the Triassic–Jurassic extinction event on the body-size distribution of bivalves, based on data from three study sites in the United Kingdom, is published by Opazo & Twitchett (2022). ^[79]
• Wolkenstein (2022) presents evidence of abundance and diversity in the color patterns of Pleuronectites laevigatus from the Middle Triassic Muschelkalk of Central Europe. ^[80]
• Asato, Nakayama & Imai (2022) study color patterns of freshwater bivalves from the Lower Cretaceous Kitadani Formation (Japan), providing evidence of the presence of similar shell color patterns in extant and Early Cretaceous bivalves. ^[81]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Akburunella laminaris [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [82]
Akburunella nefanda archaica [82]	Ssp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [82]
Akburunella sinuosa [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [82]
Akburunella spinosa [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [82]
Ampezzopleura barremica [83]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Angularia corallina [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of Caenogastropoda belonging to the family Purpuroideidae.
Angulomphalus senowbarii [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of Vetigastropoda belonging to the family Ptychomphalidae.
Anisocycla subcylindrica [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A member of the family Murchisonellidae.
Antiphora aurora [83]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Aplexa canculae [86]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian)		France	A species of Aplexa .
Aplexa cotiensis [86]	Nom. nov	Valid	Pacaud	Eocene		France	A species of Aplexa; a replacement name for Physa (Aplecta) brevispirata Cossmann & Pissarro (1913).
Aplexa iota [86]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Aplexa.
Aplexa oryza [86]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Aplexa.
Aplexa risleius [86]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian)		France	A species of Aplexa.
Aplexa zhui [86]	Nom. nov	Valid	Pacaud	Early Cretaceous (Albian)	Dalazi Formation	China	A species of Aplexa; a replacement name for Physa fusiformis Zhu (1980).
Arachnotropis [87]	Gen. et comb. nov	Valid	Pérez, Ferrari & Ezcurra	Paleocene–Eocene		Antarctica   Australia	A member of the family Eucyclidae. The type species is "Calliotropis" microglyptophorus Darragh (1997); genus also includes "Calliotropis" antarchais Stilwell (2005).
Archaeocyclotus brevivillosus [88]	Sp. nov	In press	Bichain et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Cyclophoridae.
Archaeostylus [89]	Gen. et sp. nov	Valid	Brook & Hayward	Pliocene (Waipipian)	Kaawa Formation	New Zealand	A member of the family Bothriembryontidae belonging to the subfamily Placostylinae. The type species is A. manukauensis.
Astralium ena [90]	Sp. nov	Valid	Tomida, Inoue & Kase	Miocene	Ena Limestone	Japan	A species of Astralium .
Austrocypraea jimgracei [91]	Sp. nov	Valid	Southgate & Roberts	Pliocene		Australia	A member of the family Cypraeidae.
Barycypraea iungo [92]	Sp. nov	Valid	Seccombe & Aiken	Pleistocene		South Africa	A species of Barycypraea .
Bathrotomaria teodorii [93]	Sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)		France	A member of the family Pleurotomariidae.
Baylea tenera [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States (  Oklahoma)	A member of the family Eotomariidae.
Brachystomia succineiformis [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	A species of Brachystomia . Published online in 2023, but the issue date is listed as December 2022. [82]
Ceritella convexa [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of the family Ceritellidae.
Choshipleura [83]	Gen. et sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	Genus includes new species C. striata.
Clathromangelia acuticostata [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Clathromangelia .
Clathromangelia mulderi [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Clathromangelia.
Clathromangelia oliverioi [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Clathromangelia.
"Clavatula" ariejansseni [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation	Turkey	A member of the family Clavatulidae.
"Clavatula" atatuerki [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation	Turkey	A member of the family Clavatulidae.
"Clavatula" iberica [98]	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)		Spain	A member of the family Clavatulidae.
"Clavatula" irisae [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria   Slovenia	A member of the family Clavatulidae.
"Clavatula" jarzynkae [97]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria   Poland	A member of the family Clavatulidae; a replacement name for Clavatula auingeri Finlay (1927).
"Clavatula" nana [98]	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)		Spain	A member of the family Clavatulidae.
"Clavatula" pliogradata [98]	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)		Spain	A member of the family Clavatulidae.
Clavatula sorini [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Romania   Italy?	A species of Clavatula .
Clithon cooncreekense [99]	Sp. nov	Valid	Garvie & Symonds	Late Cretaceous (Maastrichtian)	Coon Creek Formation	United States (  Tennessee)	A species of Clithon .
Clithon duomillias [99]	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation	United States (  Texas)	A species of Clithon.
Clithon lisae [99]	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation	United States (  Texas)	A species of Clithon.
Clithon traceyi [99]	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation	United States (  Texas)	A species of Clithon.
Columbellaria rara [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Colombellinidae.
Conotomaria lecatae [101]	Sp. nov	Valid	Chaix & Grenier	Late Cretaceous (Campanian)		France	A member of the family Pleurotomariidae.
Conus damianakisi [102]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece	A species of Conus .
Conus davolii [102]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece   Italy   Hungary?	A species of Conus.
Conus dellabellai [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Conus.
Conus ictini [102]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Serravallian and Tortonian)		Greece   Turkey	A species of Conus.
Conus lauriatragei [102]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Langhian to Tortonian)		Greece   Hungary	A species of Conus.
Conus moissettei [102]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece	A species of Conus.
Conus villalvernensis [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Conus.
Coptocheilus kachinensis [103]	Sp. nov		Yu, Zhuo & Páll-Gergely	Cretaceous	Burmese amber	Myanmar	A species of Coptocheilus .
Cornirostra anistratenkorum umbilicaris [82]	Ssp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [82]
Crepidula bellardii [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Crepidula .
Cretadiostoma [103]	Gen. et 2 sp. nov		Yu, Zhuo & Páll-Gergely	Cretaceous	Burmese amber	Myanmar	A member of the family Pupinidae. Genus includes new species C. caperatum and C. umbilicarinatum.
Cryptoptyxis? fortiter [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Maoraxidae.
Cyrillia clathrataeformis [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Cyrillia .
Cyrillia saldubensis [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Cyrillia.
Dentitriangularum [104]	Gen. et sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)		France	A member of the family Cypraeidae. The type species is D. pacaudi.
Diaugasma mediterranea [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Diaugasma .
Dictyotomaria turrisbabel [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Caddo Creek Formation	United States (  Texas)	A member of the family Phymatopleuridae.
Diempterus? multicostatus [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Aporrhaidae.
Edrozeba [82]	Gen. et 4 sp. et comb. nov	Valid	Guzhov	Miocene		Russia	A member of the family Hydrobiidae. The type species is E. caeca; genus also includes new species E. angulata, E. minuta and E. striata, as well as "Hydrobia" enikalensis Kolesnikov (1935). Published online in 2023, but the issue date is listed as December 2022. [82]
Eirlysella [94]	Gen. et sp. et comb. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States (  Oklahoma)	A member of the family Luciellidae. The type species is E. buckhornensis; genus also includes "Trochus" hissingerianus de Koninck (1843), "Pleurotomaria" squamula Phillips (1836) and "Luciellina" ocultabanda Kues & Batten (2001).
Endoplocus acutus [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of the family Nerineidae.
Endoplocus inflatus [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of the family Nerineidae.
Ericusa ngayawang [105]	Sp. nov	Valid	Yates	Miocene (Langhian)	Cadell Formation	Australia	A species of Ericusa .
Exelissa ursannella [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of Cerithioidea belonging to the family Cryptaulacidae.
Faureicerithium [104]	Gen. et sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)		France	A member of the family Campanilidae. The type species is F. bidentata.
Favartia plioelata [106]	Sp. nov	Valid	Brunetti & Della Bella	Pliocene		Italy	A species of Favartia .
Granulatocincta [97]	Gen. et comb. et 3 sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene and Pliocene		Austria   Bulgaria   Czech Republic   France   Hungary   Italy   Romania   Slovakia   Spain   Turkey	A member of the family Clavatulidae. The type species is "Pleurotoma" granulato-cincta Münster in Goldfuss (1841); genus also includes new species G. callim, G. theoderichi and G. pelliscrocodili, as well as "Pleurotoma (Clavatula)" angelae Hoernes & Auinger (1891), "Clavatula" contorta Švagrovský (1958), "Pleurotoma (Clavatula)" nataliae Hoernes & Auinger (1891), "Pleurotoma" schreibersi Hörnes (1854), "Clavatula" calcarai Bellardi (1877), "Clavatula" curionii Bellardi (1877), "Clavatula" pellegrinii Simonelli (1896), "Pleurotoma" sotterii Michelotti (1847), "Clavatula" turbinata Bellardi (1877), "Clavatula" turgidula Bellardi (1877), "Clavatula" vigolensis Bellardi (1877), "Clavatula" rugata Bellardi (1877), "Murex" rusticus Brocchi (1814), "Pleurotoma" capgrandi Tournouër (1873), "Pleurotoma" obtruta Millet (1865), "Clavatula" delgadoi Vera-Peláez & Lozano-Francisco (2001) and possibly "Clavatula" boreointerrupta Kautsky (1925).
Hologyra callosa [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of the family Neritopsidae.
Hydrobia neofrauenfeldi [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	A species of Hydrobia . Published online in 2023, but the issue date is listed as December 2022. [82]
Hydrobia orientalis [107]	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation	China	A species of Hydrobia.
Iberellus colladoi [108]	Sp. nov	Valid	Juárez-Ruiz & Altaba	Pliocene		Spain	A species of Iberellus .
Itieroptygmatis cylindrata [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of Nerineoidea belonging to the family Itieriidae.
Jenneria sandrae [109]	Sp. nov	Valid	Daughenbaugh	Pleistocene (Chibanian)	Probably Bermont Formation	United States (  Florida)	A species of Jenneria .
Leufroyia gradata [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Leufroyia .
Loriolotrema? nodosa [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Brachytrematidae.
Lyria estotiensis [110]	Sp. nov	Valid	Bail, Ledon & Pacaud	Oligocene (Chattian)		France	A species of Lyria .
Lyria lozoueti [110]	Sp. nov	Valid	Bail, Ledon & Pacaud	Miocene (Aquitanian)		France	A species of Lyria.
Maoristylus pliocenicus [89]	Sp. nov	Valid	Brook & Hayward	Pliocene (Waipipian)	Kaawa Formation	New Zealand	A member of the family Bothriembryontidae belonging to the subfamily Placostylinae.
Megaclavatula [97]	Gen. et 2 sp. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene		Austria   Czech Republic   Hungary   Poland   Romania   Slovakia   Slovenia   Ukraine   Italy?	A member of the family Clavatulidae. The type species is M. grunerti; genus also includes new species M. pilleri, as well as "Pleurotoma (Clavatula)" amaliae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" antoniae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" evae Hoernes & Auinger (1891), "Pleurotoma" laevigatum Eichwald (1830), "Perrona" nemethi Kovács & Vicián (2021), "Pleurotoma" neudorfensis Schaffer (1898), "Pleurotoma tuberculosa" polonica Pusch (1837), "Pleurotoma (Clavatula) asperulata" var. subsculpta Schaffer (1912), "Pleurotoma" ernesti Toula (1901), "Pleurotoma" asperulata Lamarck (1822), Clavatula saubrigiana laurensii Peyrot (1931) and possibly "Pleurotoma (Clavatula)" szontaghi Strausz (1926), "Clavatula" taurofusulata Sacco (1890), "Clavatula" subdepressa Sacco (1890) and "Clavatula" tortonica Peyrot (1931).
Metacerithium boshuae [83]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	A member of the family Metacerithiidae.
Metalycaeus fossilis [111]	Sp. nov	In press	Yu, Páll-Gergely & Salvador	Miocene	Zhangpu amber	China	A species of Metalycaeus .
Microandonia [96]	Gen. et sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A member of the family Raphitomidae. The type species is M. minutissima.
Microdrillia plioserratula [112]	Sp. nov	Valid	Landau & Harzhauser	Pliocene		Italy   Spain	A species of Microdrillia .
Microschiza rhaetica [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of the family Purpurinidae.
Mitromorpha (Mitrolumna) velerinensis [112]	Sp. nov	Valid	Landau & Harzhauser	Pliocene		Spain   Italy?	A species of Mitromorpha .
Muricopsis landaui [113]	Sp. nov	Valid	Merle in Merle, Garrigues & Pointier	Miocene		Turkey	A species of Muricopsis .
Napaeus guanche [114]	Sp. nov	In press	Miller et al.	Middle Pleistocene		Spain (  Canary Islands)	A species of Napaeus .
Napaeus lipauges [114]	Sp. nov	In press	Miller et al.	Middle Pleistocene		Spain (  Canary Islands)	A species of Napaeus.
Narrimania saldubensis [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain
Neoperrona [97]	Gen. et comb. et sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)		Croatia   Hungary   Italy	A member of the family Clavatulidae. The type species is "Perrona" harzhauseri Kovács & Vicián (2021); genus also includes new species N. zoltanorum, as well as "Clavatula" taurinensis Bellardi (1877).
Neridomus mccallae [99]	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Edwards Formation	United States (  Texas)	A member of the family Neridomidae.
Neridomus minutissimus [99]	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Walnut Formation	United States (  Texas)	A member of the family Neridomidae.
Nerinea donosa [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of the family Nerineidae.
Neuburgensia [100]	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian to Tithonian)		Germany    Switzerland	Possibly a member of Cerithioidea. The type species is "Gymnocerithium" concavum Janicke (1966); genus also includes "Cerithium" amabile Zittel (1873), "Gymnocerithium"? convexoconcavum Gründel, Keupp & Lang (2019), "Cerithium" crenato-cinctum Zittel (1873), "Cerithium" involvens Zittel (1873) and "Cerithium" perrotundum Cossmann (1913).
Nystia conoidalis [116]	Sp. nov	Valid	Kadolsky & Morton	Oligocene (Rupelian)		United Kingdom	A species of Nystia .
Odetta chirlii [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Odetta .
Odostomia caucasica [82]	Sp. nov	Valid	Guzhov	Miocene		Russia	A species of Odostomia . Published online in 2023, but the issue date is listed as December 2022. [82]
Oleginina [97] [96]	Gen. et sp. et comb. nov	Valid	Harzhauser, Landau & Janssen in Landau, Harzhauser & Giannuzzi-Savelli	Miocene (Burdigalian to Tortonian)		Austria   Bosnia and Herzegovina   Bulgaria   Croatia   Czech Republic   Italy   Moldova   Poland   Romania   Slovakia   Ukraine	A member of the family Clavatulidae. The type species is O. mandici; genus also includes "Pleurotoma (Clavatula)" agathae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" brigittae Hoernes & Auinger (1891), "Pleurotoma" doderleini Hörnes (1854), "Pleurotoma (Clavatula)" dorotheae Hoernes & Auinger (1891), "Pleurotoma" rumana Simionescu & Barbu (1940), Clavatula schreibersi szokolyensis Strausz (1960), "Pleurotoma" winterlingensis Quenstedt (1884), "Clavatula" aradasi Bellardi (1877), "Clavatula" baccifera Bellardi (1877), "Clavatula" consularis Bellardi (1877) and "Clavatula" stazzanensis Bellardi (1877). The original generic name was Olegia, [97] but this name turned out to be preoccupied by Olegia Shaposhnikov (1979), necessitating the creation of a replacement name. [96]
Oligoptyxis khorasanica [117]	Sp. nov	Valid	Guzhov & Nazemi in Raisossadat et al.	Early Cretaceous (Aptian)		Iran	A member of the family Nerineidae.
Ondina curta [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina .
Ondina elongata [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina.
Ondina pseudovitrea [85]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina.
Ophieulima lobilloensis [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Ophieulima .
Otostoma cattoi [99]	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Edwards Formation	United States (  Texas)	A member of the family Otostomidae.
Ovalotrema [100]	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)		France    Switzerland	A member of the family Brachytrematidae. The type species is "Turbo" cotteausius d'Orbigny (1853); genus might also include "Turbo" greppini De Loriol in De Loriol & Koby (1895).
Palaeoleuca edeni [118]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian) to Eocene (Ypresian)		France	A member of the family Ellobiidae.
Palaeoleuca gradata [118]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian) to Eocene (Ypresian)		France	A member of the family Ellobiidae.
Palaeoleuca hulvizensis [118]	Nom. nov	Valid	Pacaud	Paleocene		United Kingdom	A member of the family Ellobiidae; a replacement name for Auricula (Conovulus) pygmaea Morris (1854).
Paleothyca [119]	Gen. et sp. nov	Valid	Forner, Domènech & Martinell	Early Cretaceous (Albian)		Spain	A member of the family Eulimidae. The type species is P. quinionarii.
Panneritina [120]	Gen. et comb. et sp. nov	In press	Salvador & Yu	Jurassic and Cretaceous		China   Myanmar	A member of the new family Neritidae. Genus includes Chinese species formerly assigned to the genus Mesoneritina ("M." crassa Pan, 1982, "M." dakangensis Pan, 1978, "M." gansuensis Yu & Guo, 1982, "M." nanshanensis Pan, 1984, "M." opima Yu, 1974, "M." pustula Pan, 1980), as well as new species P. ambrae from the Kachin amber.
Papuliscala spinosa [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Papuliscala .
Paragoniozona ornata [94]	Nom. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Lazy Bend Formation	United States (  Texas)	A member of the family Phymatopleuridae; a replacement name for Pleurotomaria aspera Girty (1934).
Paragoniozona yanceyi [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States (  Oklahoma)	A member of the family Phymatopleuridae.
Perrona grossi [97]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Florian Formation	Austria	A species of Perrona ; a replacement name for Pleurotoma (Clavatula) auingeri Hilber (1879).
Perrona ilonae [97]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Sámsonháza Formation	Hungary   Ukraine	A species of Perrona; a replacement name for Clavatula vindobonensis nodosa Csepreghy-Meznerics (1954).
Perrona koeberli [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria	A species of Perrona.
Perrona loetschi [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria	A species of Perrona.
Perrona wanzenboecki [97]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria	A species of Perrona; a replacement name for Pleurotoma concinna Handmann (1883).
Peruvispira kanchanaburiensis [121]	Sp. nov	In press	Karapunar, Nützel & Ketwetsuriya	Permian	Ratburi Group	Thailand	A member of the family Goniasmatidae.
Peruvispira oklahomaensis [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States (  Oklahoma)	A member of the family Goniasmatidae.
Phymatopleura conica [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States (  Oklahoma)	A member of the family Phymatopleuridae.
Phymatopleura girtyi [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wewoka Formation	United States (  Oklahoma)	A member of the family Phymatopleuridae.
Platyphysa nagensis [86]	Nom. nov	Valid	Pacaud	Cretaceous		India	A replacement name for Physa prinsepii elongata Hislop (1860).
Pleurotomaria cloutetsensis [93]	Sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)		France
Pleurotomella obesula [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Pleurotomella .
Pleurotomella turrita [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Pleurotomella.
Pomatiasia [82]	Gen. et comb. nov	Valid	Guzhov	Miocene		Russia	A member of the family Hydrobiidae. Genus includes "Amnicola" cyclostomoides Sinzov (1880). Published online in 2023, but the issue date is listed as December 2022. [82]
Propefavartia [113]	Gen. et comb. nov	Valid	Merle in Merle, Garrigues & Pointier	Eocene		France	A member of the family Muricidae belonging to the subfamily Muricopsinae. The type species is "Murex" distans Deshayes (1835).
Pseudamnicola roblesi [122]	Sp. nov	Valid	Albesa, López & Crespo	Early Miocene	Ribesalbes-Alcora Basin	Spain	A species of Pseudamnicola .
Pseudamnicola taodonggouensis [107]	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation	China	A species of Pseudamnicola.
Pseudodaphnella iberica [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Pseudodaphnella .
Pseudodaphnella velerinensis [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Pseudodaphnella.
Pseudodostia lisbonensis [99]	Sp. nov	Valid	Garvie & Symonds	Eocene	Lisbon Formation	United States (  Alabama)	A member of the family Neritidae.
Pseudomelania yamadai [83]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Pseudonerinea ? pseudomelaniformis [95]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)		Germany	A member of Nerineoidea belonging to the family Pseudonerineidae.
Purpuripullina [117]	Gen. et comb. nov	Valid	Guzhov & Raisossadat in Raisossadat et al.	Cretaceous		Iran   Lebanon	The type species is "Natica" scalaris Conrad (1852).
Purpuroidea moosleitneri [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of Caenogastropoda belonging to the family Purpuroideidae.
Pusionella hofmanni [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Romania	A species of Pusionella .
Ringicula costata chokrakensis [123]	Ssp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [123]
Ringicula costata paratethica [123]	Ssp. nov	Valid	Guzhov	Miocene		Russia	Published online in 2023, but the issue date is listed as December 2022. [123]
Ringicula subglobosa [123]	Sp. nov	Valid	Guzhov	Miocene		Russia	A species of Ringicula . Published online in 2023, but the issue date is listed as December 2022. [123]
Saalensia lorioli [100]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Brachytrematidae.
Shansiella (Oklahomaella) globilineata [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States (  Oklahoma)	A member of the family Portlockiellidae.
Sigmesalia anatole [124]	Sp. nov	Valid	Hoşgör & Pacaud	Paleocene (Danian)	Çaldağ Formation	Turkey	A member of the family Turritellidae.
Sigmesalia erifensis [124]	Sp. nov	Valid	Hoşgör & Pacaud	Paleocene (Danian)	Çaldağ Formation	Turkey	A member of the family Turritellidae.
Skenea gofasi [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Skenea .
Specula estebbunensis [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Specula .
Spinatus ruizi [125]	Sp. nov	Valid	Liverani & Wieneke	Miocene (Langhian)		Indonesia	A member of the family Strombidae.
Spiroscala quasipulchra [94]	Nom. nov	Valid	Karapunar et al.	Permian		United States	A member of the family Eotomariidae; a replacement name for Euconospira pulchra Batten (1958).
Spiroscala shwedagoniformis [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States (  Oklahoma)	A member of the family Eotomariidae.
Striopusionella [97]	Gen. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)		Hungary	A member of the family Clavatulidae. The type species is "Clavatula" szekelyhidiae Kovács & Vicián (2021).
Stuoraxis kasei [83]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	A member of the family Stuoraxidae.
Stuorella zapfei [84]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation	Austria	A member of Vetigastropoda belonging to the family Stuorellidae.
Subpterynotus eocaenicus [126]	Sp. nov		Merle, Lesport & Ledon	Eocene (Bartonian)		France	A species of Subpterynotus .
Subpterynotus mainotensis [126]	Sp. nov		Merle, Lesport & Ledon	Miocene (Aquitanian)		France	A species of Subpterynotus.
Talassia plioalboranensis [115]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Talassia .
Teretia inflatissima [96]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)		Spain	A species of Teretia .
Thalassonerita hagai [60]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation	Philippines	A member of the family Phenacolepadidae.
Tomellana aueri [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria   Bulgaria   Hungary   Romania   Slovakia   Slovenia	A species of Tomellana .
Tomellana dulaii [97]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Hungary   Romania	A species of Tomellana.
Tomellana onubensis [98]	Sp. nov	Valid	Landau & Harzhauser	Early Pliocene		Spain	A species of Tomellana.
Tomellana postjouannetii [98]	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)		Spain	A species of Tomellana.
Tortaxis zhangpuensis [111]	Sp. nov	In press	Yu, Páll-Gergely & Salvador	Miocene	Zhangpu amber	China
Tosapusia peregrina [127]	Sp. nov	Valid	Bertaccini et al.	Pliocene		Italy	A species of Tosapusia .
Trigonostoma ariejansseni [128]	Sp. nov	Valid	Roosen	Miocene		Germany	A species of Trigonostoma .
Tudorella lenneae [129]	Sp. nov	Valid	Groh et al.	Pliocene		Spain (  Canary Islands)	A species of Tudorella .
Tudorella rothei [129]	Sp. nov	Valid	Groh et al.	Pliocene		Spain (  Canary Islands)	A species of Tudorella.
Turbonilla yadongensis [130]	Sp. nov		Li, Li & Garvie	Eocene (Ypresian)	Zhupure Formation	China	A species of Turbonilla .
Valvata complanusa [107]	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation	China	A species of Valvata .
Varicoturbo [131]	Gen. et sp. nov	Valid	Nützel, Schweigert & Karapunar	Late Triassic (Carnian)	San Cassiano Formation	Italy	A member of the family Seguenziidae. Genus includes new species V. microstriatus.
Worthenia (Yochelsonospira) kuesi [94]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wetumka Formation	United States (  Oklahoma)	A species of Worthenia .
Xinjiangospira habita [107]	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation	China	A member of the family Hydrobiidae.

Gastropod research

• A study on transitions from water to land throughout the evolutionary history of gastropods is published by Vermeij & Watson-Zink (2022). ^[132]
• A study on changes in gastropod diversity at the genus/subgenus level from the Norian to the Pliensbachian is published by Ferrari & Hautmann (2022), who report that gastropods lost 56% of genera/subgenera during the Triassic–Jurassic extinction event, which was much more than the average loss of marine life at the time, and attempt to determine potential causes of the end-Triassic extinctions of gastropods. ^[133]
• A study aiming to determine the drivers of diversification for European freshwater gastropods over the past 100 million years is published by Neubauer et al. (2022). ^[134]
• A study aiming to determine the factors which caused the diversification of European freshwater gastropods in the Late Cretaceous is published by Neubauer & Harzhauser (2022). ^[135]
• Revision of the fossil material of Ancistrolepis from the Eocene (Ypresian) Llajas Formation (California, United States), representing the oldest occurrence of this genus reported to date, is published by Squires (2022). ^[136]

Other molluscs

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Calloplax roederi [137]	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene		United States (  California)	A chiton.
Chiton solaris [137]	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene		United States (  California)	A chiton.
Dorispira avannga [138]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira? lemdadensis [139]	Sp. nov		Mghazli et al.	Cambrian		Morocco
Dorispira septentrionalis [138]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira tavsenensis [138]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira tippik [138]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Erugoconus [138]	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A member of Helcionelloida of uncertain affinities. The type species is E. acuminatus.
Hoarepora [140]	Nom. nov	Valid	Mapes in Mazaev	Carboniferous (Pennsylvanian)		United States	A rostroconch; a replacement name for Oxyprora Hoare, Mapes & Yancey (2002).
Ischnochiton loureiroi [141]	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin	Portugal	A chiton, a species of Ischnochiton .
Lepidochitona rochae [141]	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin	Portugal	A chiton, a species of Lepidochitona .
Scenella? siku [138]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A member of Helcionelloida. Originally tentatively assigned to the genus Scenella , but subsequently made the type species of the separate genus Hensoniconus. [142]
Sermeqiconus [138]	Gen. et comb. nov	Valid	Peel & Kouchinsky	Cambrian	Aftenstjernesø Formation	Greenland	A helcionellid. The type species is "Figurina" polaris Peel (2021).
Tavseniconus [138]	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A member of Helcionelloida of uncertain affinities. The type species is T. erectus.
Vendrascospira [138]	Gen. et 2 sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid. The type species is V. troelseni; genus also includes V. frykmani.
Yochelcionella gracilituba [139]	Sp. nov		Mghazli et al.	Cambrian		Morocco
Yochelcionella longituba [139]	Sp. nov		Mghazli et al.	Cambrian		Morocco

Other mollusc research

• Revision of the oldest known specimens of Jinonicella kolebabai from the Middle Ordovician strata of Ukraine and Belarus is published by Gubanov et al. (2022), who report the preservation of growth lines which were previously documented only in Silurian specimens. ^[143]
• New information on the anatomy of Typhloesus wellsi is presented by Conway Morris & Caron (2022), who consider it plausible that Typhloesus was a pelagic mollusc with possible affinities with gastropods. ^[144]

References

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