2021 in paleoichthyology

Last updated

This list of fossil fish research presented in 2021 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2021.

Contents

Jawless vertebrates

NameNoveltyStatusAuthorsAgeType localityLocationNotesImages

Falxcornus [1]

Gen. et sp. nov

In press

Meng & Gai

Devonian (Lochkovian)

Xishancun Formation

Flag of the People's Republic of China.svg China

A member of Galeaspida belonging to the family Tridensaspidae. Genus includes new species F. liui.

Hongshanaspis [2]

Gen. et sp. nov

Valid

Liu et al.

Silurian (Telychian)

Qingshui Formation

Flag of the People's Republic of China.svg China

A member of Galeaspida belonging to the family Hanyangaspidae. Genus includes new species H. inexpectatus.

Jiangxialepis [3]

Gen. et sp. nov

Valid

Liu et al.

Silurian (Telychian)

Fentou Formation

Flag of the People's Republic of China.svg China

A member of Galeaspida belonging to the group Eugaleaspidiformes. The type species is J. retrospina.

Qushiaspis [4]

Gen. et sp. nov

Valid

Jiang et al.

Early Devonian

Xujiachong Formation

Flag of the People's Republic of China.svg China

A member of Galeaspida. Genus includes new species Q. elaia.

Jawless fishes research

Placoderms

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Bianchengichthys [13]

Gen. et sp. nov

Valid

Li, Zhu & Zhu in Li et al.

Silurian (Ludfordian)

Xiaoxi Formation

Flag of the People's Republic of China.svg China

A placoderm closely related to the last common ancestor of bony and cartilaginous fishes. The type species is B. micros.

Bianchengichthys.png

Leptodontichthys [14]

Gen. et sp. nov

Jobbins et al.

Devonian (Givetian)

Taboumakhlouf Formation

Flag of Morocco.svg Morocco

A member of Arthrodira belonging to the family Plourdosteidae. The type species is L. ziregensis.

Placoderm research

Acanthodians

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Dobunnacanthus [18]

Gen. et comb. nov

Valid

Dearden et al.

Devonian

Flag of the United Kingdom.svg United Kingdom

A new genus for "Vernicomacanthus" waynensis Miles (1973)

Nostolepis digitus [19]

Sp. nov

Valid

Li et al.

Devonian (Lochkovian)

Xitun Formation

Flag of the People's Republic of China.svg China

Nostolepis qujingensis [19]

Sp. nov

Valid

Li et al.

Devonian (Lochkovian)

Xitun Formation

Flag of the People's Republic of China.svg China

Acanthodian research

Cartilaginous fishes

NameNoveltyStatusAuthorsAgeType localityLocationNotesImages

Aquilolamna [22]

Gen. et sp. nov

Vullo et al.

Late Cretaceous (Turonian)

Agua Nueva Formation

Flag of Mexico.svg Mexico

A probable planktivorous shark placed in the new family Aquilolamnidae, of uncertain placement. Possibly a member of Lamniformes. The type species is A. milarcae.

Aquilolamna milarcae restoration.jpg

Carcharhinus dudoni [23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

Flag of France.svg France

A species of Carcharhinus .

Carcharhinus pedronii [23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

Flag of France.svg France

A species of Carcharhinus.

Carcharhinus tingae [24]

Sp. nov

Valid

Cicimurri & Ebersole

Eocene (Bartonian)

Flag of the United States.svg United States
(Flag of Louisiana.svg Louisiana)

A species of Carcharhinus.

Cladodus gailensis [25]

Sp. nov

Valid

Feichtinger et al.

Carboniferous (Serpukhovian)

Flag of Austria.svg Austria

Dracopristis [26]

Gen. et sp. nov

Valid

Hodnett et al.

Late Carboniferous (Kasimovian)

Atrasado Formation

Flag of the United States.svg United States
(Flag of New Mexico.svg New Mexico)

A medium-sized ctenacanthiform shark known from a complete skeleton with soft tissue. The type species is D. hoffmanorum.

Dracopristis hoffmanorum.png

Durnonovariaodus [27]

Gen. et sp. nov

Valid

Stumpf et al.

Late Jurassic (Tithonian)

Kimmeridge Clay

Flag of the United Kingdom.svg United Kingdom

A member of the family Hybodontidae. The type species is D. maiseyi.

Favusodus [28]

Gen. et sp. nov

Valid

Li et al.

Triassic (LadinianCarnian)

Flag of the People's Republic of China.svg China

A member of Euselachii. Genus includes new species F. orientalis.

Junggarensis [29]

Gen. et sp. nov

Valid

Roelofs et al.

Devonian (Famennian)

Flag of Mongolia.svg Mongolia

Genus includes new species J. ambiguus.

Keichouodus [28]

Gen. et sp. nov

Valid

Li et al.

Triassic (Ladinian–Carnian)

Flag of the People's Republic of China.svg China

A member of Euselachii. Genus includes new species K. nimaiguensis.

Keuperodus [30]

Gen. et comb. nov

Valid

Ivanov, Duffin & Richter

Late Triassic (Carnian)

Arden Sandstone Formation
Grabfeld Formation

Flag of Germany.svg Germany
Flag of the United Kingdom.svg United Kingdom

A member of the family Jalodontidae. Genus includes "Phoebodus" brodiei Woodward (1893) (interpreted by Ivanov, Duffin & Richter, 2021 as a senior synonym of "Phoebodus" keuperinus Seilacher, 1948).

Lilamna [31]

Gen. et comb. nov

Valid

Greenfield

late Eocene or Cretaceous (uncertain)

Flag of the People's Republic of China.svg China

A possible member of the family Pseudoscapanorhynchidae. The type species is "Archaeolamna" apophysata Li (1997).

Lilamna tooth.png

Maiseyacanthus [32]

Gen. et sp. nov

Valid

Bronson

Carboniferous (Mississippian)

Fayetteville Shale

Flag of the United States.svg United States
(Flag of Arkansas.svg Arkansas)

A symmoriiform. The type species is M. ozarkanum.

Maiseyodus [33]

Gen. et comb. nov

Valid

Long et al.

Devonian (Emsian)

Cravens Peak Beds

Flag of Australia (converted).svg Australia

A member of the family Mcmurdodontidae; a new genus for "Mcmurdodus" whitei Turner & Young (1987).

Moskovirhynchus [34]

Gen. et sp. nov

Valid

Popov & Shapovalov

Late Jurassic

Flag of Russia.svg Russia

A chimaera belonging to the family Callorhinchidae. Genus includes new species M. robustus.

Nebriimimus [35]

Gen. et sp. nov

Valid

Collareta et al.

Pliocene (Zanclean)

Flag of Italy.svg Italy
Flag of Spain.svg Spain [36]

A member of Rajiformes, possibly a skate. The type species is N. wardi.

Negaprion cossardi [23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

Flag of France.svg France

A species of Negaprion .

Phoebodus curvatus [37]

Sp. nov

Valid

Ivanov

Devonian (GivetianFrasnian)

Flag of Australia (converted).svg Australia
Flag of Poland.svg Poland
Flag of Russia.svg Russia

Protochimaera [38]

Gen. et sp. nov

Valid

Lebedev & Popov in Lebedev et al.

Carboniferous (ViséanSerpukhovian)

Dashkovo Formation

Flag of Russia.svg Russia
(Flag of Moscow oblast.svg Moscow Oblast)

A chimaera. Genus includes new species P. mirabilis.

Pseudocorax kindlimanni [39]

Sp. nov

In press

Jambura, Stumpf & Kriwet

Late Cretaceous (Cenomanian)

Sannine Formation

Flag of Lebanon.svg Lebanon

Ptychodus maghrebianus [40]

Sp. nov

In press

Amadori et al.

Late Cretaceous (Turonian)

Flag of Morocco.svg Morocco

Reifella [41]

Gen. et sp. nov

Valid

Ivanov in Ivanov et al.

Permian (Roadian)

Cutoff Formation

Flag of the United States.svg United States
(Flag of Texas.svg Texas)

A member of the family Anachronistidae. Genus includes new species R. lata.

Rosaodus [28]

Gen. et sp. nov

Valid

Li et al.

Triassic (Ladinian–Carnian)

Flag of the People's Republic of China.svg China

A member of Elasmobranchii of uncertain phylogenetic placement. Genus includes new species R. xingyiensis.

Strophodus indicus [42]

Sp. nov

Valid

Sharma & Singh

Middle Jurassic (Bathonian)

Jaisalmer Formation

Flag of India.svg India

Marine hybodont shark

Strophodus jaisalmerensis [43]

Sp. nov

In press

Kumar et al.

Jurassic

Jaisalmer Formation

Flag of India.svg India

A hybodont shark.

Taeniurops sergiomorai [44]

Sp. nov

Laurito & Valerio

Miocene–Pliocene (MessinianPiacenzian)

Uscari Formation

Flag of Costa Rica.svg Costa Rica

A stingray, a species of Taeniurops .

Triodus aeduorum [45]

Sp. nov

Valid

Luccisano et al.

Early Permian

Autun Basin

Flag of France.svg France

Vallisodus [46]

Nom. nov

Valid

Duffin

Late Triassic

Penarth Group

Flag of the United Kingdom.svg United Kingdom

A member of Neoselachii; a replacement name for Vallisia Duffin (1982).

Cartilaginous fish research

Ray-finned fishes

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Achirus australis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Achirus .

Achirus chungkuz [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Achirus.

Agonopsis cume [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Agonopsis .

Agoultpycnodus [69]

Gen. et comb. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian-Turonian)

Akrabou Formation

Flag of Morocco.svg Morocco

A member of the family Pycnodontidae. The type species is A. aldrovandii.

Alilepis texasensis [41]

Sp. nov

Valid

Bakaev in Ivanov et al.

Permian (Roadian)

Cutoff Formation

Flag of the United States.svg United States
(Flag of Texas.svg Texas)

A member of the family Elonichthyidae.

Anomoeodus aegypticus [70]

Sp. nov

Valid

Capasso et al.

Late Cretaceous (Maastrichtian)

Dakhla Formation

Flag of Egypt.svg Egypt

A member of Pycnodontiformes.

Anomoeodus caddoi [71]

Sp. nov

Valid

Suarez et al.

Early Cretaceous (Albian)

Holly Creek Formation

Flag of the United States.svg United States
(Flag of Arkansas.svg Arkansas)

A member of Pycnodontiformes.

Aphanolebias bettinae [72]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

Flag of Serbia.svg Serbia

A member of the family Valenciidae.

Apuliadercetis gonzalezae [73]

Sp. nov

In press

Díaz-Cruz, Alvarado-Ortega & Cantalice

Late Cretaceous (Campanian)

Angostura Formation

Flag of Mexico.svg Mexico

A member of Aulopiformes belonging to the family Dercetidae.

Archaemacruroides vanknippenbergi [74]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

Flag of Belgium (civil).svg Belgium
Flag of the Netherlands.svg Netherlands

A member of Gadiformes of uncertain phylogenetic placement.

Ariosoma mesohellenica [75]

Sp. nov

Valid

Agiadi, Koskeridou & Thivaiou

Miocene (Aquitanian)

Flag of Greece.svg Greece

A species of Ariosoma .

Austelliscus [76]

Gen. et sp. nov

Figueroa, Weinschütz & Friedman

Middle Devonian or older

Paraná Basin

Flag of Brazil.svg Brazil

An early ray-finned fish. Genus includes new species A. ferox.

Auxis koreanus [77]

Sp. nov

Valid

Nam, Nazarkin & Bannikov

Middle Miocene

Duho Formation

Flag of South Korea.svg South Korea

A species of Auxis .

Bardackichthys [78]

Gen. et sp. nov

In press

Hacker & Shimada

Late Cretaceous (Cenomanian)

Woodbine Formation

Flag of the United States.svg United States
(Flag of Texas.svg Texas)

A member of Ichthyodectiformes. Genus includes new species B. carteri.

Bobbitichthys [79]

Gen. et comb. nov

Valid

Schwarzhans, Milàn & Carnevale

Paleocene (Selandian)

Kerteminde Marl

Flag of Denmark.svg Denmark

A member of the family Macrouridae. The type species is "Hymenocephalus" rosenkrantzi Schwarzhans (2003).

Brauccipycnodus [80]

Gen. et comb. nov

Valid

Taverne & Capasso

Early Cretaceous (Albian)

Flag of Italy.svg Italy

A member of the family Pycnodontidae. The type species is "Proscinetes" pillae Capasso (2007).

Briveichthys [81]

Gen. et sp. nov

Valid

Štamberg & Steyer

Permian

Brive Basin

Flag of France.svg France

A pygopterid. Genus includes new species B. chantepieorum.

Capromimus undulatus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Capromimus .

Cheirolepis jonesi [82]

Sp. nov

In press

Newman et al.

Devonian (Givetian)

Tordalen Formation

Flag of Norway.svg Norway

Chiloconger chilensis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Chiloconger .

Choichix [83]

Gen. et sp. nov

Valid

Cantalice, Than-Marchese & Villalobos-Segura

Late Cretaceous (Cenomanian)

Flag of Mexico.svg Mexico

A member of Acanthopterygii of uncertain phylogenetic placement. Genus includes new species C. alvaradoi.

Citharichthys parvisulcus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds
Lacui Formation
Navidad Formation
Ranquil Formation

Flag of Chile.svg Chile

A species of Citharichthys .

Citharichthys vergens [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Citharichthys.

Clarotes eocenicus [84]

Sp. nov

Valid

Murray & Holmes

Late Eocene

Jebel Qatrani Formation

Flag of Egypt.svg Egypt

A species of Clarotes .

Coccolepis solnhofensis [85]

Sp. nov

Valid

López-Arbarello & Ebert

Late Jurassic (Tithonian)

Altmühltal Formation

Flag of Germany.svg Germany

A member of Chondrostei belonging to the family Coccolepididae.

Coelorinchus fidelis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún Beds
Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Coelorinchus .

Coelorinchus rapelanus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Coelorinchus.

Cottunculus primaevus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Cottunculus .

Cretaserranus [74]

Gen. et sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

Flag of Belgium (civil).svg Belgium
Flag of the Netherlands.svg Netherlands

A member of Perciformes, possibly belonging to the family Serranidae. Genus includes new species C. maastrichtensis.

Ctenopharyngodon orientalis [86]

Sp. nov

Valid

Su, Chang & Chen

Miocene

Xiacaowan Formation

Flag of the People's Republic of China.svg China

A species of Ctenopharyngodon .

Ctenopharyngodon xiejiaensis [86]

Sp. nov

Valid

Su, Chang & Chen

Miocene

Xiejia Formation

Flag of the People's Republic of China.svg China

A species of Ctenopharyngodon.

Dezaoia [86]

Gen. et sp. nov

Valid

Su, Chang & Chen

Oligocene

Ulanbulage Formation

Flag of the People's Republic of China.svg China

A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is D. saintjaquensis.

Diastemapycnodus [87]

Gen. et sp. nov

In press

Abu El-Kheir et al.

Late Cretaceous (Maastrichtian)

Dakhla Formation

Flag of Egypt.svg Egypt

A member of Pycnodontiformes. Genus includes new species D. tavernensis.

Elongofuro augustilgi [88]

Sp. nov

Valid

Ebert

Late Jurassic (Kimmeridgian)

Nusplingen Limestone

Flag of Germany.svg Germany

A member of Ophiopsiformes.

Eoctenopharyngodon [86]

Gen. et sp. nov

Valid

Su, Chang & Chen

Oligocene

Dongying Formation

Flag of the People's Republic of China.svg China

A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is E. liui.

Eotexachara [89]

Gen. et sp. nov

In press

Wick

Late Cretaceous (Campanian)

Aguja Formation

Flag of the United States.svg United States
(Flag of Texas.svg Texas)

A member of Characiformes. Genus includes new species E. malateres.

Feroxichthys panzhouensis [90]

Sp. nov

Valid

Ma, Xu & Geng

Middle Triassic (Anisian)

Guanling Formation

Flag of the People's Republic of China.svg China

A member of the family Colobodontidae.

Garganomyctophum [91]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Santonian)

Flag of Italy.svg Italy

A lanternfish. The type species is G. sorbinii.

Gnathophis elongatus [75]

Sp. nov

Valid

Agiadi, Koskeridou & Thivaiou

Miocene (Aquitanian)

Flag of Greece.svg Greece

A species of Gnathophis .

Gnathophis quinzioi [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Gnathophis.

Gobiosoma? axsmithi [92]

Sp. nov

Valid

Ebersole, Cicimurri & Stringer

Oligocene (Rupelian)

Byram Formation

Flag of the United States.svg United States
(Flag of Alabama.svg Alabama)

A member of the family Gobiidae.

Guiclupea [93]

Gen. et sp. nov

Valid

Chen et al.

Oligocene

Flag of the People's Republic of China.svg China

A member of Clupeomorpha belonging to the group Ellimmichthyiformes. The type species is G. superstes.

Hellenigobius [94]

Gen. et sp. et comb. nov

Valid

Schwarzhans, Agiadi & Thivaiou

Miocene

Flag of the Czech Republic.svg Czech Republic
Flag of Greece.svg Greece
Flag of Italy.svg Italy
Flag of Kazakhstan.svg Kazakhstan
Flag of Serbia.svg Serbia
Flag of Hungary.svg Hungary?

A member of the subfamily Gobionellinae. The type species is H. praeschismatus; genus also includes "Pomatoschistus" bunyatovi Bratishko et al. (2015).

Katyagobius [95]

Gen. et sp. nov

Valid

Reichenbacher & Bannikov

Miocene

Flag of Moldova.svg Moldova

A member of the family Gobiidae. The type species is K. prikryli.

Kelliaichthys [96]

Nom. nov

Valid

Schultze in Schultze et al.

Flag of Kazakhstan.svg Kazakhstan

A replacement name for Kellia Kazantseva-Selezneva (1980).

Klincigobus haraldahnelti [72]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

Flag of Serbia.svg Serbia

A member of the family Gobiidae.

Krumvirichthys [97]

Gen. et sp. nov

Valid

Přikryl

Oligocene–early Miocene

Flag of the Czech Republic.svg Czech Republic

A deep-sea smelt. The type species is K. brzobohatyi.

Kuhlia orientalis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A flagtail.

Lampanyctus ipunensis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds

Flag of Chile.svg Chile
Flag of New Zealand.svg New Zealand

A species of Lampanyctus .

Larimichthys koae [98]

Sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

Flag of the Republic of China.svg Taiwan

A species of Larimichthys .

Lepophidium chonorum [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Lepophidium .

Lepophidium mapucheorum [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Lepophidium.

Maurolicus brevirostris [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Maurolicus .

Navidadichthys [68]

Gen. et sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

Possibly a member of the family Prototroctidae. The type species is N. mirus.

Neuburgichthys [96]

Nom. nov

Valid

Schultze in Schultze et al.

Flag of Kazakhstan.svg Kazakhstan

A replacement name for Neuburgia Kazantseva-Selezneva (1980).

Nezumia epuge [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

Flag of Chile.svg Chile

A species of Nezumia .

Nibea chaoi [98]

Sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

Flag of the Republic of China.svg Taiwan

A species of Nibea .

Oshiaichthys [96]

Nom. nov

Valid

Schultze in Schultze et al.

Triassic

Flag of Kyrgyzstan.svg Kyrgyzstan

A replacement name for Oshia Sytchevskaya (1999).

Palaeopantodon [99]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

Flag of Lebanon.svg Lebanon

A member of the family Pantodontidae. The type species is P. vandersypeni.

Pankowskipiscis [100]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

Flag of Lebanon.svg Lebanon

A member of the family Pantodontidae. The type species is P. haqelensis.

Paracarapus [68]

Gen. et sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A pearlfish. The type species is P. chilensis.

Peltoperleidus asiaticus [101]

Sp. nov

Valid

Xu

Middle Triassic (Anisian)

Guanling Formation

Flag of the People's Republic of China.svg China

A member of Neopterygii belonging to the group Louwoichthyiformes.

Petersichthys [102]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

Flag of Lebanon.svg Lebanon

A member of the family Pantodontidae. The type species is P. libanicus.

Physiculus pichi [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Physiculus .

Pinichthys shirvanensis [103]

Sp. nov

Valid

Bannikov

Miocene

Flag of Russia.svg Russia
(Flag of Krasnodar Krai.svg Krasnodar Krai)

A member of Perciformes belonging to the group Stromateoidei.

Plesiogobius [94]

Gen. et sp. et comb. nov

Valid

Schwarzhans, Agiadi & Thivaiou

Miocene (Aquitanian)

Flag of Greece.svg Greece

A member of the family Gobiidae belonging to the subfamily Gobiinae. The type species is P. felliensis.

Polyipnus bandeli [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Polyipnus .

Portentosoceros [104]

Gen. et comb. nov

Valid

Nazarkin & Bannikov

Miocene

Flag of Russia.svg Russia
(Flag of Sakhalin Oblast.svg Sakhalin Oblast)

A member of Percoidei of uncertain phylogenetic placement; a new genus for "Pentaceros" sakhaliniсus Gretchina (1975).

Primuluchara [89]

Gen. et sp. nov

In press

Wick

Late Cretaceous (Campanian)

Aguja Formation

Dinosaur Park Formation

Flag of the United States.svg United States
(Flag of Texas.svg Texas)

Flag of Canada (Pantone).svg Canada (Flag of Alberta.svg Alberta)

A member of Characiformes. Genus includes new species P. laramidensis.

Pseudohilsa nikosi [105]

Sp. nov

Valid

Kevrekidis, Arratia & Reichenbacher in Kevrekidis et al.

Late Miocene

Flag of Greece.svg Greece

A member of the family Clupeidae.

Pseudolesueurigobius [95]

Gen. et sp. nov

Valid

Reichenbacher & Bannikov

Miocene

Flag of Moldova.svg Moldova

A member of the family Gobiidae. The type species is P. manfredi.

Pseudonus humilis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds
Navidad Formation

Flag of Chile.svg Chile

A species of Pseudonus .

Pteronisculus changae [106]

Sp. nov

Valid

Ren & Xu

Middle Triassic (Anisian)

Guanling Formation

Flag of the People's Republic of China.svg China

Pterothrissus transpacificus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Pterothrissus .

Pythonichthys panulus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation

Flag of Chile.svg Chile

A species of Pythonichthys .

Raususetarches [107]

Gen. et sp. nov

Valid

Yabumoto & Nazarkin

Late Miocene

Koshikawa Formation

Flag of Japan.svg Japan

A member of the family Scorpaenidae. Genus includes new species R. sakurai.

Rhinocephalus cretaceus [74]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

Flag of Belgium (civil).svg Belgium

A member of the family Merlucciidae.

Rhynchoconger chiloensis [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Rhynchoconger .

Sarmatigobius [95]

Gen. et sp. et comb. nov

Valid

Reichenbacher & Bannikov

Miocene

Flag of Moldova.svg Moldova

A member of the family Gobiidae. The type species is S. compactus; genus also includes "Hesperichthys" iugosus Schwarzhans, Brzobohatý & Radwańska (2020).

Saurichthys sceltrichensis [108]

Sp. nov

Valid

Renesto, Magnani & Stockar

Middle Triassic (Ladinian)

Meride Limestone

Flag of Switzerland (Pantone).svg  Switzerland

Severnichthus [109] [110]

Gen. et sp. nov

In press

Stringer & Schwarzhans

Late Cretaceous (Maastrichtian)

Severn Formation

Flag of the United States.svg United States
(Flag of Maryland.svg Maryland)

Possibly a member of Polymixiiformes. Genus includes new species S. bourdoni.

Sirembola supersa [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation

Flag of Chile.svg Chile

A cusk-eel.

Spectrunculus sparsus [68]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

Flag of Chile.svg Chile

A species of Spectrunculus .

Taosciaena [98]

Gen. et 2 sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

Flag of the Republic of China.svg Taiwan

A member of the family Sciaenidae. The type species is T. jiangi; genus also includes T. hui.

Toxopyge vracevicensis [72]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

Flag of Serbia.svg Serbia

A member of the family Gobiidae.

Wilsonium [111]

Gen. et comb. nov

Valid

Liu

Early Eocene

Allenby Formation

Flag of Canada (Pantone).svg Canada
(Flag of British Columbia.svg British Columbia)

A Catostomidae sucker.
The type species is "Amyzon" brevipinne (1893).

Yarigobius [95]

Gen. et 2 sp. nov

Valid

Reichenbacher & Bannikov

Miocene

Flag of Moldova.svg Moldova

A member of the family Gobiidae. The type species is Y. decoratus; genus also includes Y. naslavcensis.

Ray-finned fish research

Lobe-finned fishes

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Ceratodus guanganensis [129]

Sp. nov

Valid

Wang et al.

Late Jurassic

Shaximiao Formation

Flag of the People's Republic of China.svg China

Announced in 2021; the final version of the article naming it was published in 2022.

Eusthenodon bourdoni [130]

Sp. nov

Valid

Downs, Barbosa & Daeschler

Devonian (Famennian)

Catskill Formation

Flag of the United States.svg United States
(Flag of Pennsylvania.svg Pennsylvania)

Lobe-finned fish research

General research

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