2018 in paleoentomology

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2018 in paleoentomology is a list of new fossil insect taxa that were described during the year 2018, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Contents

Newly named taxa

Coleopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Acalyptopygus [1]

Gen. et 4 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Aepyceratinae. The type species is A. brevicornis; genus also includes A. lingziae, A. elongatus and A. astriatus.

Acalyptopygus brevicornis Acalyptopygus brevicornis fig1 A.jpg
Acalyptopygus brevicornis

Allostrophus [2]

Gen. et sp. nov

Valid

Hsiao et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Tetratomidae. Genus includes new species A. cretaceus.

Amberocula [3]

Gen. et 3 sp. nov

Valid

Batelka, Engel & Prokop

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ripiphoridae. The type species is A. muelleri; genus also includes A. costata and A. fallax.

Amplectister [4]

Gen. et sp. nov

Valid

Caterino & Maddison

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Histeridae. The type species is A. tenax.

Amplectister tenax Amplectister tenax Fig4.jpg
Amplectister tenax

Aphelonyssus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is A. latus.

Apotomoura [5]

Gen. et sp. nov

Valid

Bao et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Tenebrionoidea belonging to the new family Apotomouridae. Genus includes new species A. fortiscrura.

Archaeoheilus [6]

Gen. et 2 sp. et 4 comb. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil. The type species is A. scudderi
genus also includes new species A. ovalis
Also the moved species Hylobius deleticius (2015), Hylobius packardii ( 1893 ), Hylobius provectus ( 1876 )
Additionally includes Hylobius lacoei ( 1893 ) from the Florissant Formation.

Ataenius damzeni [7]

Sp. nov

Valid

Bukejs & Alekseev

Eocene

Baltic amber

Europe (Baltic Sea coast)

A species of Ataenius .

Atomaria saxonica [8]

Sp. nov

Valid

Lyubarsky & Perkovsky

Late Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

A species of Atomaria .

Autosilis annisettaekoppelae [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Silinae.

Baltocoeliodes [10]

Gen. et sp. nov

Valid

Legalov & Bukejs

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A weevil belonging to the family Curculionidae and the tribe Ceutorhynchini. The type species is B. sontagae.

Baltocyba [11]

Gen. et sp. nov

Valid

Legalov

Eocene

Prussian Formation
(Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A member of the family Brentidae, the subfamily Apioninae and the tribe Rhadinocybini. The type species is B. electrinus.

Baltonanophyes [11]

Gen. et sp. nov

Valid

Legalov

Eocene

Prussian Formation
(Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A member of the family Brentidae, the subfamily Nanophyinae and the tribe Nanophyini. The type species is B. crassirostre.

Baltostigus striatipennis [12]

Sp. nov

Valid

Jałoszyński et al.

Late Eocene

Baltic amber

Europe (Baltic Sea coast)

A scydmaenine rove beetle belonging to the supertribe Mastigitae.

Bowangius [1]

Gen. et 4 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is B. cyclops; genus also includes B. tanaops, B. zhenuai and B. glabratus.

Burmalestes [13]

Gen. et sp. nov

Valid

Tomaszewska & Ślipiński in Tomaszewska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Endomychidae. Genus includes new species B. albertalleni.

Burmitoma [3]

Gen. et sp. nov

Valid

Batelka, Engel & Prokop

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ripiphoridae. The type species is B. nalae.

Burmocorynus [14]

Gen. et 2 sp. nov

Valid

Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

Originally described as a member of the family Belidae, [14] but subsequently transferred to the weevil family Mesophyletidae and the subfamily Mesophyletinae. [1] The type species is B. jarzembowskii Legalov (2018); genus also includes B. longus Clarke & Oberprieler in Clarke et al. (2018). [1]

Burmomacer [14]

Gen. et sp. nov

Valid

Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Nemonychidae. The type species is B. kirejtshuki.

Burmomiles [15]

Gen. et sp. nov

Valid

Fanti, Damgaard & Ellenberger

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A soldier beetle. Genus includes new species B. willerslevorum.

Burmorhinus [16]

Gen. et 2 sp. nov

Valid

Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil, originally assigned to the subfamily Erirhininae and the tribe Arthrostenini; subsequently transferred to the weevil family Mesophyletidae and the subfamily Mesophyletinae. [1] Genus includes new species B. georgei Legalov (2018) and B. setosus Clarke & Oberprieler in Clarke et al. (2018). [1]

Cacomorphocerus bentifabrici [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae.

Cacomorphocerus madseni [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae.

Cacomorphocerus wiszniewskii [17]

Sp. nov

Valid

Fanti & Kupryjanowicz

Eocene

Baltic amber

Europe (Baltic Sea coast)

A soldier beetle.

Calyptocis [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Aepyceratinae. The type species is C. brevirostris.

Calyptocis brevirostris Calyptocis brevirostris Fig1 A.jpg
Calyptocis brevirostris

Cantharis hanswerneri [18]

Sp. nov

Valid

Kazantsev

Eocene

Baltic amber

Europe (Baltic region)

A species of Cantharis .

Cantharis hoffeinsorum [18]

Sp. nov

Valid

Kazantsev

Eocene

Baltic amber

Europe (Baltic region)

A species of Cantharis .

Cantharis mikkelsenorum [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Cantharis .

Cascomastigus minor [19]

Sp. nov

Valid

Yin, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Scydmaeninae.

Cetionyx [1]

Gen. et 3 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is C. batiatus; genus also includes C. terebrans and C. ursinus.

Colon burmiticum [20]

Sp. nov

Valid

Yamamoto in Yamamoto & Takahashi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Leiodidae belonging to the subfamily Coloninae.

Compsopsarus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is C. reneae.

Cretaparamecus [13]

Gen. et sp. nov

Valid

Tomaszewska, Ślipiński, Bai & Zhang in Tomaszewska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Endomychidae. Genus includes new species C. tarsalis.

Cretohister [21]

Gen. et sp. nov

Valid

Zhou et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Histeroidea belonging to the new family Cretohisteridae. Genus includes new species C. sinensis.

Cretohypsilara [22]

Gen. et sp. nov

Valid

Cai, Maier & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A riffle beetle. Genus includes new species C. parva.

Cretolestes [13]

Gen. et sp. nov

Valid

Tomaszewska, Ślipiński & Ren in Tomaszewska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Endomychidae. Genus includes new species C. niger.

Cretoparacucujus [23]

Gen. et sp. nov

Valid

Cai & Escalona in Cai et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Boganiidae belonging to the subfamily Paracucujinae. The type species is C. cycadophilus.

Cretozenodosus [24]

Gen. et sp. nov

Valid

Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Cleroidea belonging to the family Thanerocleridae and the subfamily Zenodosinae. Genus includes new species C. fossilis.

?Cryptarcha subglobosa [25]

Sp. nov

Valid

Kirejtshuk & Nel

Paleocene

Flag of France.svg  France

Possibly a species of Cryptarcha .

Cyclaxyra cretacea [26]

Sp. nov

Valid

Wu, Li & Ding

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Cucujoidea belonging to the family Cyclaxyridae. Originally described as a species of Cyclaxyra ; Gimmel et al. (2019) made it the type species of a separate genus Electroxyra. [27]

Cyrtocis [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is C. gibbus.

Debbia [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is D. gracilirostris.

Dignomus francescovitalii [28]

Sp. nov

Valid

Bukejs, Bellés & Alekseev

Eocene

Baltic amber

Europe (Baltic Sea region)

A spider beetle.

Dinoharpalus martynovi [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Permosynidae.

Dysanabatium aenaum [30]

Sp. nov

Valid

Bogri, Solodovnikov & Żyła

Eocene

Baltic amber

Flag of Denmark.svg  Denmark

A rove beetle belonging to the subfamily Paederinae.

Dysanabatium damgaardi [30]

Sp. nov

Valid

Bogri, Solodovnikov & Żyła

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A rove beetle belonging to the subfamily Paederinae.

Dysanabatium johannesi [30]

Sp. nov

Valid

Bogri, Solodovnikov & Żyła

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A rove beetle belonging to the subfamily Paederinae.

Dysanabatium kechrimparense [30]

Sp. nov

Valid

Bogri, Solodovnikov & Żyła

Eocene

Baltic amber

Flag of Denmark.svg  Denmark
Flag of Poland.svg  Poland
Flag of Russia.svg  Russia
Flag of Ukraine.svg  Ukraine

A rove beetle belonging to the subfamily Paederinae.

Echogomphus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is E. viridescens.

Eduardoxenus [31]

Gen. et sp. nov

Valid

Legalov, Nazarenko & Perkovsky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A member of the family Anthribidae belonging to the subfamily Choraginae and the tribe Valenfriesiini. The type species is E. unicus.

Electrocis [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is E. dentitibialis.

Electrorubesopsis [32]

Gen. et sp. nov

Valid

Bai & Wang in Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Scarabaeidae. Genus includes new species E. beuteli.

Elwoodius [1]

Gen. et sp. nov

Junior homonym

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is E. conicops. The generic name is preoccupied by Elwoodius Colonnelli (2014); Clarke & Oberprieler (2019) coined a replacement name Zimmiorhinus. [33]

Erichia cretacea [34]

Sp. nov

Valid

Yu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Limnichidae.

Eridanula [9]

Gen. et sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae. The type species is E. susannaebierae.

Eudiagogus vossi [6]

Sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil, a species of Eudiagogus .

Euroleptochromus setifer [12]

Sp. nov

Valid

Jałoszyński et al.

Late Eocene

Baltic amber

Europe (Baltic Sea coast)

A scydmaenine rove beetle belonging to the supertribe Mastigitae.

Euroleptochromus tuberculatus [35]

Sp. nov

Valid

Yin & Cai

Eocene

Baltic amber

Europe (Baltic Sea region)

A scydmaenine rove beetle belonging to the supertribe Mastigitae.

Euryepomus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is E. lophomerus.

Extinctocentrinus [36]

Gen. et sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States
Flag of Colorado.svg  Colorado

An Eudiagogini tribe entimine Curculionidae weevil.
The type species is E. brevirostris.

Frankencupes [37]

Gen. et sp. nov

Valid

Ponomarenko & Bashkuev

Middle Triassic (Anisian)

Röt Formation

Flag of Germany.svg  Germany

A member of Archostemata belonging to the family Permocupedidae. Genus includes new species F. ultimus.

Furcalabratum [38]

Gen. et sp. nov

Valid

Poinar & Brown

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

Originally described as a member of the family Kateretidae; subsequently argued to be a sap beetle belonging to the subfamily Apophisandrinae [39] or a member of the separate family Apophisandridae. [40] Genus includes new species F. burmanicum.

Geraeus anvilis [36]

Sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States
Flag of Colorado.svg  Colorado

A species of Geraeus .

Geraeus fossilis [36]

Sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A species of Geraeus .

Giltine [41]

Gen. et sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of Coccinelloidea belonging to the family Anamorphidae. The type species is G. ampeensis.

Glaresis burmitica [42]

Sp. nov

Valid

Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A species of Glaresis .

Globicornis groehni [43]

Sp. nov

Valid

Bukejs & Háva

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Globicornis .

Gnomus [1]

Gen. et 2 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is G. brevis; genus also includes G. spinipes.

Gollandia [44]

Gen. et sp. nov

Valid

Makranczy, Yamamoto & Engel

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Oxytelinae, tentatively placed in the tribe Coprophilini. The type species is G. planata.

Gonodera baygushevae [45]

Sp. nov

Valid

Nabozhenko & Chigray

Eocene

Baltic amber

Europe (Baltic Sea region)

A darkling beetle belonging to the subfamily Alleculinae.

Gramboale [41]

Gen. et sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of Coccinelloidea belonging to the family Anamorphidae. The type species is G. prutenorum.

Guillermorhinus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Nemonychidae and the subfamily Rhinorhynchinae. The type species is G. longitarsis.

Guillermorhinus longitarsis Guillermorhinus longitarsis fig 15 A.jpg
Guillermorhinus longitarsis

Habropezus kimpulleni [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Habropezus ncoxatirostris [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Habropezus tenuicornis [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Hukawngichthyurus [46]

Gen. et sp. nov

Valid

Fanti & Ellenberger

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A soldier beetle. Genus includes new species H. kyawkhaingwini.

Hukawngius [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is H. crassipes.

Hydrobiites permianus [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Permosynidae.

Hymenorus campbelli [47]

Nom. nov

Valid

Bouchard in Bousquet et al.

Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A darkling beetle; replacement name for Hymenorus oculatus Doyen & Poinar (1994).

Juratelacrima [9]

Gen. et sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae. The type species is J. ballingi.

Kekveus [48]

Gen. et sp. nov

Valid

Yamamoto, Grebennikov & Takahashi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ptiliidae. Genus includes new species K. jason.

Kleinzaches [41]

Gen. et sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

A member of Coccinelloidea belonging to the family Anamorphidae. The type species is K. germanicianus.

Klesovia [49]

Gen. et sp. nov

Valid

Petrov & Perkovsky

Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A bark beetle. Genus includes new species K. pubescens.

Kuafu [50]

Gen. et sp. nov

Valid

Yin et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An ant-like stone beetle belonging to the stem group of the tribe Scydmaenini. Genus includes new species K. borealis.

Kuskaella bajerae [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Malthininae.

Leptopezus [1]

Gen. et 2 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is L. rastellipes; genus also includes L. barbatus.

Limodromus emetikos [51]

Sp. nov

Valid

Gamboa & Ortuño

Eocene

Baltic amber

Europe (Baltic Sea coast)

A ground beetle belonging to the tribe Platynini.

Lithogeraeus [36]

Gen. et sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil. The type species is L. greenriverensis.

Louwiocis [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is L. megalops.

Lycocerus christelae [18]

Sp. nov

Valid

Kazantsev

Eocene

Baltic amber

Europe (Baltic region)

A soldier beetle.

Lycocerus dentantennatus [18]

Sp. nov

Valid

Kazantsev

Eocene

Baltic amber

Europe (Baltic region)

A soldier beetle.

Malthinus rifbjergi [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Malthinus .

Malthodes aphidiphagus [52]

Sp. nov

Valid

Fanti & Michalski

Eocene

Baltic amber

Flag of Poland.svg  Poland

A species of Malthodes .

Malthodes henningseni [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Malthodes .

Malthodes meriae [53]

Sp. nov

Valid

Fanti

Eocene (Lutetian to Priabonian)

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Malthodes .

Malthodes moellehavei [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Malthodes .

Markus [54]

Gen. et sp. nov

Valid

Fanti & Pankowski

Eocene (Lutetian to Priabonian)

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Silinae and the tribe Silini. The type species is M. karenae.

Malthodes (Malthodes) josephi [55]

Sp. nov

Valid

Fanti & Pankowski

Eocene (Priabonian)

Prussian Formation
(Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Malthodes .

Mekorhamphus beatae [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Mekorhamphus gracilipes [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Mekorhamphus poinari [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Mekorhamphus tenuicornis [1]

Sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae.

Menatoraea [25]

Gen. et 4 sp. nov

Valid

Kirejtshuk & Nel

Paleocene

Flag of France.svg  France

A sap beetle. The type species is M. typica; genus also includes M. angustitibialis, M. laticollis and M. gracilis.

Microzavaljus [56]

Gen. et sp. nov

Valid

Lyubarsky & Perkovsky

Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

A member of the family Erotylidae belonging to the subfamily Xenoscelinae. The type species is M. saxonicum.

Miocenocylas [57]

Gen. et sp. nov

Valid

Legalov

Middle Miocene

Upper Freshwater-Molasse Formation

Flag of Germany.svg  Germany

A member of the family Brentidae belonging to the subfamily Cyladinae. Genus includes new species M. heeri.

Molliberus [58]

Gen. et sp. nov

Valid

Peris & Fanti

Early Cretaceous (Albian)

Flag of Spain.svg  Spain

A soldier beetle known from amber from the Cave of El Soplao. Genus includes new species M. albae.

Multispinus [5]

Gen. et sp. nov

Valid

Bao et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Tenebrionoidea belonging to the new family Apotomouridae. Genus includes new species M. multispinosus.

Myanmarus [1]

Gen. et 4 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is M. caviventris; genus also includes M. robustus, M. dentifer and M. diversiunguis.

Nicentrus curvirostris [36]

Sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A species of Nicentrus .

Nitoraeopsis [25]

Gen. et sp. nov

Valid

Kirejtshuk & Nel

Paleocene

Flag of France.svg  France

A sap beetle. Genus includes new species N. mixta.

Noergaardia [9]

Gen. et sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae. The type species is N. dinae.

Nuegua [59]

Gen. et sp. nov

Valid

Yin, Cai & Newton

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A glandulariine ant-like stone beetle. Genus includes new species N. elongata.

Nugatorhinus [1]

Gen. et 2 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Aepyceratinae. The type species is N. chenyangi; genus also includes N. albomaculatus.

Nugatorhinus chenyangi Nugatorhinus chenyangi Fig1 A.jpg
Nugatorhinus chenyangi

Ocriocis [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is O. binodosus.

Opeatorhynchus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is O. comans.

Ornatomalthinus ruicheni [60]

Sp. nov

Valid

Hsiao & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A soldier beetle. Originally described as a species of Ornatomalthinus ; Fanti (2018) transferred this species to the genus Sanaungulus. [61]

Orsunius electronefelus [62]

Sp. nov

Valid

Kypke & Solodovnikov

Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A rove beetle belonging to the subfamily Paederinae, tribe Lathrobiini and subtribe Medonina.

Oxycorynoides bucklowae [63]

Sp. nov

Valid

Legalov & Jarzembowski

Early Cretaceous (Barremian)

Upper Weald Clay Formation

Flag of the United Kingdom.svg  United Kingdom

A weevil belonging to the family Nemonychidae.

Palaecoryphus [41]

Gen. et sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of Coccinelloidea belonging to the family Anamorphidae. The type species is P. viktori.

Palaeolycra [25]

Gen. et sp. nov

Valid

Kirejtshuk & Nel

Paleocene

Flag of France.svg  France

A sap beetle. Genus includes new species P. palaeocenica.

Palaeomallerus [36]

Gen. et sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil. The type species is P. longirostris.

Palaeomycetes [13]

Gen. et sp. nov

Valid

Tomaszewska, Ślipiński & Ren in Tomaszewska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Endomychidae. Genus includes new species P. foveolatus.

Palaeotylus [64]

Gen. et sp. nov

Valid

Poinar, Vega & Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

Originally described as an ambrosia beetle; subsequently considered to be unlikely to belong to Curculionoidea by Clarke et al. (2018). [1] Genus includes new species P. femoralis.

Palmnickeneoceras [9]

Gen. et sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae. The type species is P. ejersboi.

Pangusyndicus [65]

Gen. et sp. nov

Valid

Yin, Zhou & Cai in Yin et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An ant-like stone beetle. Genus includes new species P. excavatus.

Paraodontomma szwedoi [66]

Sp. nov

Valid

Jarzembowski, Wang & Zheng

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ommatidae belonging to the tribe Brochocoleini.

Paraodontomma szwedoi Paraodontomma szwedoi.jpg
Paraodontomma szwedoi

Perapion rasnitsyni [6]

Sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil, a species of Perapion .

Periosocerus [1]

Gen. et 2 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is P. deplanatus; genus also includes P. crenulatus.

Perroudia manuherikia [67]

Sp. nov

Valid

Kaulfuss et al.

Early Miocene

Manuherikia Group

Flag of New Zealand.svg  New Zealand

A member of the family Brentidae.

Petalotarsus [1]

Gen. et 3 sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is P. oxycorynoides; genus also includes P. curculionoides and P. cylindricus.

Platychirus [1]

Gen. et sp. nov

Junior homonym

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Aepyceratinae. The type species is P. beloides. The generic name is preoccupied by Platychirus Agassiz (1846); Clarke & Oberprieler (2019) coined a replacement name Burmophyletis. [33]

Burmophyletis beloides Burmophyletis beloides holotye fig1 A.jpg
Burmophyletis beloides

Platypelochares electricus [68]

Sp. nov.

Valid

Hernando, Szawaryn & Ribera

Eocene (Lutetian)

Baltic amber

Europe (Gdańsk Bay area)

A member of the family Limnichidae. Originally described as a species of Platypelochares , but subsequently made the type species of the separate genus Hernandochares. [69]

Plesiotoma [3]

Gen. et sp. nov

Valid

Batelka, Engel & Prokop

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ripiphoridae. The type species is P. alissae.

Podistra kloevedali [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae.

Poinarinius [70]

Gen. et sp. nov

Valid

Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An auger beetle belonging to the subfamily Dinoderinae. Genus includes new species P. burmaensis.

Ponomarenkium [71]

Nom. nov

Valid

Yan et al.

Late Permian

Newcastle Coal Measures

Flag of Australia (converted).svg  Australia

A beetle of uncertain phylogenetic placement, assigned to the new family Ponomarenkiidae; a replacement name for Ponomarenkia Yan et al. (2017).

Praphennium [72]

Gen. et sp. nov

Valid

Jałoszyński

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Scydmaeninae and the tribe Cephenniini. Genus includes new species P. carinatum.

Primocentron [6]

Gen. et sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A weevil. The type species is P. wickhami.

Procleomenes gouverneuri [73]

Sp. nov

Valid

Vitali

Eocene

Baltic amber

Europe (Baltic Sea region)

A longhorn beetle.

Propiestus [74]

Gen. et sp. nov

Valid

Yamamoto, Caron & Bortoluzzi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Piestinae. Genus includes new species P. archaicus.

Protoripidius [75]

Gen. et sp. nov

Valid

Cai, Yin & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ripiphoridae and the subfamily Ripidiinae. Genus includes new species P. burmiticus.

Pseudochirotenon [6]

Gen. et sp. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

A fungus weevil.
The type species is P. eocaenicus.

Pseudochrysomelites dvinensis [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Schizocoleidae.

Pseudophaops [6]

Gen. et comb. nov

Valid

Legalov

Eocene

Green River Formation

Flag of the United States.svg  United States

An Eustylini tribe entimine weevil.
The type species is Otiorhynchus perditus (1876).

Rhadinomycter [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Mesophyletinae. The type species is R. perplexus.

Rhagonycha (Rhagonycha) maryae [55]

Sp. nov

Valid

Fanti & Pankowski

Eocene (Priabonian)

Prussian Formation
(Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Rhagonycha .

Rhagonycha nielsenae [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A species of Rhagonycha .

Rhynchitomimus [1]

Gen. et sp. nov

Valid

Clarke & Oberprieler in Clarke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A weevil belonging to the family Mesophyletidae and the subfamily Aepyceratinae. The type species is R. chalybeus.

Rhynchitomimus chalybeus Rhynchitomimus chalybeus holotype Fig1 B.jpg
Rhynchitomimus chalybeus

Sanaungulus [15]

Gen. et 2 sp. nov

Valid

Fanti, Damgaard & Ellenberger

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A soldier beetle. Genus includes new species S. curtipennis and S. ghitaenoerbyae.

Scydmobisetia loebli [76]

Sp. nov

Valid

Yin, Zhou & Cai

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An ant-like stone beetle.

Scydmobisetia mengjiae [76]

Sp. nov

Valid

Yin, Zhou & Cai

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An ant-like stone beetle.

Seidlitzella hoffeinsorum [77]

Sp. nov

Valid

Kolibáč & Alekseev

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of the family Trogossitidae belonging to the tribe Gymnochilini.

Serangium gedanicum [78]

Sp. nov

Valid

Szawaryn & Szwedo

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of the family Coccinellidae.

Serangium twardowskii [78]

Sp. nov

Valid

Szawaryn & Szwedo

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of the family Coccinellidae.

Shixitomaria [8]

Gen. et comb. nov

Valid

Lyubarsky & Perkovsky

Early Cretaceous (Aptian-Albian)

Shixi Formation

Flag of the People's Republic of China.svg  China

A member of the family Cryptophagidae belonging to the subfamily Atomariinae; a new genus for "Atomaria" cretacea Cai & Wang (2013).

Sinochaetodus [79]

Gen. et sp. nov

Valid

Lu et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of the family Hybosoridae. Genus includes new species S. tridentatus.

Sinohybosorus [79]

Gen. et sp. nov

Valid

Lu et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of the family Hybosoridae. Genus includes new species S. cheni.

Sorodites [80]

Gen. et sp. nov

Valid

Kirejtshuk & Chetverikov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A sap beetle. Genus includes new species S. angustipes.

Soronia menatensis [25]

Sp. nov

Valid

Kirejtshuk & Nel

Paleocene

Flag of France.svg  France

A sap beetle.

Spinotoma [81]

Gen. et sp. nov

Valid

Hsiao & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ripiphoridae belonging to the subfamily Pelecotominae. The type species is S. ruicheni.

Succinumanax [82]

Gen. et sp. nov

Valid

Bao, Rust & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A water-penny beetle belonging to the subfamily Eubrianacinae. Genus includes new species is S. birmaniasis.

Sucinorhagonycha samsockorum [55]

Sp. nov

Valid

Fanti & Pankowski

Eocene (Priabonian)

Prussian Formation
(Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae and the tribe Cantharini.

Symbiotes borussiaeorientalis [41]

Sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of Coccinelloidea belonging to the family Anamorphidae.

Taimyraltica [83]

Gen. et sp. nov

Valid

Nadein & Perkovsky

Late Cretaceous (Santonian)

Taimyr amber

Flag of Russia.svg  Russia

A leaf beetle belonging to the subfamily Galerucinae. Genus includes new species T. calcarata.

Tetracoleus aristovi [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Asiocoleidae.

Tetracoleus meyeni [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Asiocoleidae.

Tetracoleus vasilenkoi [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Asiocoleidae.

Themus bennyianderseni [9]

Sp. nov

Valid

Fanti & Damgaard

Eocene

Baltic amber

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae.

Tomoxia succinea [84]

Sp. nov

Valid

Bao et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Tomoxia .

Trichodesma fennosarmatica [85]

Sp. nov

Valid

Bukejs, Háva & Alekseev

Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A species of Trichodesma .

Trochoideus koenigsbergicus [41]

Sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of the family Endomychidae.

Trochoideus resinatissimus [41]

Sp. nov

Valid

Alekseev & Tomaszewska

Late Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

A member of the family Endomychidae.

Tunguskagyrus [86]

Gen. et sp. et comb. nov

Yan, Beutel & Lawrence

Permian (Changhsingian)

Lebedevskian Horizon

Flag of Russia.svg  Russia

A beetle of uncertain phylogenetic placement; originally described as a whirligig beetle, but subsequently argued to be a member of the family Triaplidae instead. [87] The type species is T. planus; genus might also include "Triaplus" sibiricus Volkov (2013), [87] though this species was also argued to be a member of the genus Tomiaplus instead. [88]

Tychtocoleus popovi [29]

Sp. nov

Valid

Ponomarenko

Late Permian

Komarity Beds of the Salaraevo Formation

Flag of Russia.svg  Russia

A member of the family Permosynidae.

Vetatrecus [89]

Gen. et 2 sp. nov

Valid

Kypke et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Staphylininae and the tribe Othiini. Genus includes new species V. adelfiae and V. secretum.

Vetudasycerus [90]

Gen. et sp. nov

Valid

Cai et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A rove beetle belonging to the subfamily Dasycerinae. Genus includes new species V. burmiticus.

Vetujinbrianax [91]

Gen. et sp. nov

Valid

Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A water-penny beetle. Genus includes new species V. cretaceus.

Zemyna [13]

Gen. et sp. nov

Valid

Tomaszewska in Tomaszewska et al.

Late Eocene

Baltic amber

Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of the family Endomychidae. Genus was originally named Laima Alekseev & Tomaszewska (2018); [41] however, this generic name turned out to be preoccupied by Laima Gravitis (1981). The type species is Z. andreei (Alekseev & Tomaszewska, 2018).

Dermapterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Acanthodiplatys [92]

Gen. et sp. nov

Valid

Ren et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Diplatyidae. Genus includes new species A. leptocercus.

Brevicula maculata [93]

Sp. nov

Valid

Kelly, Ross & Jarzembowski

Early Jurassic (Sinemurian)

Charmouth Mudstone Formation

Flag of the United Kingdom.svg  United Kingdom

An earwig belonging to the family Protodiplatyidae.

Dimapteron [93]

Gen. et sp. nov

Valid

Kelly, Ross & Jarzembowski

Early Cretaceous (Berriasian)

Durlston Formation

Flag of the United Kingdom.svg  United Kingdom

An earwig, possibly a member of the family Dermapteridae. The type species is D. corami.

Hirtidiplatys [92]

Gen. et sp. nov

Valid

Ren et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Diplatyidae. Genus includes new species H. cardiophyllus.

Phanerogramma australis [93]

Sp. nov

Valid

Kelly, Ross & Jarzembowski

Late Triassic (Carnian)

Blackstone Formation

Flag of Australia (converted).svg  Australia

An earwig, possibly a member of the family Dermapteridae.

Phanerogramma dunstani [93]

Sp. nov

Valid

Kelly, Ross & Jarzembowski

Late Triassic (Carnian)

Blackstone Formation

Flag of Australia (converted).svg  Australia

An earwig, possibly a member of the family Dermapteridae.

Phanerogramma gouldsbroughi [93]

Sp. nov

Valid

Kelly, Ross & Jarzembowski

Rhaetian or Hettangian

Flag of the United Kingdom.svg  United Kingdom

An earwig, possibly a member of the family Dermapteridae.

Robustipygia [92]

Gen. et sp. nov

Valid

Ren et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Pygidicranidae. Genus includes new species R. calvata.

Trivenapteron [93]

Gen. et sp. nov

Valid

Kelly, Ross & Jarzembowski

Early Jurassic (Toarcian)

Flag of the United Kingdom.svg  United Kingdom

An earwig of uncertain phylogenetic placement. The type species is T. moorei.

Tytthodiplatys ortholabis [92]

Sp. nov

Valid

Ren et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Diplatyidae.

Valdopteron [93]

Gen. et sp. nov

Valid

Kelly, Ross & Jarzembowski

Early Cretaceous (Barremian)

Upper Weald Clay

Flag of the United Kingdom.svg  United Kingdom

An earwig, possibly a member of the family Dermapteridae. The type species is V. woodi.

Dictyopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Alienopterella [94]

Gen. et sp. nov

Valid

Kočárek

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of Alienopterus . Genus includes new species A. stigmatica.

Alienopterix [95]

Gen. et sp. nov

Valid

Mlynský, Vršanský & Wang in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of Alienopterus [95] or a member of the family Cratovitismidae. [96] The type species is A. ocularis.

Apiblatta [95]

Gen. et sp. nov

Valid

Barna & Bigalk in Vršanský et al.

Early Cretaceous (Albian)

Crato Formation

Flag of Brazil.svg  Brazil

A relative of Alienopterus . The type species is A. muratai.

Burmantis hexispinea [97]

Sp. nov

Valid

Li & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A mantis.

Caputoraptor [98] [99]

Gen. et 2 sp. nov

Valid

Bai, Beutel & Wipfler in Bai et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of Alienopterus . The type species is C. elegans; genus also includes C. vidit Šmídová, Vršanský & Wang in Vršanský et al. (2018). [95]

Chimaeroblattina [95]

Gen. et sp. nov

Valid

Barna in Vršanský et al.

Middle Eocene

Green River Formation

Flag of the United States.svg  United States

A relative of Alienopterus . The type species is C. brevipes.

Cratovitisma bechlyi [100]

Sp. nov

Valid

Podstrelená in Podstrelená & Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A beetle-like cockroach belonging to the family Umenocoleidae.

Cratovitisma cortexi [100]

Sp. nov

Valid

Sendi in Podstrelená & Sendi

Early Cretaceous

Lebanese amber

Flag of Lebanon.svg  Lebanon

A beetle-like cockroach belonging to the family Umenocoleidae.

Falcatusiblatta [101]

Gen. et 2 sp. et comb. nov

Valid

Liang, Shih & Ren

Middle and Late Jurassic

Daohugou Beds
Karabastau Svita

Flag of the People's Republic of China.svg  China
Flag of Kazakhstan.svg  Kazakhstan

A cockroach belonging to the family Raphidiomimidae. Genus includes new species F. gracilis and F. qiandaohua, as well as "Rhipidoblattina" karatavica Vishniakova (1968).

Grant [95]

Gen. et sp. nov

Valid

Aristov in Vršanský et al.

Middle Eocene

Green River Formation

Flag of the United States.svg  United States

A relative of Alienopterus . The type species is G. viridifluvius.

Meilia [95]

Gen. et sp. nov

Valid

Vršanský et Wang in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of Alienopterus . The type species is M. jinghanae.

Nodosigalea [102]

Gen. et sp. nov

Valid

Li & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species N. burmanica.

Phyloblatta beijingensis [103]

Sp. nov

Valid

Huang et al.

Carboniferous-Permian

Taiyuan Formation

Flag of the People's Republic of China.svg  China

A cockroach.

Piniblattella yixianensis [104]

Sp. nov

Valid

Gao, Shih & Ren in Gao et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A cockroach. Transferred to the genus Spinaeblattina by Hinkelman (2019). [105]

Teyia [95]

Gen. et 2 sp. nov

Valid

Vršanský, Mlynský & Wang in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of Alienopterus . The type species is T. branislav Vršanský & Wang (2018); genus also includes T. huangi Vršanský, Mlynský & Wang (2018).

Vcelesvab [95]

Gen. et sp. nov

Valid

Vršanský, Barna & Bigalk in Vršanský et al.

Early Cretaceous (Albian)

Crato Formation

Flag of Brazil.svg  Brazil

A relative of Alienopterus . The type species is V. cratocretokrat.

Dipterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Acartophthalmites willii [106]

Sp. nov

Valid

Pérez-de la Fuente, Hoffeins & Roháček

Eocene

Baltic amber

Baltic Sea coast, probably Flag of Russia.svg  Russia (Kaliningrad Oblast)

A member of Opomyzoidea of uncertain phylogenetic placement.

Alonchoptera [107]

Gen. et sp. nov

Valid

Grimaldi

Early Cretaceous

Flag of Lebanon.svg  Lebanon

A member of the stem group of Lonchopteroidea. The type species is A. lebanica.

Archilimonia grauvogeliana [108]

Sp. nov

Valid

Lukashevich & Ribeiro

Middle Triassic (Anisian)

Flag of France.svg  France

An early member of Tipuloidea.

Archipleciomima germanica [109]

Sp. nov

Valid

Kopeć et al.

Early Jurassic (Toarcian)

Flag of Germany.svg  Germany

A member of Mycetophiloidea.

Aschizomyia [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Syrphoidea of uncertain phylogenetic placement. The type species is A. burmensis.

Austroconops perrichoti [110]

Sp. nov

Valid

Dominiak, Szadziewski & Nel

Cretaceous (latest Albian-earliest Cenomanian)

Flag of France.svg  France

A member of the family Ceratopogonidae belonging to the subfamily Leptoconopinae.

Burmapeza [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Platypezidae. The type species is B. radicis.

Burmazelmira grimaldii [111]

Sp. nov

Valid

Arillo, Blagoderov & Peñalver

Early Cretaceous (Albian)

Maestrat Basin

Flag of Spain.svg  Spain

A member of Archizelmiridae.

Calvopeza [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Platypezidae. The type species is C. divergens.

Canadopeza [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous

Flag of Canada (Pantone).svg  Canada

A member of the family Platypezidae. The type species is C. biacrosticha.

Chandleromyia [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Platypezidae. The type species is C. anomala.

Dacochile browni [112]

Sp. nov

Valid

Skibińska & Krzemiński

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Tanyderidae.

Dacochile poinari [112]

Sp. nov

Valid

Skibińska & Krzemiński

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Tanyderidae.

Dicranomyia (Dicranomyia) colombiana [113]

Sp. nov

Valid

Krzemiński et al.

Colombian copal

Flag of Colombia.svg  Colombia

A species of Dicranomyia .

Dicranomyia (Dicranomyia) indica [114]

Sp. nov

Valid

Kania et al.

Early Eocene

Cambay amber

Flag of India.svg  India

A species of Dicranomyia .

Dicranomyia (Melanolimonia) kukulai [115]

Sp. nov

Valid

Krzemiński, Kania & Wojtoń

Eocene

Baltic amber

Europe (Baltic Sea coast)

A species of Dicranomyia .

Eodromyia [116]

Gen. et sp. nov

Valid

Myskowiak, Garrouste & Nel

Early Eocene

Oise amber

Flag of France.svg  France

A member of the family Hybotidae. Genus includes new species E. pumilio.

Eoprocladius [117]

Gen. et sp. nov

Valid

Szadziewski, Sontag & Dominiak

Eocene

Baltic amber

Baltic Sea region

A member of the family Chironomidae belonging to the subfamily Tanypodinae and the tribe Procladiini. Genus includes new species E. hoffeinsorum.

Eosciadocera pauciseta [107]

Sp. nov

Valid

Grimaldi

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Phoridae belonging to the subfamily Sciadocerinae.

Gujaratomyia [118]

Gen. et sp. nov

Valid

Giłka & Zakrzewska in Zakrzewska et al.

Early Eocene

Cambay amber

Flag of India.svg  India

A non-biting midge belonging to the tribe Tanytarsini. Genus includes new species G. miripes.

Helius spiralensis [119]

Sp. nov

Valid

Kania, Krzemiński & Arillo

Early Cretaceous

Álava amber

Flag of Spain.svg  Spain

A species of Helius .

Irwinimyia [120]

Gen. et sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Apsilocephalidae. Genus includes new species I. spinosa.

Langtonius [121]

Gen. et sp. nov

Valid

Lukashevich & Przhiboro

Early Cretaceous

Tsagan Tsab Formation

Flag of Mongolia.svg  Mongolia

A member of the family Chironomidae. Genus includes new species L. cynaricaudatus.

Lebanopeza [107]

Gen. et sp. nov

Valid

Grimaldi

Early Cretaceous

Flag of Lebanon.svg  Lebanon

A member of the family Platypezidae. The type species is L. azari.

Libanosycorax [122]

Gen. et sp. nov

Valid

Azar, Azar & Maksoud

Early Cretaceous

Flag of Lebanon.svg  Lebanon

A relative of members of the genus Sycorax . Genus includes new species L. dimyi.

Lindneromyia dominicana [107]

Sp. nov

Valid

Grimaldi

Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A species of Lindneromyia .

Lindneromyia neomedialis [107]

Sp. nov

Valid

Grimaldi

Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A species of Lindneromyia .

Lonchopterites burmensis [107]

Sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the stem group of Lonchopteroidea.

Macalpinomyia [123]

Gen. et sp. nov

Valid

Li & Yeates

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ironomyiidae. Genus includes new species M. jiewenae.

Mailotrichocera praedicta [124]

Sp. nov

Valid

Krzemińska & Lukashevich

Early Jurassic (possibly Sinemurian)

Dzil Formation

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of the family Trichoceridae.

Mesorhyphus ulrichi [109]

Sp. nov

Valid

Kopeć et al.

Early Jurassic (Toarcian)

Flag of Germany.svg  Germany

A member of the family Anisopodidae.

Mesotipula slatteri [125]

Sp. nov

Valid

Kopeć

Early Jurassic

Flag of the United Kingdom.svg  United Kingdom

A member of Limoniidae.

Myanmarpsilocephala [120]

Gen. et sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Apsilocephalidae. Genus includes new species M. grimaldii.

Neuseptychoptera [126]

Gen. et sp. nov

Valid

Szadziewski, Krynicki & Krzemiński

Late Cretaceous (Campanian)

Tar Heel Formation

Flag of the United States.svg  United States

A member of Ptychopteridae belonging to the subfamily Eoptychopterinae. Genus includes new species N. carolinensis.

Oligopipiza [127]

Gen et sp. nov

Valid

Nidergras, Hadrava & Nel in Nidergas et al.

Oligocene

Flag of France.svg  France

A hoverfly. The type species is O. quadriguttata.

Palaeopetia dorsalis [107]

Sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ironomyiidae.

Palaeopetia terminus [107]

Sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ironomyiidae.

Proironia [107]

Gen. et 2 sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Ironomyiidae. The type species is P. gibbera; genus also includes P. burmitica.

Prophora [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomianian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Phoridae of uncertain phylogenetic placement. The type species is P. dimorion.

Prosyrphus [107]

Gen. et sp. nov

Valid

Grimaldi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A possible stem-hoverfly. The type species is P. thompsoni.

Serromyia errata [128]

Sp. nov

Valid

Szadziewski

Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

A species of Serromyia ; a new species established for the specimens originally assigned to the species Serromyia alphea (Heyden, 1870).

Similinannotanyderus zbigniewi [129]

Sp. nov

Valid

Skibińska & Krzemiński

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Tanyderidae.

Sylvicola baltica [130]

Sp. nov

Valid

Wojtoń, Kania & Kopeć

Eocene

Baltic amber

Baltic Sea region

A species of Sylvicola .

Sylvicola hoffeinsorum [130]

Sp. nov

Valid

Wojtoń, Kania & Kopeć

Eocene

Baltic amber
Bitterfeld amber

Baltic Sea region
Flag of Germany.svg  Germany

A species of Sylvicola .

Sylvicola punctata [130]

Sp. nov

Junior homonym

Wojtoń, Kania & Kopeć

Eocene

Baltic amber

Baltic Sea region

A species of Sylvicola . The name is preoccupied by Sylvicola punctatus (Fabricius, 1787); Hancock & Kania (2019) coined a replacement name Sylvicola harrisi. [131]

Tipula americana [132]

Sp. nov

Valid

Kania et al.

Eocene

Green River Formation

Flag of the United States.svg  United States

A species of Tipula .

Ugolyakia [133]

Gen. et sp. nov

Valid

Perkovsky, Sukhomlin & Zelenkov

Late Cretaceous (Santonian)

Taimyr amber

Flag of Russia.svg  Russia

A black fly. Genus includes new species U. kaluginae.

Hemipterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Anthoscytina daidaleos [134]

Sp. nov

Valid

Fu, Huang & Engel

Middle or Late Jurassic

Haifanggou Formation

Flag of the People's Republic of China.svg  China

A froghopper belonging to the family Procercopidae.

Aphrastomedes [135]

Gen. et sp. nov

Valid

Yamada & Yamamoto in Yamada, Yamamoto & Takahashi

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Cimicomorpha, probably belonging to the family Velocipedidae. Genus includes new species A. anthocoroides.

Archetingis [136]

Gen. et sp. nov

Valid

Montagna et al.

Middle Triassic (Ladinian)

Meride Limestone

Flag of Switzerland (Pantone).svg   Switzerland

A member of the family Tingidae. The type species is A. ladinica.

Archipedionis [137]

Gen. et sp. nov

Valid

Dietrich & Thomas

Eocene

Baltic amber

Europe (Baltic Sea coast)

A leafhopper belonging to the subfamily Eurymelinae and the tribe Macropsini. The type species is A. obscurus.

Burmacader lativentris [138]

Sp. nov

Valid

Heiss & Guilbert

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Tingidae.

Camuracicada [139]

Gen. et comb. nov

Valid

Moulds

Miocene (Serravallian)

Flag of Croatia.svg  Croatia

A cicada; a new genus for "Cicada" aichhorni Heer (1853).

Chilamnestocoris [140]

Gen. et sp. nov

Valid

Lis, Lis & Heiss

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Cydnidae. Genus includes new species C. mixtus.

Criniverticillus [141]

Gen. et sp. nov

Valid

Lin, Yao & Ren

Cretaceous (Albian-Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of scale insects belonging to the family Weitschatidae. Genus includes new species C. longicumulus.

Cucullitingis [142]

Gen. et sp. nov

Valid

Du & Yao

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Tingidae. The type species is C. biacantha.

Dachibangus [143]

Gen. et sp. nov

Valid

Jiang, Szwedo & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A planthopper belonging to the family Mimarachnidae. Genus includes new species D. trimaculatus.

Dorytocus [144]

Gen. et sp. nov

Valid

Emeljanov & Shcherbakov

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A planthopper belonging to the new family Dorytocidae. Genus includes new species D. ornithorhynchus.

Eoidiocerus [137]

Gen. et sp. nov

Valid

Dietrich & Thomas

Eocene

Baltic amber

Europe (Baltic Sea coast)

A leafhopper belonging to the subfamily Eurymelinae and the tribe Idiocerini. The type species is E. emarginatus.

Fangyuania [145]

Gen. et sp. nov

Valid

Chen, Szwedo & Wang in Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A froghopper belonging to the family Sinoalidae. Genus includes new species F. xiai.

Hormatalis [146]

Gen. et sp. nov

Valid

Węgierek & Wang in Węgierek et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An aphid belonging to the family Isolitaphidae. Genus includes new species H. lancigerens.

Jaculistilus [147]

Gen. et sp. nov

Valid

Zhang, Ren & Yao

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A planthopper belonging to the family Mimarachnidae. Genus includes new species J. oligotrichus.

Leptopharsa antica [148]

Sp. nov

Valid

Golub & Heiss

Eocene or Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A species of Leptopharsa .

Leptopharsa colombiana [148]

Sp. nov

Valid

Golub & Heiss

Colombian copal

Flag of Colombia.svg  Colombia

A species of Leptopharsa .

Luanpingia daohugouensis [149]

Sp. nov

Valid

Fu, Cai & Huang

Middle or Late Jurassic

Daohugou Beds

Flag of the People's Republic of China.svg  China

A froghopper belonging to the family Sinoalidae.

Paleopsalta [139]

Gen. et comb. nov

Valid

Moulds

Miocene (Serravallian)

Flag of Croatia.svg  Croatia

A cicada; a new genus for "Cicada" ungeri Heer (1853).

Phatnoma pulchra [148]

Sp. nov

Valid

Golub & Heiss

Eocene or Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A member of the family Tingidae.

Prolavexillaphis [150]

Gen. et sp. nov

Valid

Liu, Qiao & Yao in Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An aphid belonging to the family Juraphididae. Genus includes new species P. munditia.

Stictocercopis [151]

Gen. et sp. nov

Valid

Fu & Huang

Jurassic (Callovian or Oxfordian)

Haifanggou Formation

Flag of the People's Republic of China.svg  China

A froghopper belonging to the family Sinoalidae. The type species is S. wuhuaensis.

Tanyaulus [152]

Gen. et sp. nov

Valid

Poinar

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

An aphid belonging to the family Burmitaphidae. Genus includes new species T. caudisetula.

Hymenopterans

NameNoveltyStatusAuthorship of new nameAgeUnitType LocationNotesImages

Amplicella minor [153]

Sp. nov

Valid

Kopylov

Early Cretaceous

Flag of Russia.svg  Russia

A member of the family Ichneumonidae.

Angarosphex saxosus [154]

Sp. nov

Valid

Zhang, Rasnitsyn & Zhang

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Apoidea belonging to the family Angarosphecidae.

Apis (Synapis) dalica [155]

Sp. nov

Valid

Engel & Wappler in Engel et al.

Middle Miocene

Huazhige Formation

Flag of the People's Republic of China.svg  China

A honey bee.

Aptenoperissus amabilis [156]

Sp. nov

Valid

Zhang et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus delicatus [156]

Sp. nov

Valid

Zhang et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus etius [157]

Sp. nov

Valid

Rasnitsyn & Öhm-Kühnle

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus formosus [156]

Sp. nov

Valid

Zhang et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus magnifemoris [157]

Sp. nov

Valid

Rasnitsyn & Öhm-Kühnle

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus pusillus [157]

Sp. nov

Valid

Rasnitsyn & Öhm-Kühnle

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Aptenoperissus zonalis [158]

Sp. nov

Valid

Zhang & Rasnitsyn in Zhang, Rasnitsyn & Zhang

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Aptenoperissidae.

Archaeocercus [159]

Gen. et sp. nov

Valid

Simutnik & Perkovsky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A member of the family Encyrtidae. Genus includes new species A. schuvachinae.

Auricleptes [160]

Gen. et sp. nov

Valid

Lucena & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cuckoo wasp. Genus includes new species A. nebulosus.

Azanichrum [160]

Gen. et sp. nov

Valid

Lucena & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cuckoo wasp. Genus includes new species A. pilosum.

Baeomorpha avamica [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha baikurenis [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Cretaceous (late Albian–?early Cenomanian)

Ognevka Formation?
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha bianellus [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha caeleps [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha gracilis [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha ingens [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha popovi [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha quattorduo [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha quattoruno [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha yantardakh [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Baeomorpha zherikhini [161]

Sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Cretaceous (late Albian–?early Cenomanian)

Ognevka Formation?
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae.

Bohartiura [160]

Gen. et sp. nov

Valid

Lucena & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cuckoo wasp. Genus includes new species B. glabrata.

Brachysyntexis tenebrosa [162]

Sp. nov

Valid

Kopylov

MiddleLate Jurassic

Flag of Kazakhstan.svg  Kazakhstan

A member of the family Anaxyelidae.

Brachysyntexis tigris [162]

Sp. nov

Valid

Kopylov

MiddleLate Jurassic

Flag of Kazakhstan.svg  Kazakhstan

A member of the family Anaxyelidae.

Burmasega [160]

Gen. et sp. nov

Valid

Lucena & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cuckoo wasp. Genus includes new species B. ammirabilis.

Burmasphex [163]

Gen. et 2 sp. nov

Valid

Melo & Rosa

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Apoidea belonging to the family Angarosphecidae. The type species is B. sulcatus; genus also includes B. pilosus.

Burminata [164]

Gen. et sp. nov

Valid

Haas, Burks & Krogmann

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A chalcid wasp belonging to the new family Diversinitidae. The type species is B. caputaeria.

Burmusculus [165]

Gen. et 3 sp. nov

Valid

Zhang & Rasnitsyn in Zhang, Rasnitsyn & Zhang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the group Pompiloidea. Genus includes new species B. nuwae, B. fuxii and B. shennongii.

Carinibus [166]

Gen. et sp. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A member of the family Ichneumonidae. The type species is C. molestus.

Chifengilyda [167]

Gen. et sp. nov

Valid

Zheng & Chen

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A sawfly belonging to the family Xyelydidae. Genus includes new species C. robusta.

Colmepsiterona [168]

Gen. et sp. nov

Valid

Cockx & McKellar

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A wasp belonging to the family Crabronidae and the subfamily Pemphredoninae. Genus includes new species C. cumcarena.

Conostigmus talamasi [169]

Sp. nov

Valid

Mikó & Trietsch in Mikó et al.

Eocene

Baltic amber

Europe
(Gdańsk Bay)

A species of Conostigmus .

Coptera anka [170]

Sp. nov

Valid

Krogmann, van de Kamp & Schwermann in van de Kamp et al.

Paleogene

Quercy Phosphorites Formation

Flag of France.svg  France

A member of the family Diapriidae.

Cretevania tenuis [171]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea.

Cretevania venae [171]

Sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Evanioidea.

Cretosirex [172]

Gen. et sp. nov

Valid

Wang et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A horntail. Genus includes new species C. xiaoi.

Crusopimpla [173]

Gen. et sp. nov

Valid

Kopylov, Spasojevic & Klopfstein

Early Eocene

Tadushi Formation

Flag of Russia.svg  Russia

A member of the family Ichneumonidae. Genus includes new species C. tadushensis.

Curtevania [171]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea. Genus includes new species C. enervia.

Dencyrtus [174]

Gen. et sp. nov

Valid

Simutnik & Perkovsky

Late Eocene

Danish amber

Flag of Denmark.svg  Denmark

A member of the family Encyrtidae. Genus includes new species D. vilhelmseni.

Dolichomitus? saxeus [173]

Sp. nov

Valid

Kopylov, Spasojevic & Klopfstein

Early Eocene

Tadushi Formation

Flag of Russia.svg  Russia

Possibly a species of Dolichomitus .

Dryinus maderai [175]

Sp. nov

Valid

Guglielmino et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Dryinidae.

Dryinus maderai Dryinus maderai holotype Fig1.jpg
Dryinus maderai

Diversinitus [164]

Gen. et sp. nov

Valid

Haas, Burks & Krogmann

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A chalcid wasp belonging to the new family Diversinitidae. The type species is D. attenboroughi.

Diversinitus attenboroughi Diversinitus attenboroughi-1 (cropped).png
Diversinitus attenboroughi

Electromyrmex wheeleri [176]

Sp. nov

Valid

Radchenko & Dlussky

Late Eocene

Baltic amber
Bitterfeld amber

Baltic Sea region
Flag of Germany.svg  Germany

Epyris moulyi [177]

Sp. nov

Valid

Falières & Nel

Early Eocene

Oise amber

Flag of France.svg  France

A species of Epyris .

Epyris neolongiceps [178]

Nom. nov

Valid

Azevedo in Azevedo et al.

Oligocene

Baltic amber

Europe (Baltic Sea region)

A species of Epyris ; a replacement name for Calyoza longiceps Brues (1923).

Eupsenella neoeocenica [178]

Nom. nov

Valid

Ramos & Azevedo in Azevedo et al.

Eocene

Fushun amber

Flag of the People's Republic of China.svg  China

A member of the family Bethylidae; a replacement name for Fushunochrysites eocenicus Hong (2002).

Eupsenella neohongi [178]

Nom. nov

Valid

Ramos & Azevedo in Azevedo et al.

Eocene

Fushun amber

Flag of the People's Republic of China.svg  China

A member of the family Bethylidae; a replacement name for Sinibethylus eocenicus Hong (2002).

Exilaulacus [171]

Gen. et 2 sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea. Genus includes new species E. loculatus and E. latus.

Fallomyrma anodonta [179]

Sp. nov

Valid

Radchenko & Dlussky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

An ant belonging to the subfamily Myrmicinae.

Fallomyrma marginata [179]

Sp. nov

Valid

Radchenko & Dlussky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

An ant belonging to the subfamily Myrmicinae.

Fallomyrma robusta [179]

Sp. nov

Valid

Radchenko & Dlussky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

An ant belonging to the subfamily Myrmicinae.

Glabiala [164]

Gen. et sp. nov

Valid

Haas, Burks & Krogmann

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A chalcid wasp belonging to the new family Diversinitidae. The type species is G. barbata.

Heterobaissa [171]

Gen. et sp. nov

Valid

Li et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Evanioidea. Genus includes new species H. apetiola.

Ichninsum [166]

Gen. et sp. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A pimpline Ichneumonidae. The type species is I. appendicrassum.

Kulbastavia grandis [162]

Sp. nov

Valid

Kopylov

MiddleLate Jurassic

Flag of Kazakhstan.svg  Kazakhstan

A member of the family Anaxyelidae.

Mesoclistus? yamataroti [166]

Sp. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A possible Coleocentrini tribe acaenitine darwin wasp. [166]

Mesornatus [180]

Gen. et sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae of uncertain phylogenetic placement. The type species is M. markovici.

Miracorium [160]

Gen. et sp. nov

Valid

Lucena & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A cuckoo wasp. Genus includes new species M. tetrafoveolatum.

Myanmarina [181]

Gen. et 4 sp. nov

Valid

Zhang & Rasnitsyn in Zhang et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Apocrita belonging to the superfamily Stephanoidea. The type species is M. lisu; genus also includes new species M. kachin, M. lahu [181] and M. jeannineae Li et al. (2018). [182]

Napakimyrma [183]

Gen. et sp. nov

Valid

Lapolla & Barden

Paleocene

Paskapoo Formation

Flag of Canada (Pantone).svg  Canada

An ant belonging to the subfamily Aneuretinae. The type species is N. paskapooensis.

Newjersevania brevis [171]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea.

Newjersevania longa [171]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea.

Palaeortona [170]

Gen. et sp. nov

Valid

Krogmann, van de Kamp & Schwermann in van de Kamp et al.

Paleogene

Quercy Phosphorites Formation

Flag of France.svg  France

A member of the family Diapriidae. The type species is P. quercyensis.

Parevania oculiseparata [184]

Sp. nov

Valid

Jennings et al.

Eocene

Baltic amber

Flag of Europe.svg  Europe

A member of the family Evaniidae.

Peleproctus [185]

Gen. et sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A basal member of Proctotrupomorpha. Genus includes new species P. dolichurus.

Peleserphus [185]

Gen. et 2 sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A basal member of Proctotrupomorpha. The type species is P. brachyurus; genus also includes P. genalis.

Phanerotomella brevivena [186]

Sp. nov

Valid

Kittel

Eocene

Baltic amber

Europe
(Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Cheloninae.

Polyhelictes [180]

Gen. et sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae of uncertain phylogenetic placement. The type species is P. bipolarus.

Praeaulacus rectus [171]

Sp. nov

Valid

Li et al.

Middle Jurassic

Jiulongshan Formation

Flag of the People's Republic of China.svg  China

A member of Evanioidea.

Pristomyrmex archaios [187]

Sp. nov

Valid

Radchenko & Dlussky

Eocene

Bitterfeld amber

Flag of Germany.svg  Germany

An ant, a species of Pristomyrmex .

Pristomyrmex elmesi [187]

Sp. nov

Valid

Radchenko & Dlussky

Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

An ant, a species of Pristomyrmex .

Proleptothorax [188]

Gen. et sp. nov

Valid

Radchenko, Dlussky & Perfilieva

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

An ant. Genus includes new species P. primitivus.

Rhyssella vera [180]

Sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae, a species of Rhyssella .

Scambus fossilobus [180]

Sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae, a species of Scambus .

Scambus? mandibularis [166]

Sp. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A possible Ephialtini tribe pimpline darwin wasp.

Scambus? parachuti [166]

Sp. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A possible Ephialtini tribe pimpline darwin wasp.

Seneciobracon [189]

Gen. et sp. nov

Valid

Engel & Huang in Engel et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Braconidae. The type species is S. novalatus.

Seneciobracon novalatus Seneciobracon novalatus holoytpe Zookeys.jpg
Seneciobracon novalatus

Sinuevania [171]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Evanioidea. Genus includes new species S. mira.

Taimyromorpha [161]

Gen. et sp. nov

Valid

Gumovsky in Gumovsky, Perkovsky & Rasnitsyn

Late Cretaceous (late Santonian)

Kheta Formation
(Taimyr amber)

Flag of Russia.svg  Russia

A member of the family Rotoitidae. Genus includes new species T. pusilla.

Trigonator [180]

Gen. et sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae belonging to the subfamily Labeninae. The type species is T. macrocheirus.

Trjapitzion [190]

Gen. et sp. nov

Valid

Simutnik in Simutnik & Perkovsky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A member of the family Encyrtidae. Genus includes new species T. cylindrocerus.

Trymectus [166]

Gen. et comb. nov

Valid

Spasojevic et al.

Early Eocene

Green River Formation

Flag of the United States.svg  United States

A darwin wasp of uncertain subfamily placement.
The type species is Tryphon amasidis (1931).

Versolabrum [191]

Gen. et sp. nov

Valid

Burks & Krogmann in Burks, Krogmann & Heraty

Eocene

Baltic amber

Europe
(Baltic Sea region)

A member of the family Pteromalidae. Genus includes new species V. coriaceum.

Vitimosphex vividus [154]

Sp. nov

Valid

Zhang, Rasnitsyn & Zhang

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Apoidea belonging to the family Angarosphecidae.

Xanthopimpla messelensis [180]

Sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae belonging to the subfamily Pimplinae and the tribe Pimplini.

Xanthopimpla messelensis Xanthopimpla messelensis sp. nov., holotype SF MeI 16988.tiff
Xanthopimpla messelensis

Xanthopimpla praeclara [180]

Sp. nov

Valid

Spasojevic, Wedmann & Klopfstein

Eocene

Messel pit

Flag of Germany.svg  Germany

A member of the family Ichneumonidae belonging to the subfamily Pimplinae and the tribe Pimplini.

Xenomorphia [170]

Gen. et 2 sp. nov

Valid

Krogmann, van de Kamp & Schwermann in van de Kamp et al.

Paleogene

Quercy Phosphorites Formation

Flag of France.svg  France

A member of the family Diapriidae. The type species is X. resurrecta; genus also includes X. handschini.

Xenomorphia resurrecta Xenomorphia resurrecta fossil tomography.png
Xenomorphia resurrecta

Mecopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Bittacus lepiduscretaceus [192]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A hangingfly, a species of Bittacus .

Eomerope eonearctica [193]

Sp. nov

Valid

Archibald & Rasnitsyn

Eocene (Ypresian)

Flag of Canada (Pantone).svg  Canada

A member of the family Eomeropidae.

Eomerope simpkinsae [193]

Sp. nov

Valid

Archibald & Rasnitsyn

Eocene (Ypresian)

Allenby Formation

Flag of Canada (Pantone).svg  Canada

A member of the family Eomeropidae.

Gigaphlebia [194]

Gen. et comb. nov

Valid

Soszyńska-Maj et al.

Jurassic

Dzil Formation
Jiulongshan Formation
Karabastau Svita

Flag of the People's Republic of China.svg  China
Flag of Kazakhstan.svg  Kazakhstan
Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of the family Orthophlebiidae. Genus includes "Orthophlebia" riccardii Petrulevičius & Ren (2012), "Orthophlebia" grandis Martynov (1927) and "Mesopanorpa" palmaris Martynova (1948).

Hongchoristites [195]

Nom. nov

Valid

Hernández

Middle Triassic

Tongchuan Formation

Flag of the People's Republic of China.svg  China

A member of the family Permochoristidae; a replacement name for Choristites Hong (2005).

Longiphlebia [194]

Gen. et comb. nov

Valid

Soszyńska-Maj et al.

Middle Jurassic

Jiulongshan Formation

Flag of the People's Republic of China.svg  China

A member of the family Orthophlebiidae. Genus includes "Orthophlebia" stigmosa Qiao, Shih & Ren (2012).

Mesopanorpa brooksorum [196]

Sp. nov

Valid

Jarzembowski & Soszyńska-Maj

Early Cretaceous

Wealden Supergroup

Flag of the United Kingdom.svg  United Kingdom

A member of the family Orthophlebiidae.

Orthobittacus suni [197]

Sp. nov

Valid

Kopeć et al.

Middle Jurassic

Flag of the People's Republic of China.svg  China

A hangingfly.

Worcestobia [198]

Gen. et comb. nov

Valid

Soszyńska-Maj et al.

Triassic

Flag of Japan.svg  Japan
Flag of the United Kingdom.svg  United Kingdom

A new genus for "Orthophlebia" gigantea Tillyard (1933) and "O." haradai Ueda (1991).

Neuropterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Aberrantochrysa [199]

Gen. et 2 sp. nov

Valid

Khramov

Early Cretaceous

Flag of Russia.svg  Russia

A green lacewing. Genus includes new species A. buryatica and A. pulchella.

Acanthopsychops [200]

Gen. et sp. nov

Valid

Badano & Engel in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychopsidae. The type species is A. triaina.

Acanthopsychops triaina Acanthopsychops triaina holotype Sup Fig 1 F.jpg
Acanthopsychops triaina

Adelpholeon [200]

Gen. et sp. nov

Valid

Badano & Engel in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of antlions and owlflies. The type species is A. lithophorus.

Adelpholeon lithophorus Adelpholeon lithophorus holotype fig 1 A.jpg
Adelpholeon lithophorus

Aphthartopsychops [200]

Gen. et sp. nov

Valid

Badano & Engel in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychopsidae. The type species is A. scutatus.

Aphthartopsychops scutatus Aphthartopsychops scutatus holotype fig 1 A.jpg
Aphthartopsychops scutatus

Burmitus [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of antlions and owlflies. The type species is B. tubulifer.

Burmitus tubulifer Burmitus tubulifer holotype Sup Fig 2 D.jpg
Burmitus tubulifer

Burmogramma [201]

Gen. et sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Kalligrammatidae belonging to the subfamily Cretanallachiinae. The type species is B. liui.

Burmogramma liui Burmogramma liui Fig1 A.jpg
Burmogramma liui

Burmopsychops labandeirai [201]

Sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Kalligrammatidae belonging to the subfamily Cretanallachiinae.

Burmopsychops labandeirai Burmopsychops labandeirai holotype.jpg
Burmopsychops labandeirai

Caririneura macrothoracica [202]

Sp. nov

Valid

Makarkin, Wedmann & Heads

Early Cretaceous

Crato Formation

Flag of Brazil.svg  Brazil

A member of Myrmeleontoidea belonging to the family Araripeneuridae.

Chorilingia bakharica [203]

Sp. nov

Valid

Khramov & Vasilenko

Jurassic

Flag of Mongolia.svg  Mongolia

A member of the family Grammolingiidae.

Cladofer [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the stem group of Myrmeleontiformia. The type species is C. huangi.

Cladofer huangi Cladofer huangi holotype Sup Fig 1 D.jpg
Cladofer huangi

Cratoneura minor [202]

Sp. nov

Valid

Makarkin, Wedmann & Heads

Early Cretaceous

Crato Formation

Flag of Brazil.svg  Brazil

A member of Myrmeleontoidea belonging to the family Araripeneuridae.

Cretogramma [201]

Gen. et sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Kalligrammatidae belonging to the subfamily Cretanallachiinae. The type species is C. engeli.

Cretogramma engeli Cretogramma engeli holotype.jpg
Cretogramma engeli

Diodontognathus [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of antlions and owlflies. The type species is D. papillatus.

Diodontognathus papillatus Diodontognathus papillatus holotype fig 1 A.jpg
Diodontognathus papillatus

Electrocaptivus [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of antlions and owlflies. The type species is E. xui.

Electrocaptivus xui Electrocaptivus xui holotype fig 1 A.jpg
Electrocaptivus xui

Haploberotha carsteni [204]

Sp. nov

Valid

Makarkin

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Berothidae.

Kujiberotha [205]

Gen. et sp. nov

Valid

Nakamine & Yamamoto

Late Cretaceous (Santonian)

Tamagawa Formation
(Kuji amber)

Flag of Japan.svg  Japan

A thorny lacewing. The type species is K. teruyukii.

Kujiberotha teruyukii Kujiberotha teruyukii (10.3897-zookeys.802.28754) Figure 2.jpg
Kujiberotha teruyukii

Laccosmylus cicatricatus [206]

Sp. nov

Valid

Fang et al.

Middle Jurassic (Bathonian-Callovian boundary)

Jiulongshan Formation

Flag of the People's Republic of China.svg  China

A member of the family Saucrosmylidae.

Laccosmylus latizonus [206]

Sp. nov

Valid

Fang et al.

Middle Jurassic (Bathonian-Callovian boundary)

Jiulongshan Formation

Flag of the People's Republic of China.svg  China

A member of the family Saucrosmylidae.

Longipronotum [207] [208]

Gen. et sp. nov

Valid

Jepson, Khramov & Ohl

Late Jurassic

Flag of Kazakhstan.svg  Kazakhstan

A member of the family Mantispidae. Genus includes new species L. benmaddoxi. The original generic name, Longicollum, turned out to be preoccupied by Longicollum Yamaguti (1935), necessitating creation of a replacement name.

Macleodiella [200]

Gen. et sp. nov

Valid

Badano & Engel in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the stem group of Myrmeleontiformia. The type species is M. electrina.

Macleodiella electrina Macleodiella electrina holotype fig 1 A.jpg
Macleodiella electrina

Mesoptynx [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A relative of antlions and owlflies. The type species is M. unguiculatus.

Mesoptynx unguiculatus Mesoptynx unguiculatus holotype Sup Fig 2 E.jpg
Mesoptynx unguiculatus

Mesypochrysa cannabina [199]

Sp. nov

Valid

Khramov

Early Cretaceous

Flag of Russia.svg  Russia

A green lacewing.

Mesypochrysa naranica [199]

Sp. nov

Valid

Khramov

Early Cretaceous

Flag of Russia.svg  Russia

A green lacewing.

Nilionympha shantouensis [209]

Sp. nov

Valid

Li, Ren & Wang

Middle Jurassic

Jiulongshan Formation

Flag of the People's Republic of China.svg  China

A member of the family Osmylidae.

Nymphavus [200]

Gen. et sp. nov

Valid

Badano, Engel & Wang in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Nymphidae. The type species is N. progenitor.

Nymphavus progenitor Nymphavus progenitor paratype fig 1 A.jpg
Nymphavus progenitor

Oligopsychopsis [210]

Gen. et 2 sp. et comb. nov

Valid

Chang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Psychopsoidea, possibly belonging to the family Kalligrammatidae. [201] Genus includes new species O. penniformis Chang et al. (2018) and O. grandis Liu et al. (2018), as well as "Burmopsychops" groehni Makarkin (2017). [201]

Oligopsychopsis grandis Oligopsychopsis grandis holotype.jpg
Oligopsychopsis grandis

Opapanfilovia [203]

Gen. et sp. nov

Valid

Khramov & Vasilenko

Jurassic

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of the family Panfiloviidae. Genus includes new species O. bonata.

Ovalofemora [207]

Gen. et sp. et comb. nov

Valid

Jepson, Khramov & Ohl

Late Jurassic

Flag of Kazakhstan.svg  Kazakhstan

A member of the family Mantispidae. Genus includes new species O. abbottae, as well as "Mesithone" monstruosa Khramov (2013).

Parababinskaia makarkini [211]

Sp. nov

Valid

Hu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Myrmeleontoidea belonging to the family Babinskaiidae.

Parababinskaia makarkini Parababinskaia makarkini holotype Fig1 A.jpg
Parababinskaia makarkini

Parabaisochrysa [212]

Gen. et sp. nov

Valid

Lu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A green lacewing. Genus includes new species P. xingkei.

Parabaisochrysa xingkei Parabaisochrysa xingkei holotype fig1 A.jpg
Parabaisochrysa xingkei

Phyllochrysa [213]

Gen. et sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Chrysopoidea. The type species is P. huangi.

Pristinofossor [200]

Gen. et sp. nov

Valid

Badano & Engel in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A stem-antlion. The type species is P. rictus.

Pristinofossor rictus Pristinofossor rictus holotype fig 1 A.jpg
Pristinofossor rictus

Pseudosencera [214]

Gen. et sp. nov

Valid

Makarkin, Wedmann & Weiterschan

Late Eocene

Baltic amber

Europe (Baltic Sea coast)

A green lacewing. The type species is P. baltica.

Stictosisyra [215]

Gen. et sp. nov

Valid

Yang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Sisyridae. The type species is S. pennyi.

Stictosisyra pennyi Stictosisyra pennyi fig 1A.jpg
Stictosisyra pennyi

Tragichrysa [216]

Gen. et sp. nov

Valid

Pérez-de la Fuente et al.

Early Cretaceous (early Barremian)

Lebanese amber

Flag of Lebanon.svg  Lebanon

A member of Chrysopoidea of uncertain phylogenetic placement. The type species is T. ovoruptora.

Tragichrysa ovoruptora Tragichrysa ovoruptora Lebanese amber fig4 a.jpg
Tragichrysa ovoruptora

Tyruschrysa [217]

Gen. et sp. nov

Valid

Pérez-de la Fuente et al.

Early Cretaceous (Barremian)

Lebanese amber

Flag of Lebanon.svg  Lebanon

A member of the superfamily Chrysopoidea. The type species is T. melqart.

Tyruschrysa melqart Tyruschrysa melqart holotype Fig2.jpg
Tyruschrysa melqart

Wesmaelius makarkini [218]

Sp. nov

Valid

Yang, Pang & Ren in Yang et al.

Early Miocene

Garang Formation

Flag of the People's Republic of China.svg  China

A brown lacewing belonging to the subfamily Hemerobiinae.

Odonatans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Anglophlebia [219]

Gen. et comb. nov

Valid

Kelly & Nel

Early Jurassic (Sinemurian)

Charmouth Mudstone Formation

Flag of the United Kingdom.svg  United Kingdom

A member of Epiprocta belonging to the superfamily Heterophlebioidea and to the family Anglophlebiidae. The type species is "Liassophlebia" gigantea Zeuner (1962).

Araripegomphus shai [220]

Sp. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A dragonfly.

Burmahemiphlebia hui [221]

Sp. nov

Valid

Zheng & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A damselfly belonging to the family Hemiphlebiidae.

Cretamegaloprepus [222]

Gen. et sp. nov

Valid

Zheng, Nel & Wang in Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A damselfly. Genus includes new species C. zhouae.

Cretapodagrion [223]

Gen. et sp. nov

Valid

Huang, Azar & Nel

Early Cretaceous (BarremianAptian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A damselfly. Genus includes new species C. sibelleae.

Cretastenophlebia jiuquanensis [224]

Sp. nov

Valid

Zheng et al.

Early Cretaceous (Lower Albian)

Zhonggou Formation

Flag of the People's Republic of China.svg  China

A member of the family Stenophlebiidae.

Cymatophlebia yixianensis [225]

Sp. nov

Valid

Zheng et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A dragonfly belonging to the family Cymatophlebiidae.

Erpetogomphus shii [226]

Sp. nov

Valid

Zheng et al.

Early Miocene

La Quinta Formation
(Mexican amber)

Flag of Mexico.svg  Mexico

A dragonfly, a species of Erpetogomphus .

Honghea [227]

Gen. et sp. nov

Valid

Zheng et al.

Early Jurassic

Badaowan Formation

Flag of the People's Republic of China.svg  China

A damsel-dragonfly belonging to the family Campterophlebiidae. Genus includes new species H. xui.

Jurathemis [228]

Gen. et sp. nov

Valid

Huang et al.

Late Jurassic (Oxfordian)

Yangshuzhuang Formation

Flag of the People's Republic of China.svg  China

A damsel-dragonfly belonging to the group Isophlebioptera and the family Selenothemistidae. The type species is J. incompletus.

Lateophlebia [229]

Gen. et comb. nov

Valid

Kelly & Nel

Early Jurassic (Sinemurian)

Flag of the United Kingdom.svg  United Kingdom

A damsel-dragonfly belonging to the family Campterophlebiidae. The type species is "Petrophlebia" anglicanopsis Zeuner (1962).

Madres [230]

Gen. et sp. nov

Valid

Petrulevičius

Eocene (Lutetian)

Flag of Argentina.svg  Argentina

A damselfly belonging to the family Synlestidae. The type species is M. delpueblo.

Paraburmagomphides [231]

Gen. et sp. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A dragonfly. Genus includes new species P. zhaoi.

Paracoryphagrion [232]

Gen. et sp. nov

Valid

Zheng, Nel & Wang in Zheng et al.

Cenomanian

Burmese amber

Flag of Myanmar.svg  Myanmar

A damselfly belonging to the group Pseudostigmatoidea. Genus includes new species P. deltoides.

Paradecoraeshna [233]

Gen. et sp. nov

Valid

Zheng, Nel & Zhang in Zheng et al.

Early Cretaceous (Aptian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A dragonfly belonging to the family Progobiaeshnidae. Genus includes new species P. liaoningensis.

Parazygokaratawia [234]

Gen. et sp. nov

Valid

Huang, Cai & Nel

Middle Jurassic

Daohugou Beds

Flag of the People's Republic of China.svg  China

A damsel-dragonfly belonging to the family Campterophlebiidae. Genus includes new species P. azari.

Proinogomphus kreuzerorum [235]

Sp. nov

Valid

Bechly

Early Jurassic

Flag of Luxembourg.svg  Luxembourg

A dragonfly belonging to the family Liassogomphidae.

Rossiphlebia [219]

Gen. et comb. nov

Valid

Kelly & Nel

Early Jurassic (Sinemurian)

Charmouth Mudstone Formation

Flag of the United Kingdom.svg  United Kingdom

A member of Epiprocta belonging to the superfamily Heterophlebioidea and to the family Liassophlebiidae. The type species is "Liassophlebia" jacksoni Zeuner (1962).

Sinothemis [236]

Gen. et sp. nov

Valid

Huang, Cai & Nel

Late Jurassic (late Oxfordian or early Kimmeridgian)

Tiaojishan Formation

Flag of the People's Republic of China.svg  China

A damsel-dragonfly belonging to the group Isophlebioptera and the family Selenothemistidae. Genus includes new species S. difficilis.

Thairia [237]

Gen. et sp. nov

Valid

Felker & Vasilenko

Early Cretaceous

Flag of Russia.svg  Russia

A damselfly belonging to the family Ponomarenkiidae. Genus includes new species T. transbaikalica.

Valdaeshna mikei [225]

Sp. nov

Valid

Zheng et al.

Early Cretaceous

Upper Weald Clay

Flag of the United Kingdom.svg  United Kingdom

A dragonfly belonging to the family Cymatophlebiidae.

Zygokaratawia incompleta [234]

Sp. nov

Valid

Huang, Cai & Nel

Middle Jurassic

Daohugou Beds

Flag of the People's Republic of China.svg  China

A damsel-dragonfly belonging to the family Campterophlebiidae.

Orthopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Cascadelcana [238]

Gen. et sp. nov

Valid

Fang et al.

Late Triassic (Norian)

Cow Branch Formation

Flag of the United States.svg  United States

A member of the family Elcanidae. Genus includes new species C. virginiana.

Cascogryllus [239]

Gen. et sp. nov

Valid

Poinar

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Tridactylidae. Genus includes new species C. lobiferus.

Elcanonympha [240]

Gen. et sp. nov

Valid

Heads, Thomas & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Elcanidae. Genus includes new species E. diana.

Elcanonympha diana Elcanonympha diana holotype left lateral.jpg
Elcanonympha diana

Ellipes dominicana [239]

Sp. nov

Valid

Poinar

Eocene or Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A member of the family Tridactylidae.

Jeholelcana [241]

Gen. et sp. nov

Valid

Fang et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Orthoptera belonging to the family Elcanidae. Genus includes new species J. yanensis.

Pseudaboilus ningchengensis [242]

Sp. nov

Valid

Wang et al.

Early Cretaceous

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of the family Prophalangopsidae belonging to the subfamily Termitidiinae.

Sigmaboilus calophlebius [243]

Sp. nov

Valid

Wang et al.

Middle–Late Jurassic

Daohugou Beds

Flag of the People's Republic of China.svg  China

An orthopteran belonging to the family Prophalangopsidae.

Plecopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Balticopteryx [244]

Gen. et sp. nov

Valid

Chen

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Taeniopterygidae. Genus includes new species B. dui.

Baltileuctra [245]

Gen. et sp. nov

Valid

Chen

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of Plecoptera belonging to the family Leuctridae. Genus includes new species B. foraminis.

Boreoperlidium callopterus [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Eustheniidae.

Boreoperlidium certus [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Eustheniidae.

Boreoperlidium ramificans [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Eustheniidae.

Electroneuria [247]

Gen. et sp. nov

Valid

Sroka, Staniczek & Kondratieff

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Plecoptera belonging to the family Perlidae. The type species is E. ronwoodi.

Electroneuria ronwoodi Electroneuria ronwoodi holotype Fig5 A.jpg
Electroneuria ronwoodi

Euroleuctra [248]

Gen. et sp. nov

Valid

Chen

Eocene

Baltic amber

Flag of Poland.svg  Poland

A member of Plecoptera belonging to the family Leuctridae. Genus includes new species E. gillesi.

Kargaloperla fibrosa [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Kargaloperla gratum [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Kargaloperla postica [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Kargaloperla queata [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Lapisperla [247]

Gen. et sp. nov

Valid

Sroka, Staniczek & Kondratieff

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Plecoptera belonging to the group Arctoperlaria, assigned to the new family Petroperlidae. The type species is L. keithrichardsi.

Lapisperla keithrichardsi Lapisperla keithrichardsi holotype Fig3 A.jpg
Lapisperla keithrichardsi

Largusoperla [249]

Gen. et 11 sp. nov

Valid

Chen, Wang & Du

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Plecoptera belonging to the family Perlidae. Genus includes new species L. acus Chen, Wang & Du (2018), L. arcus Chen, Wang & Du (2018), L. flata Chen, Wang & Du (2018), L. difformitatem Chen (2018), [250] L. dewalti Chen (2018), L. borisi Chen (2018), [251] L. charliewattsi Sroka, Staniczek & Staniczek (2018), L. brianjonesi Sroka, Staniczek & Staniczek (2018), L. micktaylori Sroka, Staniczek & Staniczek (2018), L. billwymani Sroka, Staniczek & Staniczek (2018) [247] and L. crassus Chen (2018). [252]

Largusoperla billwymani Largusoperla billwymani holotype fig9 A.jpg
Largusoperla billwymani

Palaeonemoura benevola [253]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

Palaeonemourisca corrugata [253]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

Palaeotaeniopteryx alligata [253]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

Palaeotaeniopteryx trinerva [253]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

Paranotonemoura [254]

Gen. et sp. et comb. nov

Valid

Cui & Béthoux in Cui, Ren & Béthoux

Middle Jurassic

Jiulongshan Formation

Flag of the People's Republic of China.svg  China
Flag of Mongolia.svg  Mongolia

A member of Plecoptera belonging to the family Notonemouridae. The type species is P. zwicki; genus also includes "Perlariopsis" fidelis Sinitshenkova (1987).

Petroperla [247]

Gen. et sp. nov

Valid

Sroka, Staniczek & Kondratieff

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Plecoptera belonging to the group Arctoperlaria, assigned to the new family Petroperlidae. The type species is P. mickjaggeri.

Petroperla mickjaggeri Petroperla mickjaggeri holotype Fig1 A.jpg
Petroperla mickjaggeri

Pinguisoperla [255]

Gen. et sp. nov

Valid

Chen

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Plecoptera belonging to the family Perlidae. Genus includes new species P. yangzhouensis.

Properla incrassata [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Properla umbrosa [246]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

A member of the family Palaeoperlidae.

Votaknemoura persuasibil [253]

Sp. nov

Valid

Sinitshenkova

Middle Permian

Flag of Russia.svg  Russia

Trichopterans

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Anisocalamus [256]

Gen. et sp. nov

Valid

Sukacheva & Vasilenko

Probably Early Cretaceous

Doronino Formation

Flag of Russia.svg  Russia

A member of the family Calamoceratidae. Genus includes new species A. mixtus.

Austaulius [257]

Gen. et comb. et sp. nov

Valid

Kelly, Ross & Coram

Late Triassic (Rhaetian) and Early Jurassic (Hettangian and Sinemurian)

Penarth Group

Flag of the United Kingdom.svg  United Kingdom

A member of the family Necrotauliidae. The type species is "Orthophlebia" furcata Giebel (1856); genus also includes new species A. haustrum.

Burmapsyche [258]

Gen. et 2 sp. nov

Valid

Wichard et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Integripalpia belonging to the family Dysoneuridae. Genus includes new species B. comosa and B. palpsfurcata.

Calamostavropolia [259]

Gen. et sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A member of the family Calamoceratidae. Genus includes new species C. revolutionaria.

Cretaganonema [260]

Gen. et sp. nov

Valid

Wichard, Espeland & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Calamoceratidae. The type species is C. dongi.

Cretahelicopsyche [260]

Gen. et sp. nov

Valid

Wichard, Espeland & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A snail-case caddisfly. The type species is C. liuyani.

Cretapsyche [258]

Gen. et 3 sp. nov

Valid

Wichard et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Integripalpia belonging to the family Dysoneuridae. Genus includes new species C. circula,C. elegans and C. insueta.

Ecnomus cretacia [261]

Sp. nov

Valid

Wichard & Azar

Early Cretaceous

Lebanese amber

Flag of Lebanon.svg  Lebanon

A member of the family Ecnomidae.

Electroadicella kuenowi [262]

Sp. nov

Valid

Wichard, Neumann & Werneburg

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of the family Leptoceridae.

Halesus miocenicus [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

Kempia [263]

Gen. et sp. nov

Junior homonym

Sukatcheva in Aristov & Sukatcheva

Late JurassicEarly Cretaceous

Flag of Russia.svg  Russia

A member of the family Philopotamidae. The type species is K. piotri. The generic name is preoccupied by Kempia Mathews (1912), Kempia Kieffer (1913) and Kempia Preston (1913).

Limnephilus antiquastepposus [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A species of Limnephilus .

Limnephilus metakaspievi [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A species of Limnephilus.

Limnephilus parakaspievi [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A species of Limnephilus.

Limnephilus valliculacerasinus [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A species of Limnephilus.

Litholimnephilops [264]

Gen. et sp. nov

Valid

Robinson, Thomas & Heads

Eocene (Ypresian)

Green River Formation

Flag of the United States.svg  United States

A caddisfly of uncertain phylogenetic placement.
Genus includes new species L. yinani.

Mesoviatrix [263]

Gen. et sp. nov

Valid

Sukatcheva in Aristov & Sukatcheva

Late JurassicEarly Cretaceous

Flag of Russia.svg  Russia

A member of the family Philopotamidae. The type species is M. paradoxa.

Miocenocosmoecus [259]

Gen. et sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

A member of the family Limnephilidae. Genus includes new species M. silvacaliginosus.

Palaeorheithrus [256]

Gen. et sp. nov

Valid

Sukacheva & Vasilenko

Probably Early Cretaceous

Doronino Formation

Flag of Russia.svg  Russia

A member of the family Philorheithridae. Genus includes new species P. sibiricus.

Palerasnitsynus furcatis [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus gracilis [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus lepidus [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus spinosus [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus subgrandis [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus subglobolus [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus sukatchevae [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Palerasnitsynus vulgaris [265]

Sp. nov

Valid

Wichard, Müller & Wang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of the family Psychomyiidae.

Phylocentropus succinolebanensis [261]

Sp. nov

Valid

Wichard & Azar

Early Cretaceous

Lebanese amber

Flag of Lebanon.svg  Lebanon

A member of the family Dipseudopsidae.

Polylongaevus [263]

Gen. et sp. nov

Valid

Sukatcheva in Aristov & Sukatcheva

Late JurassicEarly Cretaceous

Flag of Russia.svg  Russia

A member of the family Polycentropodidae. The type species is P. eskovi.

Potamophylax martynovorum [259]

Sp. nov

Valid

Sukatsheva et al.

Middle Miocene

Flag of Russia.svg  Russia

Triaenodes simoni [262]

Sp. nov

Valid

Wichard, Neumann & Werneburg

Eocene

Baltic amber

Europe (Baltic Sea coast)

A member of the family Leptoceridae.

Other insects

NameNoveltyStatusAuthorsAgeUnitType LocationNotesImages

Aiban [266]

Gen. et sp. nov

Valid

Aristov

Middle Triassic (Ladinian)

Madygen Formation

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of Cnemidolestida/Cnemidolestodea (an extinct group of insects of uncertain phylogenetic placement, might be related to plecopterans or orthopterans) belonging to the family Sylvabestiidae. Genus includes new species A. kichineis.

Annulipsyllipsocus [267]

Gen. et 2 sp. nov

Valid

Hakim et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Psocodea belonging to the family Psyllipsocidae. Genus includes new species A. andreneli and A. inexspectatus.

Antiquatortia [268]

Gen. et sp. nov

Valid

Brown & Baixeras in Heikkilä et al.

Miocene (Burdigalian)

La Toca Formation (Dominican amber)

Flag of the Dominican Republic.svg  Dominican Republic

A Tortricidae moth. The type species is A. histuroides.

Antiquatortia histuroides Antiquatortia histuroides AMNH DR8-43 Dominican amber fig A.jpg
Antiquatortia histuroides

Archaemegaptilus blakelyi [269]

Sp. nov

Valid

Beckemeyer & Engel

Carboniferous (Pennsylvanian)

Pottsville Formation

Flag of the United States.svg  United States

A member of Palaeodictyoptera belonging to the family Archaemegaptilidae.

Baltianania [270]

Gen. et sp. nov

Valid

Solis in Heikkilä, Simonsen & Solis

Eocene (Lutetian)

Prussian Formation (Baltic amber)

Flag of Russia.svg  Russia
(Kaliningrad Oblast)

A lepidopteran, probably a member of the family Crambidae. The type species is B. yantarnia.

Carbohymen [271]

Gen. et sp. nov

Valid

Pecharová & Prokop

Late Carboniferous

Mazon Creek fossil beds

Flag of the United States.svg  United States

A member of Megasecoptera belonging to the family Protohymenidae. Genus includes new species C. testai.

Chiastogryllus [272]

Gen. et sp. nov

Valid

Beckemeyer

Early Permian

Wellington Formation

Flag of the United States.svg  United States

A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae. The type species is C. taschi.

Cretaceomachilis longa [273]

Sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Archaeognatha belonging to the family Meinertellidae.

Dinmopsylla [274]

Gen. et sp. nov

Valid

Lambkin

Late Triassic (Norian)

Dinmore fossil insect locality

Flag of Australia (converted).svg  Australia

A member of Paraneoptera belonging to the group Permopsocida and the family Archipsyllidae. Genus includes new species D. semota.

Echmepteryx (Loxopholia) dominicanus [275]

Sp. nov

Valid

Hakim, Huang & Azar

Miocene

Dominican amber

Flag of the Dominican Republic.svg  Dominican Republic

A member of the family Lepidopsocidae, a species of Echmepteryx .

Elasmophasma [276]

Gen. et sp. nov

Valid

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Phasmatodea belonging to the group Euphasmatodea. Genus includes new species E. stictum.

Eopyralis [270]

Gen. et sp. nov

Valid

Simonsen in Heikkilä, Simonsen & Solis

Eocene (Ypresian)

Fur Formation

Flag of Denmark.svg  Denmark

A lepidopteran, probably a member of the family Pyralidae. The type species is E. morsae.

Griphopteron iya [263]

Sp. nov

Valid

Aristov in Aristov & Sukatcheva

Middle Jurassic

Cheremkhovo Formation

Flag of Russia.svg  Russia

A member of Eoblattida belonging to the family Blattogryllidae.

Hexameropsis elongatus [277]

Sp. nov

Valid

Lin et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A mayfly belonging to the family Hexagenitidae.

Iblatta [278]

Gen. et sp. nov

Valid

Aristov

Permian (Kungurian)

Flag of Russia.svg  Russia

A member of Polyneoptera belonging to the group Eoblattida. Genus includes new species I. attrepida.

Issatermes [266]

Gen. et sp. nov

Valid

Aristov

Late Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the new family Issatermitidae. Genus includes new species I. parvus.

Karga [279]

Nom. nov

Valid

Aristov

Middle Permian

Flag of Russia.svg  Russia

A member of Polyneoptera belonging to the group Reculida; a replacement name for Kargalia Aristov (2009).

Kinitocelis sparsella [280]

Sp. nov

Valid

Mey et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Amphiesmenoptera belonging to the group Tarachoptera.

Lepismachilis eocaenica [281]

Sp. nov

Valid

Kaplin & Perkovsky

Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A member of Archaeognatha belonging to the family Machilidae.

Mazonopterum cooperi [282]

Sp. nov

Valid

Prokop et al.

Carboniferous (Pennsylvanian)

Flag of the United Kingdom.svg  United Kingdom

A member of Palaeodictyoptera belonging to the family Homoiopteridae.

Micropermula ceceya [266]

Sp. nov

Valid

Aristov

Late Permian

Flag of Mongolia.svg  Mongolia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Sylvabestiidae.

Myanmarothrips [283]

Gen. et sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A thrips belonging to the family Merothripidae. The type species is M. pankowskiorum.

Nullmeinertellus [273]

Gen. et sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Archaeognatha belonging to the family Meinertellidae. The type species is N. wenxuani.

Paleolepidopterites [268]

Gen. et comb. nov

Valid

Kozlov in Heikkilä et al.

Eocene (Lutetian to Priabonian)

Prussian Formation (Baltic amber)
Florissant Fossil Beds National Monument

Flag of Russia.svg  Russia
(Kaliningrad Oblast)
Flag of the United States.svg  United States

A collective group name for Lepidoptera fossils that cannot be placed with certainty in any known family.
Includes Tortrix? florissantana Cockerell (1907), Tortrix? destructus Cockerell (1916) and "Tortricites" sadilenkoi Kozlov (1988).

Paleolepidopterites florissantanus Paleolepidopterites florissantanus Cockerell 1907 UCNH-HT-no-8579.jpg
Paleolepidopterites florissantanus

Paralepinotus [284]

Gen. et sp. nov

Valid

Azar et al.

Early Eocene

Fushun amber

Flag of the People's Republic of China.svg  China

A member of Psocodea belonging to the group Trogiomorpha and to the family Trogiidae. Genus includes new species P. fushunensis.

Protohymen novokshonovi [271]

Sp. nov

Valid

Pecharová & Prokop

Early Permian

Flag of Russia.svg  Russia

A member of Megasecoptera belonging to the family Protohymenidae.

Proximicorneus [285]

Gen. et sp. nov

Valid

Lin et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A mayfly belonging to the family Prosopistomatidae. Genus includes new species P. rectivenius.

Retortocelis [280]

Gen. et 3 sp. nov

Valid

Mey et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Amphiesmenoptera belonging to the group Tarachoptera. Genus includes new species R. longella, R. minimella and R. tyloptera.

Rohrthrips burmiticus [283]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A thrips belonging to the family Rohrthripidae.

Rohrthrips burmiticus Rohrthrips burmiticus.jpg
Rohrthrips burmiticus

Sauk [263]

Gen. et sp. nov

Valid

Aristov in Aristov & Sukatcheva

Early Jurassic

Sauk-Tan'ga locality

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae. The type species is S. batkenicus.

Say [263]

Gen. et sp. nov

Valid

Aristov in Aristov & Sukatcheva

EarlyMiddle Jurassic

Sogul Formation

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae. The type species is S. kirgizicus.

Shurabia tanga [263]

Sp. nov

Valid

Aristov in Aristov & Sukatcheva

Early Jurassic

Madygen Lagerstätte

Flag of Kyrgyzstan.svg  Kyrgyzstan

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae.

Sibestia [266]

Gen. et sp. nov

Valid

Aristov

Middle Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Sylvabestiidae. Genus includes new species S. nana.

Stenoraphidia [286]

Gen. et comb. et sp. nov

Valid

Lyu, Ren & Liu

Early Cretaceous (Barremian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A snakefly belonging to the family Mesoraphidiidae. Genus includes "Alloraphidia" obliquivenatica Ren (1994) and a new species S. longioccipitalis.

Tillyardembia zalazna [266]

Sp. nov

Valid

Aristov

Late Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Tillyardembiidae.

Tshekardushka [278]

Gen. et sp. nov

Valid

Aristov

Permian (Kungurian)

Flag of Russia.svg  Russia

A member of Polyneoptera belonging to the group Reculida. Genus includes new species T. artenatis.

Udembia [266]

Gen. et sp. nov

Valid

Aristov

Middle Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Tillyardembiidae. Genus includes new species U. udmurtica.

Unimeinertellus [273]

Gen. et 2 sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Archaeognatha belonging to the family Meinertellidae. The type species is U. abundus; genus also includes U. bellus.

Vernooijia [282]

Gen. et sp. et comb. nov

Valid

Prokop et al.

Carboniferous (Pennsylvanian)

Flag of the Netherlands.svg  Netherlands
Flag of the United Kingdom.svg  United Kingdom

A member of Palaeodictyoptera belonging to the family Breyeriidae. Genus includes new species V. sassoonae, as well as "Breyeria" harlemensis Brauckmann & Gröning (1996).

Vokhmia [266]

Gen. et 2 sp. nov

Valid

Aristov

Late Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Sylvabestiidae. Genus includes new species V. pyriformis and V. laxa.

Votyak [266]

Gen. et sp. nov

Valid

Aristov

Middle Permian

Flag of Russia.svg  Russia

A member of Cnemidolestida/Cnemidolestodea belonging to the family Sylvabestiidae. Genus includes new species V. pictus.

Zorotypus (Octozoros) cenomanianus [287]

Sp. nov

Valid

Yin, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Zorotypus (s. str.) denticulatus [287]

Sp. nov

Valid

Yin, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Zorotypus dilaticeps [288]

Sp. nov

Valid

Yin et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Zorotypus hirsutus [289]

Sp. nov

Valid

Mashimo in Mashimo et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Zorotypus oligophleps [290]

Sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Zorotypus robustus [290]

Sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

A member of Zoraptera.

Research

Related Research Articles

Michael S. Engel, FLS, FRES is an American paleontologist and entomologist, notable for contributions to insect evolutionary biology and classification. In connection with his studies he has undertaken field expeditions in Central Asia, Asia Minor, the Levant, Arabia, eastern Africa, the high Arctic, and South and North America, and has published more than 925 papers in scientific journals. Some of Engel's research images were included in exhibitions on the aesthetic value of scientific imagery.

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2019 in paleoentomology is a list of new fossil insect taxa that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

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Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up to the end of 2023 can be found in Ross (2024).

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This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

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References

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  194. 1 2 Agnieszka Soszyńska-Maj; Wiesław Krzemiński; Katarzyna Kopeć; Yizi Cao; Dong Ren (2018). "Large Jurassic scorpionflies belonging to a new subfamily of the family Orthophlebiidae (Mecoptera)". Annales Zoologici. 68 (1): 85–92. doi:10.3161/00034541ANZ2018.68.1.004. S2CID   90298406.
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  198. Agnieszka Soszyńska-Maj; Wiesław Krzemiński; Katarzyna Kopeć; Robert A. Coram (2018). "Worcestobiidae – a new Triassic family of Mecoptera, based on species removed from the family Orthophlebiidae". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 107 (2–3): 145–149. doi:10.1017/S1755691017000160. S2CID   134871834.
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