Aeshnidae

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Aeshnidae
Temporal range: Berriasian to present
Lesser Tasmanian Darner Austroaeschna hardyi female.jpg
Tasmanian darner (Austroaeschna tasmanica)
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Odonata
Infraorder: Anisoptera
Superfamily: Aeshnoidea
Family: Aeshnidae
Leach, 1815 [1]
Genera

See text.

Synonyms
Mangrove darner (Coryphaeschna viriditas), Grand Cayman Mangrove darner (Coryphaeschna viriditas).JPG
Mangrove darner (Coryphaeschna viriditas), Grand Cayman

Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 species. [2] [3]

Contents

The family includes some of the largest dragonflies, and can be over three inches long.

Description

Common worldwide or nearly worldwide genera are Aeshna and Anax . Anax includes some of the largest dragonflies, including the North American A. walsinghami , Hawaiian A. strenuus , European A. imperator and A. immaculifrons , and African A. tristis , but these are all exceeded by another member of the family, the Asian Tetracanthagyna plagiata , which, by both wingspan and weight, is the world's largest dragonfly.

There are 41 North American species in 11 genera in this family. Most European species belong to Aeshna. Their American name "darner" stems from the female abdomens looking like a sewing needle, as they cut into plant stem when they lay their eggs through the ovipositor.

These dragonfly mate in flight. The eggs are deposited in water or close by. The larvae (nymphs or naiads) are generally slender compared to those of other families, with a long and flat extensible lower lip (labium). The larvae are aquatic predators, feeding on other insects and even small fish.

The adults spend large amounts of time in the air and seem to fly tirelessly with their four large and powerful wings. They can fly forwards, backwards, or hover like a helicopter. The wings are always extended horizontally.

Their abdomens are long and thin. Most are colored blue and or green, with black and occasionally yellow. Their large, hemispherical, compound eyes touch in the midline and nearly cover their heads. They have extremely good vision, and are voracious insect predators, using their sharp, biting mouthparts.

All are difficult to catch because of their flying ability and keen sight.

Taxonomy and genera

The name Aeshnidae was first proposed as a family by Elford Leach in 1815. [1]

The spelling of Aeshnidae may have resulted from a printer's error in spelling the Greek Aechma, "a spear". [4] The spelling Aeschnidae has been intermittently used over a period of time, but is now abandoned for the original name Aeshnidae. However, derived genus names (such as Rhionaeschna ) retain the 'sch' spelling, as this is how they were first cited.

The oldest known members of the family are † Gomphaeschna inferna Pritykina, 1977 and † Baissaeschna Pritykina, 1977, both from the earliest Cretaceous (Berriasian) of Buryatia, Russia. [5] Gomphaeschna displays a very ancient divergence from the rest of the genus, and many fossil dragonfly genera appear to be closely related to it. For this reason, some studies have placed it and its fossil relatives in their own family, Gomphaeschnidae. [6] However, the World Odonata Checklist presently retains it in Aeshnidae. [7]

Fossil genera

The following fossil aeshnid genera are known. They have been variously classified in the subfamilies Aeschninae, Allopetalinae or Gomphaeschninae, with the latter two sometimes treated as separate families. [8] [9]

See also

References

  1. 1 2 Leach, W.E. (1815). "Entomology". In Brewster, D. (ed.). The Edinburgh Encyclopedia. Vol. 9 (reprint 1830 ed.). Edinburgh: William Blackburn. pp. 57–172 [136]. doi:10.5962/bhl.title.30911.
  2. Dijkstra, Klaas-Douwe B.; Bechly, Günter; Bybee, Seth M.; Dow, Rory A.; Dumont, Henri J.; Fleck, Günther; Garrison, Rosser W.; Hämäläinen, Matti; Kalkman, Vincent J.; Karube, Haruki; May, Michael L.; Orr, Albert G.; Paulson, Dennis R.; Rehn, Andrew C.; Theischinger, Günther; Trueman, John W.H.; Van Tol, Jan; von Ellenrieder, Natalia; Ware, Jessica (2013). "The classification and diversity of dragonflies and damselflies (Odonata). In: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013)". Zootaxa. 3703 (1): 36–45. doi: 10.11646/zootaxa.3703.1.9 . hdl: 10072/61365 . ISSN   1175-5334.
  3. Paulson, D.; Schorr, M.; Abbott, J.; Bota-Sierra, C.; Deliry, C.; Dijkstra, K.-D.; Lozano, F. "World Odonata List". OdonataCentral. University of Alabama.
  4. "Dragonflies of the Family Aeshnidae in British Columbia" (PDF). Retrieved 25 August 2009.[ permanent dead link ]
  5. 1 2 University~mkk24@njit.edu, Manpreet Kaur Kohli~Rutgers; University~jware@amnh.org, Jessica L. Ware~Rutgers; Bechly~Eberhard-Karls-University~guenter.bechly@smns-bw.de, Günter (2016-03-18). "How to date a dragonfly: Fossil calibrations for odonates". Palaeontologia Electronica. Retrieved 2025-08-21.{{cite web}}: CS1 maint: numeric names: authors list (link)
  6. Jouault, Corentin; Coutret, Baptiste; Konhauser, Kurt O.; Nel, André (2023-01-01). "New odonatans (Odonata: Gomphaeschnidae; Synlestidae) from the Paleocene Paskapoo Formation: systematic and biogeographical implications". Journal of Systematic Palaeontology. 21 (1): 2261457. doi:10.1080/14772019.2023.2261457. ISSN   1477-2019.{{cite journal}}: CS1 maint: article number as page number (link)
  7. Paulson, D.; Schorr, M.; Abbott, J.; Bota-Sierra, C.; Deliry, C.; Dijkstra, K.-D.; Lozano, F. "World Odonata List". OdonataCentral. University of Alabama.
  8. "PBDB Taxon". paleobiodb.org. Retrieved 2025-09-03.
  9. "PBDB Taxon". paleobiodb.org. Retrieved 2025-09-03.
  10. Archibald, S. Bruce; Cannings, Robert A. (2019). "Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America". The Canadian Entomologist. 151 (6): 783–816. doi:10.4039/tce.2019.61. ISSN   0008-347X.
  11. Zheng, Daran; Jarzembowski, Edmund A.; Chang, Su-Chin; Wang, Bo (2016-11-01). "A new true dragonfly (Odonata, Anisoptera, Gomphaeschnaoidini) from mid-Cretaceous Burmese amber". Proceedings of the Geologists' Association. 127 (5): 629–632. doi:10.1016/j.pgeola.2016.07.006. ISSN   0016-7878.
  12. Pinkert, Stefan; Bechly, Günter; Nel, André (2017). "First record of hawker dragonflies from Eocene Baltic amber (Odonata: Anisoptera: Gomphaeschnidae)". Zootaxa. 4272 (2): 263. doi:10.11646/zootaxa.4272.2.7.
  13. 1 2 Zhang, Binglan; Ren, Dong; Pang, Hong (2008-01-10). "New dragonflies (Insecta: Odonata: Gomphaeschnidae) from the Yixian Formation in Inner Mongolia, China". Progress in Natural Science. 18 (1): 59–64. doi:10.1016/j.pnsc.2007.07.005. ISSN   1002-0071.
  14. Petrulevičius, Julián F.; Nel, Andre; Voisin, Jean-François (2010-01-01). "A new genus and species of darner dragonfl y (Aeshnidae: Odonata) from the lower Eocene of Laguna del Hunco, Patagonia, Argentina". Annales de la Société entomologique de France (N.S.). 46 (1–2): 271–275. doi:10.1080/00379271.2010.10697668. ISSN   0037-9271.
  15. Zheng, Daran; Nel, André; Jarzembowski, Edmund A.; Wang, Jun; Zhang, Haichun; Wang, Bo (2019-08-01). "New gomphaeschnid dragonflies (Odonata: Anisoptera: Aeshnoptera) from mid-Cretaceous Burmese amber". Cretaceous Research. 100: 138–144. doi:10.1016/j.cretres.2019.03.027. ISSN   0195-6671.

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