Osmylidae Temporal range: | |
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Porismus strigatus , Black Mountain, Canberra (Australia) | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Neuroptera |
Superfamily: | Osmyloidea |
Family: | Osmylidae Linnaeus, 1758 |
Subfamilies | |
See text |
Osmylidae are a small family of winged insects of the net-winged insect order Neuroptera. The osmylids, also called lance lacewings, stream lacewings [1] or giant lacewings [note 1] , [2] are found all over the world except North and Central America. There are around 225 extant species. [3]
Adult osmylids are small to moderately-sized net-winged insects, with wingspans ranging from 1.4 to 3 cm. Smaller members resemble typical green lacewings, and larger species resemble antlions. Many species, namely those of the type genus Osmylus , have spotted wings. The thin antennae are short. They have two compound eyes, as well as three ocelli in between. Adult osmylids, like green lacewings (some of which are colloquially known as "stinkflies"), have prothoracal glands which produce foul-smelling compounds used to deter would-be predators.
Their larvae are superficially similar to those of spongillaflies (Sisyridae). They have peculiar mouthparts which look like a thin forceps with the ends bending outwards. The body is elongated and slender and terminates in two extensible graspers bearing tiny hooks; these are used to aid in locomotion and to grasp prey. The larvae are associated with damp, mossy habitats and are amphibious. They hunt small invertebrate prey, from which they suck the body fluids with their mouthparts.
The adults are diurnal or crepuscular weak-flying insects which mostly prey on small invertebrates, supplemented with some pollen. Eggs are deposited in damp places, usually near freshwater.
Osmylids are generally placed with the extant families Sisyridae (spongillaflies) and Nevrorthidae within the clade Osmyloidea, which is the second earliest diverging clade of Neuroptera after Coniopterygidae (dustywings). The group also contains Archeosmylidae from the Permian-Triassic and Saucrosmylidae from the Middle Jurassic, both of which are thought to be closely related to Osmylidae. The earliest records of Osmylidae date to the Early Jurassic, some of which are already assignable to extant subfamilies, and were diverse during the Jurassic and Cretaceous. At least 278 species have been described in 25 extant and 38 extinct genera. [4]
The following subfamilies and genera are largely after Winterton et al., 2019 [4]
Eidoporisminae Esben-Petersen, 1917
Gumillinae Navas, 1912
Kempyninae Carpenter, 1943
Mesosmylininae Bode, 1953
Osmylinae Leach, 1815
Porisminae Krüger, 1913
Protosmylinae Krüger, 1913
Spilosmylinae Kruger, 1913
Stenosmylinae Krüger, 1913
Tanyderidae, sometimes called primitive crane flies, are long, thin, delicate flies with spotted wings, superficially similar in appearance to some Tipulidae, Trichoceridae, and Ptychopteridae. Most species are restricted in distribution. They are found in many parts of the world, including North America, South America, Africa, Australia, New Zealand, and various islands in the Pacific Ocean. Adults are usually found hanging from vegetation near streams. Larvae are found either in sandy stream margins or in wet, rotten wood. Fossil species are known.
Mantispidae, known commonly as mantidflies, mantispids, mantid lacewings, mantisflies or mantis-flies, is a family of small to moderate-sized insects in the order Neuroptera. There are many genera with around 400 species worldwide, especially in the tropics and subtropics. Only five species of Mantispa occur in Europe. As their names suggest, members of the group possess raptorial forelimbs similar to those of the praying mantis, a case of convergent evolution.
Bittacidae is a family of scorpionflies commonly called hangingflies or hanging scorpionflies.
The Ptychopteridae, phantom crane flies, are a small family of nematocerous Diptera. Superficially similar in appearance to other "tipuloid" families, they lack the ocelli of the Trichoceridae, the five-branched radial vein of the Tanyderidae, and the two anal veins that reach the wing margins of the Tipulidae. They are usually allied with the Tanyderidae based on similarities of the mesonotal suture, this group being called the Ptychopteromorpha.
Anaxyelidae is a family of incense cedar wood wasps in the order Hymenoptera. It contains only one living genus, Syntexis, which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jurassic, belonging to over a dozen extinct genera, with a particularly high diversity during the Early Cretaceous. Syntexis lay eggs in the sapwood of conifers, preferring recently burnt wood.
Hemerobiidae is a family of Neuropteran insects commonly known as brown lacewings, comprising about 500 species in 28 genera. Most are yellow to dark brown, but some species are green. They are small; most have forewings 4–10 mm long. These insects differ from the somewhat similar Chrysopidae not only by the usual coloring but also by the wing venation: hemerobiids differ from chrysopids in having numerous long veins and forked costal cross veins. Some genera are widespread, but most are restricted to a single biogeographical realm. Some species have reduced wings to the degree that they are flightless. Imagines (adults) of subfamily Drepanepteryginae mimic dead leaves. Hemerobiid larvae are usually less hairy than chrysopid larvae.
The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.
Nymphidae, sometimes called split-footed lacewings, are a family of winged insects of the order Neuroptera. There are 35 extant species native to Australia and New Guinea.
Psychopsidae is a family of winged insects of the order Neuroptera. They are commonly called silky lacewings.
Ithonidae, commonly called moth lacewings and giant lacewings, is a small family of winged insects of the insect order Neuroptera. The family contains a total of ten living genera, and over a dozen extinct genera described from fossils. The modern Ithonids have a notably disjunct distribution, while the extinct genera had a more global range. The family is considered one of the most primitive living neuropteran families. The family has been expanded twice, first to include the genus Rapisma, formerly placed in the monotypic family Rapismatidae, and then in 2010 to include the genera that had been placed into the family Polystoechotidae. Both Rapismatidae and Polystoechotidae have been shown to nest into Ithonidae sensu lato. The larvae of ithonids are grub-like, subterranean and likely phytophagous.
Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now considered to be the closest relatives of clade containing Orussoidea and Apocrita. There are about 7 genera in Karatavitidae.
Pseudopolycentropodidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Middle Triassic (Anisian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Dermapteridae is an extinct family of earwigs known from the Late Triassic to Mid Cretaceous, it is part of the extinct suborder Archidermaptera, alongside Protodiplatyidae and Turanovia. It was first named as a subfamily by Vishniakova in 1980, and elevated to family status by Engel in 2003 without discussion.
Mesoblattinidae is an extinct, problematic family of cockroaches known from the Mesozoic. It was formerly considered a wastebasket taxon for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with Blattidae and Ectobiidae, as well as possibly Blaberidae. The family first appeared by the Early Jurassic. They are considered to represent amongst the oldest groups of modern cockroaches, and like them are thought to have laid oothecae.
Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis.
Praeaulacidae is an extinct family of Mesozoic parasitic wasps in the suborder Evanioidea. It among the earliest known families of the group and is characterised by more complete wing venation in comparison to other members of the suborder. It has been found that Othniodellithidae is nested within Praeaulacidae via cladistic analysis.
Mesochrysopidae is an extinct family of lacewings known from the Jurassic and Cretaceous periods. They are considered to be closely related to green lacewings of the family Chrysopidae. The family are also alternatively considered a paraphyletic grade leading up to crown Chrysopidae.
Necrotauliidae is an extinct family Mesozoic Amphiesmenoptera. While previously considered a paraphyletic grouping of "basal Trichoptera, basal Lepidoptera, and advanced stem-Amphiesmenoptera", they have recently been considered early diverging caddisflies. Other authors have considered them to be basal amphiesiopterans.
Orthophlebiidae is an extinct family of scorpionflies known from the Triassic to Cretaceous, belonging to the superfamily Panorpoidea. The family is poorly defined and is probably paraphyletic, representing many primitive members of Panorpoidea with most species only known from isolated wings, and has such been considered a wastebasket taxon.
Coccolepididae is an extinct family of ray-finned fish, known from the Early Jurassic to Early Cretaceous, most of which were originally referred to the type genus Coccolepis. They had a widespread distribution, being found in North and South America, Australia, Asia and Europe. They are mostly known from freshwater environments, though several species have been found in marine environments. They are morphologically conservative, and have poorly ossified endo and exoskeletons, which usually results in poor preservation. This makes it difficult to distinguish species. They are generally small fish, with the largest known specimens reaching a length of 210 mm. Historically, they have been classified as members of “Palaeonisciformes”, a paraphyletic grouping of non-neopterygian fish, due to their plesiomorphic conservative morphology closely resembling those of many other groups of primitive fish. They have been suggested to be relatives of the Acipenseriformes within the Chondrostei.