Mecoptera is an order of insects in the superorder Holometabola with about six hundred species in nine families worldwide. Mecopterans are sometimes called scorpionflies after their largest family, Panorpidae, in which the males have enlarged genitals raised over the body that look similar to the stingers of scorpions, and long beaklike rostra. The Bittacidae, or hangingflies, are another prominent family and are known for their elaborate mating rituals, in which females choose mates based on the quality of gift prey offered to them by the males. A smaller group is the snow scorpionflies, family Boreidae, adults of which are sometimes seen walking on snowfields. In contrast, the majority of species in the order inhabit moist environments in tropical locations.
Meropeidae is a family of tiny scorpionflies within the order Mecoptera with only three known living species, commonly referred to as "earwigflies", based on the earwig-like forceps-shaped male genitalia. The living species are the North American Merope tuber, the Western Australian Austromerope poultoni, and the recently discovered South American A. brasiliensis. The biology of these species is essentially unknown, and their larvae have never been seen. The adults have been suggested to probably be saprophagous, though they have never been observed feeding. The fossil record of the group extends back to the Jurassic. The earliest known member is Boreomerope antiqua known from an isolated wing found in the Middle Jurassic Itat Formation of Siberia. Other extinct genera include Burmomerope with three species and Torvimerope with one species are both from the mid Cretaceous (Cenomanian) aged Burmese amber. As such, the extant members of this family can be considered living fossils. These insects are also of interest due to their presumed basal position in the order Mecoptera. Thaumatomerope with four described species all from the Madygen Formation in Kyrgyzstan has historically sometimes been included within the family, it was placed into its own monotypic family, "Thaumatomeropidae." in 2002.
Nannochoristidae is a family of scorpionflies with many unusual traits. It is a tiny, relict family with a single extant genus, Nannochorista, with eight species occurring in New Zealand, southeastern Australia, Tasmania, Argentina and Chile. Due to the group's distinctiveness from other scorpionflies, it is sometimes placed in its own order, the Nannomecoptera. Some studies have placed them as the closest living relatives of fleas. Most mecopteran larvae are eruciform, or shaped like caterpillars. Nannochoristid larvae, however, are elateriform, and have elongated and slender bodies. The larvae are aquatic, which is unique among mecopterans. The larvae are predatory, hunting on the beds of shallow streams, primarily on the larvae of aquatic Diptera like chironomids. The adults are thought probably to be adapted to liquid feeding, likely on flower nectar and/or the juice of fruits. Adults of Australian and South American species are often found in habitats like the edges of streams, lakes, as well as montane bogs. Australian species have been observed visiting the foliage and flowers of Leptospermum.
Bittacidae is a family of scorpionflies commonly called hangingflies or hanging scorpionflies.
The Panorpidae are a family of scorpionflies containing more than 480 species. The family is the largest family in Mecoptera, covering approximately 70% species of the order. Species range between 9–25 mm long.
Notiothauma is the sole living genus in the scorpionfly family Eomeropidae. The genus is monotypic with a lone species Notiothauma reedi which is native to the Valdivian temperate rain forests of central Chile, especially the forests with Nothofagus stands. N. reedi is flattened with a notedly cockroach-like appearance. They are nocturnal and inhabit the forest floor where the adults feed on plants and decaying animals. The larvae are still unknown. Because this is the last extant species of Eomeropidae, N. reedi can be characterized as a living fossil taxon.
Anaxyelidae is a family of incense cedar wood wasps in the order Hymenoptera. It contains only one living genus, Syntexis, which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jurassic, belonging to over a dozen extinct genera, with a particularly high diversity during the Early Cretaceous. Syntexis lay eggs in the sapwood of conifers, preferring recently burnt wood.
The Haifanggou Formation, also known as the Jiulongshan Formation, is a fossil-bearing rock deposit located near Daohugou village of Ningcheng County, in Inner Mongolia, northeastern China.
Palaeontinidae, commonly known as giant cicadas, is an extinct family of cicadomorphs. They existed from the Late Triassic to the Early Cretaceous. The family contains around 30 to 40 genera and around a hundred species.
Mongolbittacus is an extinct genus of hangingfly in the family Bittacidae and containing a single species Mongolbittacus daohugoensis. The species is known only from the Middle Jurassic Jiulongshan Formation, part of the Daohugou Beds, near the village of Daohugou in Ningcheng County, northeastern China.
Formosibittacus is an extinct genus of hangingfly in the family Bittacidae and containing a single species Formosibittacus macularis. The species is known only from the Middle Jurassic Jiulongshan Formation, part of the Daohugou Beds, near the village of Daohugou in Ningcheng County, northeastern China.
Jurahylobittacus is an extinct genus of hangingfly in the family Bittacidae and containing a single species Jurahylobittacus astictus. The species is known only from the Middle Jurassic Jiulongshan Formation, part of the Daohugou Beds, near the village of Daohugou in Ningcheng County, northeastern China.
Jurassipanorpa is a genus of fossil scorpionfly containing two species described in 2014 from the Jiulongshan Formation of Inner Mongolia, China. The two species, J. impuctata and J. sticta, lived in the late Middle Jurassic period. Upon description, they were claimed to represent the oldest known representatives of the scorpionfly family Panorpidae, but this was later questioned.
Pseudopolycentropodidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Middle Triassic (Anisian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Aneuretopsychidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Jurassic (Callovian-Oxfordian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops of and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Mesopsychidae is an extinct family of scorpionflies known from the Late Permian to Mid Cretaceous. It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Eodermaptera is an extinct suborder of earwigs known from the Middle Jurassic to Mid Cretaceous. Defining characteristics include "tarsi three-segmented, tegmina retain venation, 8th and 9th abdominal tergite in females are narrowed, but separate from 10th tergite and not covered by 7th tergite and exposed ovipositor" They are considered to be more closely related to Neodermaptera than the more basal Archidermaptera.
Cimbrophlebiidae is an extinct family of scorpionflies. They are considered to be the sister group to the Bittacidae, together forming the clade Raptipedia.
Orthophlebiidae is an extinct family of scorpionflies known from the Triassic to Cretaceous, belonging to the superfamily Panorpoidea. The family is poorly defined and is probably paraphyletic, representing many primitive members of Panorpoidea with most species only known from isolated wings, and has such been considered a wastebasket taxon.
Protorthophlebia is an extinct genus of scorpionflies, known from the Triassic and Jurassic periods of Eurasia. It was originally considered a member of the family Orthophlebiidae, but was later placed as the only genus within the family Protorthophlebiidae within the superfamily Panorpoidea.
