Meropeidae Temporal range: Middle Jurassic to Recent | |
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Merope tuber | |
Austromerope brasiliensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Mecoptera |
Family: | Meropeidae Handlirsch 1906 |
Genera | |
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Meropeidae is a family of tiny scorpionflies within the order Mecoptera with only three known living species, commonly referred to as "earwigflies" (or sometimes "forcepflies"), based on the earwig-like forceps-shaped male genitalia. The living species are the North American Merope tuber , the Western Australian Austromerope poultoni , and the recently discovered South American A. brasiliensis . [1] The biology of these species is essentially unknown, and their larvae have never been seen. [2] The adults have been suggested to probably be saprophagous, though they have never been observed feeding. [3] The fossil record of the group extends back to the Jurassic. The earliest known member is Boreomerope antiqua [4] known from an isolated wing found in the Middle Jurassic Itat Formation of Siberia. Other extinct genera include Burmomerope [5] [6] with three species and Torvimerope with one species are both from the mid Cretaceous (Cenomanian) aged Burmese amber. [7] As such, the extant members of this family can be considered living fossils. These insects are also of interest due to their presumed basal position in the order Mecoptera. Thaumatomerope with four described species all from the Madygen Formation in Kyrgyzstan has historically sometimes been included within the family, it was placed into its own monotypic family, "Thaumatomeropidae." in 2002. [8]
The closest relatives of the family are thought to be Eomeropidae, another relictual family of mecopterans with only 1 living species whose fossil record also extends back to the Jurassic. [7] The family name was spelt "Meropidae" in old literature but this clashes with the homonymous family name in birds for bee-eaters. The spelling of Meropeidae was adopted for the insect family by the ICZN in Opinion 140 of 1943. [9]
Phylogeny after Zhang et al. 2023: [7]
Meropeidae | |
Mecoptera is an order of insects in the superorder Holometabola with about six hundred species in nine families worldwide. Mecopterans are sometimes called scorpionflies after their largest family, Panorpidae, in which the males have enlarged genitals raised over the body that look similar to the stingers of scorpions, and long beaklike rostra. The Bittacidae, or hangingflies, are another prominent family and are known for their elaborate mating rituals, in which females choose mates based on the quality of gift prey offered to them by the males. A smaller group is the snow scorpionflies, family Boreidae, adults of which are sometimes seen walking on snowfields. In contrast, the majority of species in the order inhabit moist environments in tropical locations.
Mantispidae, known commonly as mantidflies, mantispids, mantid lacewings, mantisflies or mantis-flies, is a family of small to moderate-sized insects in the order Neuroptera. There are many genera with around 400 species worldwide, especially in the tropics and subtropics. Only five species of Mantispa occur in Europe. As their names suggest, members of the group possess raptorial forelimbs similar to those of the praying mantis, a case of convergent evolution.
Nannochoristidae is a family of scorpionflies with many unusual traits. It is a tiny, relict family with a single extant genus, Nannochorista, with eight species occurring in New Zealand, southeastern Australia, Tasmania, Argentina and Chile. Due to the group's distinctiveness from other scorpionflies, it is sometimes placed in its own order, the Nannomecoptera. Some studies have placed them as the closest living relatives of fleas. Most mecopteran larvae are eruciform, or shaped like caterpillars. Nannochoristid larvae, however, are elateriform, and have elongated and slender bodies. The larvae are aquatic, which is unique among mecopterans. The larvae are predatory, hunting on the beds of shallow streams, primarily on the larvae of aquatic Diptera like chironomids. The adults are thought probably to be adapted to liquid feeding, likely on flower nectar and/or the juice of fruits. Adults of Australian and South American species are often found in habitats like the edges of streams, lakes, as well as montane bogs. Australian species have been observed visiting the foliage and flowers of Leptospermum.
Bittacidae is a family of scorpionflies commonly called hangingflies or hanging scorpionflies.
The Panorpidae are a family of scorpionflies containing more than 480 species. The family is the largest family in Mecoptera, covering approximately 70% species of the order. Species range between 9–25 mm long.
Eomeropidae is a family of aberrant, flattened scorpionflies represented today by only a single living species, Notiothauma reedi, known from the Nothofagus forests in southern Chile, while all other recognized genera in the family are known only as fossils, with the earliest definitive fossil known from Liassic-aged strata, and the youngest from Paleogene-aged strata.
Nymphidae, sometimes called split-footed lacewings, are a family of winged insects of the order Neuroptera. There are 35 extant species native to Australia and New Guinea.
Merope tuber, the earwigfly or forcepfly, is the only species in the genus Merope, and the only living member of the family Meropeidae in North America. It occurs throughout the east from Ontario to Georgia, and west to Kansas. Recently the insect has also been found in Florida. This insect's most distinguishing feature is the segmented cerci on the male abdomen. Much is unknown about the adults, which are nocturnal and secretive, sometimes found under logs or in malaise traps near streams, or attracted to lights at nighttime. No M. tuber or Meropeid larvae have been identified. The insect is characterized by long wings with many veins and no ocelli. There is a region of interlocking sclerites that holds the jugum and scutellum on the middle thoracic segment together. This may be used to keep the wings together when pushing up through dirt. A similar apparatus is found in cicadas and ground-dwelling beetles, so it may be that the winged adults dig in soil. The flat appearance of the insect suggests that the insect dwells close to the ground in fissures and other small ground openings, as does the lack of ocelli.
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This list of fossil arthropods described in 2013 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2013. The list only includes taxa at the level of genus or species.
This list of fossil arthropods described in 2010 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2010. The list only includes taxa at the level of genus or species.
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Aneuretopsychidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Jurassic (Callovian-Oxfordian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops of and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Elcanidae are an extinct family of Mesozoic and early Cenozoic orthopterans. Members of the family are distinguished by the presence of spurs on the distal part of the metatibia, unique among orthopterans, these have been suggested to have been used for controlling gliding, swimming aids, or for jumping on water. The group combines characteristics from both major groups of orthopterans, with long antennae and nymphal morphology similar to Ensifera, but with wing venation and adult morphology more similar to Caelifera. Elcanidae is part of Elcanoidea, which is thought to have diverged from living orthopterans by the beginning of the Permian, around 300 million years ago. The family also includes Permelcanidae, known from the Early-Late Permian. The relationship of Elcanoidea to Ensifera and Caelifera is currently unresolved. Elcanids are known from the Late Triassic to Paleocene of Eurasia, North and South America. Some members of the group exhibited aposematic coloration. They are thought to have been herbivorous.
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