Permosynidae

Permosynidae Temporal range: 268.0 to 122.46 Ma PreꞒ Ꞓ O S D C P T J K Pg N [1]
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	† Protocoleoptera
Superfamily:	† Permosynoidea
Family:	† Permosynidae Tillyard, 1924
Genera
See text

Permosynidae is an extinct family of beetles known from the Middle Permian to Early Cretaceous. It is sometimes treated as a form family, because all known members were described from fossils of isolated elytra with punctate striae. ^{[2] [3] [4]}

Taxonomy

After: ^[1]

• Alveolacupes Lin, 1983 (also spelled Alveolicupes)
  • Alveolacupes primusLin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
  • Alveolacupes secundesLin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
• Artematopodites Ponomarenko, 1990
  • Artematopodites crispulusYan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites insculptus(Zhang, 1997) – Sangonghe Formation, China, Early Jurassic (Toarcian); Xishanyao Formation, China, Middle Jurassic (Aalenian/Bajocian)
  • Artematopodites latissimusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites latusPonomarenko, 1990 – Byankino Formation, Russia, Late Jurassic (Tithonian)
  • Artematopodites lepidusYan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites leskoviensisYan in Yan and Zhang, 2010 – Leskovskaya Formation, Russia, Early Cretaceous (Barremian)
  • Artematopodites longus(Hong, 1983) – Jiulongshan Formation, China, Middle-Late Jurassic (Callovian/Oxfordian)
  • Artematopodites lozowskiiPonomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
  • Artematopodites majorPonomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Artematopodites maximusYan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites prolixus(Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
  • Artematopodites propinquus(Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
  • Artematopodites shaanbeiensisHong, 1995 – Huachi-Huanhe Formation, China, Early Cretaceous (Valanginian)
• Bistrisyne Lin, 1986
  • Bistrisyne tenuaLin, 1986 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
• Delpuentesyne Martins-Neto and Gallego, 2007
  • Delpuentesyne menendeziMartins-Neto and Gallego, 2007 – Potrerillos Formation, Argentina, Middle Triassic (Ladinian)
• Diarcuipenna Lin, 1986
  • Diarcuipenna bennettitophilaPonomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Diarcuipenna heterosaLin, 1986 – Zijiachong Formation, China, Late Triassic (Norian)
• Dinoharpalus Handlirsch, 1906
  • Dinoharpalus coptoclavoidesPonomarenko, 2015 – Hassberge Formation, Germany, Late Triassic (Carnian)
  • Dinoharpalus latusPonomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
  • Dinoharpalus liasinus(Giebel, 1856) – Cotham Formation, England, Late Triassic (Rhaetian)
  • Dinoharpalus martynoviPonomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Dinoharpalus rugosusPonomarenko, 1985 – Osinovo Formation, Russia, Early-Middle Jurassic (Toarcian/Aalenian)
• Dzeregia Ponomarenko, 1985
  • Dzeregia amplaPonomarenko, 1990 – Mangutskaya Formation, Russia, Early Cretaceous (Berriasian)
  • Dzeregia byrrhoidesPonomarenko, 1990 – Ukurei Formation, Russia, Late Triassic (Lithonian)
  • Dzeregia crassaPonomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Dzeregia juxta(Lin, 1986) – Zijiachong Formation, China, Late Triassic (Norian)
  • Dzeregia lataPonomarenko, 1985 – Makarova Formation, Russia, Early Jurassic (Toarcian)
  • Dzeregia longaPonomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
  • Dzeregia pilulaPonomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Dzeregia platisPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Dzeregia striataPonomarenko, 1985 – Zhargalant Formation, Mongolia, Early Jurassic (Toarcian)
• Hydrobiites Heer, 1865
  • Hydrobiites anglicusHandlirsch, 1906 – Hasfield, England, Late Triassic (Rhaetian)
  • Hydrobiites bellus(Geinitz, 1894) – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites convexusPonomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
  • Hydrobiites crassusPonomarenko, 1985 – Cheremkhovskaya Formation, Russia, Early Jurassic (Toarcian)
  • Hydrobiites dobbertinensisHandlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites giebeliHandlirsch, 1906 – Cotham Formation, England, Late Triassic (Rhaetian)
  • Hydrobiites handlirschiPonomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Hydrobiites liasinus(Giebel, 1856) – Hasfield, England, Late Triassic (Rhaetian)
  • Hydrobiites longusPonomarenko, 1990 – Kutinskaya Formation, Russian, Early Cretaceous (Barremian)
  • Hydrobiites minorPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Hydrobiites minutissimusPonomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Hydrobiites mongolicusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Hydrobiites permianusPonomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Hydrobiites punctatostriatusHandlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites purbeccensis(Giebel, 1856) – Lulworth Formation, England, Early Cretaceous (Berriasian)
  • Hydrobiites sulcatusPonomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Early Cretaceous (Aptian)
  • Hydrobiites tillyardiPonomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
  • Hydrobiites veteranus(Heer, 1852) – Insektenmergel Formation, Switzerland, Early Jurassic (Hettangian)
  • Hydrobiites vladimiriPonomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
• Ischichucasyne Martins-Neto and Gallego, 2005
  • Ischichucasyne cladocostaMartins-Neto and Gallego, 2009 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
  • Ischichucasyne santajuanaensisMartins-Neto and Gallego, 2005 – Santa Juana Formation, Chile, Late Triassic (Carnian)
• Palademosyne Rohdendorf, 1961
  • Palademosyne martynovaeRohdendorf, 1961 – Kazankovo-Markinskaya Formation, Russia, Middle Permian (Wordian)
  • Palademosyne natalensisPonomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
• Permocrossos Martynov, 1932
  • Permocrossos elongatusMartynov, 1932 – Iva-Gora Beds Formation, Russia, Middle Permian (Roadian)
• Permosyne Tillyard, 1924
  • Permosyne affinisTillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne belmontensisTillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne dentataPonomarenko, 2003 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Permosyne elongataPonomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
  • Permosyne mitchelliTillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne rasnitsyniPonomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
• Platycrossos Dunstan, 1923
  • Platycrossos caroliPonomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Platycrossos elongatusPonomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos latusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos ligulatusDunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
  • Platycrossos longusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos loxonicusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos mongolicusPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos ovumPonomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos petalusPonomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos punctatusPonomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos subtumidusDunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
  • Platycrossos tumidus(Tillyard, 1916) – Blackstone Formation, Australia, Late Triassic (Norian)
• Pseudorhynchophora Handlirsch, 1906
  • Pseudorhynchophora convexaPonomarenko, 2008 – Nyadeita Formation, Russia, Early Triassic (Olenekian)
  • Pseudorhynchophora olliffiHandlirsch, 1906 – Blackstone Formation, Australia, Late Triassic (Norian)
• Sakmaracoleus Ponomarenko, 2013
  • Sakmaracoleus orenburgensisPonomarenko, 2013 – Vyazovka Formation, Russia, Middle Permian (Capitanian)
• Tychtocoleus Rohdendorf, 1961
  • Tychtocoleus europaeus(Ponomarenko, 2003) – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Tychtocoleus neuburgaeRohdendorf, 1961 – Gramoteinskaya Formation, Russia, Late Permian (Wuchiapingian)
  • Tychtocoleus popoviPonomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
• Umkomaasia Zeuner, 1960
  • Umkomaasia depressaZeuner, 1960 – Molteno Formation, South Africa, Late Triassic (Carnian)

References

1. "Permosynidae". Paleobiology Database . Retrieved 9 March 2023.
2. Ponomarenko, A.G. (2003). "The First Beetles (Permosynidae, Coleoptera) from the Upper Tatarian of European Russia" (PDF). Paleontological Journal . 37 (2): 170–173.
3. A. G. Ponomarenko, D. S. Aristov, A. S. Bashkuev, Yu. M. Gubin, A. V. Khramov, E. D. Lukashevich, Yu. A. Popov, L. N. Pritykina, S. M. Sinitsa, N. D. Sinitshenkova, I. D. Sukatsheva, D. V. Vassilenko, E. V. Yan (2014). "Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia". Paleontological Journal 48 (14): 1573–1682. doi:10.1134/S0031030114140160.
4. A. G. Ponomarenko (2015). "New beetles (Insecta, Coleoptera) from the Nedubrovo locality, terminal Permian or basal Triassic of European Russia". Paleontological Journal 49 (1): 39–50. doi:10.1134/S0031030115010098.