Capitanian

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Capitanian
264.28 ± 0.16 – 259.51 ± 0.21 Ma
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Chronology
Etymology
Name formalityFormal
Usage information
Celestial body Earth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unit Age
Stratigraphic unit Stage
Time span formalityFormal
Lower boundary definition FAD of the Conodont Jinogondolella postserrata
Lower boundary GSSP Nipple Hill, Guadalupe Mountains, Texas, USA
31°54′33″N104°47′21″W / 31.9091°N 104.7892°W / 31.9091; -104.7892
Lower GSSP ratified2001 [2]
Upper boundary definitionFAD of the Conodont Clarkina postbitteri postbitteri
Upper boundary GSSP Penglaitan Section, Laibin, Guangxi, China
23°41′43″N109°19′16″E / 23.6953°N 109.3211°E / 23.6953; 109.3211
Upper GSSP ratified2004 [3]

In the geologic timescale, the Capitanian is an age or stage of the Permian. It is also the uppermost or latest of three subdivisions of the Guadalupian Epoch or Series. The Capitanian lasted between 264.28 and 259.51 million years ago. It was preceded by the Wordian and followed by the Wuchiapingian. [4]

Contents

A significant mass extinction event occurred at the end of this stage, which was associated with anoxia and acidification in the oceans and possibly caused by the volcanic eruptions that produced the Emeishan Traps. [5] This extinction event may be related to the much larger Permian–Triassic extinction event that followed about 10 million years later.

Stratigraphy

The Capitanian Stage was introduced into scientific literature by George Burr Richardson in 1904. The name comes from the Capitan Reef in the Guadalupe Mountains (Texas, United States). The Capitanian was first used as a stratigraphic subdivision of the Guadalupian in 1961, [6] when both names were still only used regionally in the southern US. The stage was added to the internationally used ICS timescale in 2001. [7]

Definitions

The base of the Capitanian Stage is defined as the place in the stratigraphic record where fossils of conodont species Jinogondolella postserrata first appear. The global reference profile for this stratigraphic boundary is located at Nipple Hill in the southern Guadalupe Mountains of Texas.

The top of the Capitanian (the base of the Wuchiapingian and Lopingian series) is defined as the place in the stratigraphic record where the conodont species Clarkina postbitteri postbitteri first appears.

The Capitanian Stage was part of the time in which the Zechstein was deposited in Europe. [4] It is coeval with the old European regional Saxonian Stage. In the eastern Tethys domain, the Capitanian overlaps the regional Murgabian Stage, the Midian Stage and the lower part of the Laibinian Stage. In Russia the Capitanian equals the lower part of the regional Severodvinian Stage.

Biostratigraphy

The Capitanian contains one ammonite biozone ( Timorites ) and three conodont biozones:

Larger fusulinid species permit a division in two biozones:

Capitanian life

Olson’s Extinction, in the early Guadalupian (Roadian, Wordian), led to an extended period of low diversity when worldwide two-thirds of terrestrial vertebrate life was lost. [8] Global diversity rose dramatically in the Capitanian, probably the result of disaster taxa filling empty guilds, only to fall again when the end-Guadalupian event caused a diversity drop in the Wuchiapingian. [9]

Notable formations

Events

Carbon isotopes in marine limestone from the Capitanian age show an increase in δ13C values. The change in carbon isotopes in the sea water reflects cooling of global climates. [10]

This climatic cooling may have caused the end-Capitanian extinction event among species that lived in warm water, like larger fusulinids (Verbeekninidae), large bivalves (Alatoconchidae) and rugose corals, and Waagenophyllidae. [11]

Related Research Articles

<span class="mw-page-title-main">Permian</span> Sixth and last period of the Paleozoic Era 299–252 million years ago

The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the sixth and last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.

<span class="mw-page-title-main">Guadalupian</span> Second series and epoch of the Permian

The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian.

<span class="mw-page-title-main">Cisuralian</span> First series of the Permian

The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Ma.

<span class="mw-page-title-main">Lopingian</span> Third and final series of the Permian

The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic.

<span class="mw-page-title-main">Anisian</span> Stage of the Triassic

In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from 247.2 million years ago until 242 million years ago. The Anisian Age succeeds the Olenekian Age and precedes the Ladinian Age.

In the geologic timescale, the Kungurian is an age or stage of the Permian. It is the latest or upper of four subdivisions of the Cisuralian Epoch or Series. The Kungurian lasted between 283.5 and 273.01 million years ago (Ma). It was preceded by the Artinskian and followed by the Roadian.

In the geologic timescale, the Roadian is an age or stage of the Permian. It is the earliest or lower of three subdivisions of the Guadalupian Epoch or Series. The Roadian lasted between 273.01 and 266.9 million years ago (Ma). It was preceded by the Kungurian and followed by the Wordian.

In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between 266.9 and 264.28 million years ago (Ma). It was preceded by the Roadian and followed by the Capitanian.

<span class="mw-page-title-main">Wuchiapingian</span> Eighth stage of the Permian

In the geologic timescale, the Wuchiapingian or Wujiapingian is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between 259.51 and 254.14 million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian.

<span class="mw-page-title-main">Changhsingian</span> Ninth and last stage of the Permain

In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from 254.14 to 251.9 Ma ago. It is preceded by the Wuchiapingian age/stage and is followed by the Induan age/stage.

The Emeishan Traps constitute a flood basalt volcanic province, or large igneous province, in south-western China, centred in Sichuan province. It is sometimes referred to as the Permian Emeishan Large Igneous Province or Emeishan Flood Basalts. Like other volcanic provinces or "traps", the Emeishan Traps are multiple layers of igneous rock laid down by large mantle plume volcanic eruptions. The Emeishan Traps eruptions were serious enough to have global ecological and paleontological impact.

The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to 208.5 million years ago. It was preceded by the Carnian and succeeded by the Rhaetian.

<span class="mw-page-title-main">Permocupedidae</span> Extinct family of beetles

Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, Frankencupes ultimus from the Middle Triassic (Anisian) Röt Formation of Germany. They are thought to have been xylophagous, which is presumed to be the ancestral ecology of beetles.

Clarkina is an extinct genus of conodonts. It is considered to be an offshore, outer shelf or basinal, deep-water taxon.

Mesogondolella is an extinct genus of conodonts.

Jinogondolella is an extinct genus of conodonts.

Streptognathodus is an extinct genus of conodonts from the Late Carboniferous to Early Permian.

Conodonts are an extinct class of animals whose feeding apparatuses called teeth or elements are common microfossils found in strata dating from the Stage 10 of the Furongian, the fourth and final series of the Cambrian, to the Rhaetian stage of the Late Triassic. These elements can be used alternatively to or in correlation with other types of fossils in the subfield of the stratigraphy named biostratigraphy.

<span class="mw-page-title-main">Capitanian mass extinction event</span> Extinction event around 260 million years ago

The Capitanian mass extinction event, also known as the end-Guadalupian extinction event, the Guadalupian-Lopingian boundary mass extinction, the pre-Lopingian crisis, or the Middle Permian extinction, was an extinction event that predated the end-Permian extinction event. The mass extinction occurred during a period of decreased species richness and increased extinction rates near the end of the Middle Permian, also known as the Guadalupian epoch. It is often called the end-Guadalupian extinction event because of its initial recognition between the Guadalupian and Lopingian series; however, more refined stratigraphic study suggests that extinction peaks in many taxonomic groups occurred within the Guadalupian, in the latter half of the Capitanian age. The extinction event has been argued to have begun around 262 million years ago with the Late Guadalupian crisis, though its most intense pulse occurred 259 million years ago in what is known as the Guadalupian-Lopingian boundary event.

<span class="mw-page-title-main">Alatoconchidae</span> Extinct family of molluscs

Alatoconchidae is an extinct family of prehistoric bivalves that lived in the early to middle Permian period. Genera belonging to Alatoconchidae are characterized by their shell that is strongly compressed in the dorsoventral direction. Some species reached large sizes of as much as 1 metre (3.3 ft) long. It is hypothesized that some species in this family got energy from chemosynthetic bacteria.

References

  1. ↑ "International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy . September 2023. Retrieved November 10, 2024.
  2. ↑ "GSSP for Roadian Stage". International Commission on Stratigraphy. Retrieved 13 December 2020.
  3. ↑ Jin, Yugan; Shen, Shuzhong; Henderson, Charles; Wang, Xiangdong; Wang, Wei; Wang, Yue; Cao, Changqun; Shang, Qinghua (December 2006). "The Global Stratotype Section and Point (GSSP) for the boundary between the Capitanian and Wuchiapingian Stage (Permian)" (PDF). Episodes. 29 (4): 253–262. doi: 10.18814/epiiugs/2006/v29i4/003 . Retrieved 13 December 2020.
  4. 1 2 Gradstein, F.M.; Ogg, J.G. & Smith, A.G.; 2004: A Geologic Time Scale 2004, Cambridge University Press
  5. ↑ Bond, D. P. G.; Wignall, P. B.; Joachimski, M. M.; Sun, Y.; Savov, I.; Grasby, S. E.; Beauchamp, B.; Blomeier, D. P. G. (2015-04-14). "An abrupt extinction in the Middle Permian (Capitanian) of the Boreal Realm (Spitsbergen) and its link to anoxia and acidification" (PDF). Geological Society of America Bulletin. 127 (9–10): 1411–1421. Bibcode:2015GSAB..127.1411B. doi: 10.1130/B31216.1 . ISSN   0016-7606.
  6. ↑ Glenister, B.F. & Furnish, W.M.; 1961: The Permian ammonoids of Australia, Journal of Paleontology 35(4), pp 673–736.
  7. ↑ Glenister, B.F.; Wardlaw, B.R.; Lambert, L.L.; Spinosa, C.; Bowring, S.A.; Erwin, D.H.; Menning, M. & Wilde, G.L.; 1999: Proposal of Guadalupian and Component Roadian, Wordian and Capitanian Stages as International Standards for the Middle Permian Series, Permophiles 34: pp 3–11.
  8. ↑ Bond, David; Hilton, Jason (2010). "The Middle Permian (Capitanian) mass extinction on land and in the oceans". Earth-Science Reviews. 102 (1): 100–116. Bibcode:2010ESRv..102..100B. doi:10.1016/j.earscirev.2010.07.004.
  9. ↑ Sahney, S.; Benton, M.J. (2008). "Recovery from the most profound mass extinction of all time". Proceedings of the Royal Society B: Biological Sciences. 275 (1636): 759–65. doi:10.1098/rspb.2007.1370. PMC   2596898 . PMID   18198148.
  10. ↑ Isozaki, Yukio; Kawahata, Hodaka; Ota, Ayano (2007). "A unique carbon isotope record across the Guadalupian–Lopingian (Middle–Upper Permian) boundary in mid-oceanic paleo-atoll carbonates: The high-productivity "Kamura event" and its collapse in Panthalassa". Global and Planetary Change. 55 (1–3): 21–38. Bibcode:2007GPC....55...21I. doi:10.1016/j.gloplacha.2006.06.006.
  11. ↑ Isozaki, Yukio; Aljinović, Dunja (2009). "End-Guadalupian extinction of the Permian gigantic bivalve Alatoconchidae: End of gigantism in tropical seas by cooling". Palaeogeography, Palaeoclimatology, Palaeoecology. 284 (1–2): 11–21. Bibcode:2009PPP...284...11I. doi:10.1016/j.palaeo.2009.08.022. ISSN   0031-0182.

31°54′33″N104°47′21″W / 31.9091°N 104.7892°W / 31.9091; -104.7892

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