Aguja Formation | |
---|---|
Stratigraphic range: Lower to Middle Campanian ~ | |
Type | Geological formation |
Unit of | Tornillo Group |
Sub-units | La Basa Sandstone Member, Rattlesnake Mountain Sandstone Member, Terlingua Creek Sandstone Member, Abajo Shale Member, and Alto Shale Member |
Underlies | Javelina Formation |
Overlies | Pen Formation |
Lithology | |
Primary | Sandstone, conglomerate, claystone |
Other | Mudstone, shale, limestone |
Location | |
Coordinates | 29°18′N103°30′W / 29.3°N 103.5°W |
Approximate paleocoordinates | 35°48′N77°00′W / 35.8°N 77.0°W |
Region | Texas Chihuahua Coahuila |
Country | USA Mexico |
The Aguja Formation is a geological formation in North America, exposed in Texas, United States and Chihuahua and Coahuila in Mexico, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation. [1] Fossil palms have also been unearthed here. [2]
The ages of the Aguja Formation and its primary fossil-bearing unit, the Alto Shale, are not well understood. Due to the presence of the ammonite Baculites mclearni , which only occurs from 80.67 - 80.21 Ma, in the underlying Rattlesnake Mountain Sandstone and the Terlingua Creek Sandstone, it is likely that the Upper Shale was younger than 80.2 Ma. [3] A radiometric date of 76.9 Ma was recovered in the Alto Shale, making it likely the formation wasn't younger than 76.9 Ma. [3] The contact with the overlying Javelina Formation has been estimated at about 70 Ma ago [4] but also as recently as 68.5 million years ago. [5] This is unlikely, however, due to the presence of Bravoceratops, more primitive than an unnamed chasmosaurine from the De-na-zin Member of the Kirtland Formation, in the lowermost section of the formation. [6] The age of the La Basa Sandstone is constrained by the presence of Scaphites hippocrepis III in the overlying Pen Formation which has been dated as old as 81.53 Ma. [3] [7]
2 fragmentary caudal vertebrae of indeterminate reptiles are known from the Rattlesnake Mountain sandstone member. [8]
Ornithischians of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member [9] | Material | Notes | Images |
Agujaceratops | A. mariscalensis | Texas [10] | Lower Alto Shale | Formerly considered a species of Chasmosaurus [10] | ||
A. mavericus | West Texas [11] | Middle Alto Shale | A chasmosaurine. | |||
Angulomastacator | A. daviesi | Texas [12] | Middle Alto Shale | A left maxilla. [12] | A lambeosaurine. | |
Ankylosauridae | Indeterminate | Texas [13] | Lower Alto Shale | Osteoderms, vertebrae & limb elements. [13] | Remains of an ankylosaurid, possibly represents Euoplocephalus sp. [13] | |
Aquilarhinus [14] | A. palimentus | Texas [7] | Middle Abajo Shale | Partial skeleton | Formerly referred to Kritosaurus and Gryposaurus. [15] | |
Ceratopsidae genus & species indeterminate | Indeterminate | West Texas [16] | Middle Abajo Shale | Ilia, sacral vertebra & sacral ribs. [16] | May represent Agujaceratops , but undiagnostic. | |
Chasmosaurus [17] | C. mariscalensis [18] | Texas [18] | [Twelve] disarticulated skull (sic), postcrania, juvenile." [19] | Considered by paleontologists Lucas, Sullivan, and Hunt to be distinct enough from the Chasmosaurus type species, C. belli to warrant being split off to a new genus, Agujaceratops. [10] | ||
Edmontonia | E. sp [20] | Texas [13] | Lower Alto Shale | Osteoderm (TVP 45866-2) & skull (AMNH 3076). [13] | A nodosaurid. | |
cf. Euoplocephalus [18] | E. sp [18] | Texas [13] | Lower Alto Shale | Osteoderms, sacrum & vertebra (TL-05-14). [13] | An ankylosaurid. | |
aff. Kritosaurus | K. navajovius | Texas [15] | Upper Alto Shale | Two dentary teeth | Tooth crown morphology matches with Kritosaurus [15] | |
Malefica | M. deckerti | Texas [21] | Middle Alto Shale | A partial left maxilla. [21] | Formerly referred to Kritosaurus. [21] | |
Nodosauridae genus & species indeterminate | Indeterminate | West Texas [16] | Middle Abajo Shale | Isolated & associated osteoderms. [16] | Very similar to osteoderms of Invictarx . | |
Panoplosaurus | P. mirus | Texas [13] | Lower Alto Shale | Osteoderm (TMM 45605-4). [13] | A nodosaurid. | |
Saurolophinae | Indeterminate | Texas [15] | Alto Shale | Various cranial and appendicular elements | Material from several localities previously referred to Kritosaurus but do not have its diagnostic features. [15] | |
Saurolophinae | Indeterminate | Texas | Alto Shale | Various manual elements and vertebrae representing two individuals | Included in phylogenetic analyses as the "Big Bend OTU", most recently considered a kritosaurin [21] | |
Stegoceras | S. sp. | West Texas [22] | Middle Alto Shale | Frontal [22] | A pachycephalosaur, represents a new unnamed species | |
Texacephale | T. langstoni | Texas [23] | Lower Alto Shale | Two frontoparietal domes. [23] | A pachycephalosaur. | |
Yehuecauhceratops | Y. mudei [24] | Coahuila [25] | A centrosaurine. | |||
Indeterminate ornithomimid remains are known from the Upper Aguja Formation. [26] Indeterminate tyrannosaurid fossils are known from the Upper Aguja Formation of Texas and Mexico. [27]
Theropods of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member [9] | Material | Notes | Images |
Avialae incertae sedis | West Texas [28] | Middle Abajo Shale [28] | 4 teeth (TMM 45947-349, 350, 351). [28] | A bird. | ||
Chirostenotes | C. sp. | Texas [29] | Femur & manual ungual. [29] | A caenagnathid | ||
Dromaeosauridae genus & species indeterminate | Indeterminate | West Texas [16] | Middle Abajo Shale | Pedal phalanges (TMM 45909-2, TMM 44066-4). [16] | A dromaeosaur. | |
cf. Dromaeosaurus [30] | Indeterminate [30] | Texas [18] | ||||
Leptorhynchos | L. gaddisi | Texas [29] | Lower Alto Shale | Dentaries, caudal vertebra & limb elements. [29] | A caenagnathid | |
Ornithomimidae | New genus & species | West Texas [16] | Middle Abajo Shale | Vertebrae, ischium & limb elements. [16] | An unnamed species referred to informally as the 'Aguja ornithomimid'. | |
cf. Paronychodon | West Texas [28] | Middle Abajo Shale [28] | 2 incomplete teeth (TMM 45947-362). [28] | A maniraptoran. | ||
Richardoestesia | R. cf. gilmorei | Texas [31] | Low Upper Shale [31] | Fragment of a small tooth. [31] | A coelurosaur. | |
R. isosceles | Big Bend National Park, Texas [31] | Low Upper Shale [31] | Teeth. [31] | A coelurosaur. | ||
Saurornitholestes | S. cf. langstoni | Texas [28] [31] | Lower Alto Shale & Middle Abajo Shale [28] [31] | Teeth. [28] [31] | A dromaeosaur. | |
S. langstoni | Coahuila [32] | Teeth [32] | ||||
Saurornitholestes sp. | Coahuila [32] | Teeth [32] | ||||
Theropoda incertae sedis | Morphotype A | West Texas [28] | Alto Shale [28] | 8 teeth. [28] | Unserrated, recurved teeth. | |
Morphotype B | West Texas [28] | 5 tooth crowns. [28] | Bi-serrated teeth. | |||
Morphotype C | West Texas [28] | 7 teeth. [28] | Finely serrated, distal plication only. | |||
Morphotype D | West Texas [28] | 5 teeth. [28] | Short, coarsely serrated distal plication. | |||
Morphotype E | West Texas [28] | A tooth. [28] | Strongly recurved, serrated distal plication. | |||
Morphotype F | West Texas [28] | 2 tooth fragments. [28] | Medium size, finely serrated plications. | |||
cf. Troodon [33] | Indeterminate [33] | Texas [18] | ||||
Tyrannosauridae | Indeterminate | West Texas [28] [34] | Lower Alto Shale [34] & Middle Abajo Shale [16] [28] | Isolated teeth & a handful of non-dental specimens. [28] [34] | Postcranial remains suggest a relatively small & gracile tyrannosaurid, [34] considered a member of the Teratophoneini [35] | |
Indeterminate titanosaurid remains are known from the Upper Aguja Formation of Mexico. [36]
Sauropods of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Titanosauridae | Indeterminate | Chihuahua | Four partial posterior caudal vertebrae & dorsal or sacrocaudal vertebra | A relatively massive titanosaurid. | ||
Crocodylomorphs of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member [9] | Material | Notes | Images |
Crocodilia | Indeterminate | Ten Bits Ranch, west Texas. [8] | Rattlesnake Mountain sandstone member. [8] | Teeth. [8] | A crocodilian. | |
Deinosuchus [37] | D. riograndensis | Texas, [37] Chihuahua, and Coahuila [38] | Upper Shale | Multiple partial skulls & skeletons accounting for nearly the entire skeleton minus the tail. | A giant alligatoroid. | |
cf. D. sp. | West Texas [16] | Middle Abajo Shale [16] | Cervical osteoderm (TMM 44068-2). [16] | An alligatoroid. | ||
Goniopholididae | New genus & species | West Texas [16] | Middle Abajo Shale [16] | Partial skull & skeleton along with isolated osteoderms & teeth. [16] | May pertain to Denazinosuchus or a related taxon. | |
Phobosuchus [37] | P. riograndensis [37] | Texas, [37] Chihuahua, and Coahuila | Reclassified as a Deinosuchus species | |||
Testudines of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Adocus | A. sp. | Big Bend National Park, Texas. [39] | Upper Shale [39] | Shell fragments. [39] | An adocid. | |
Baenidae | Big Bend National Park, Texas. [39] | Upper Shale [39] | Shell fragments. [39] | 2 morphotypes present (A & B). | ||
Basilemys | B. sp. | Big Bend National Park, Texas. [39] | Upper Shale & Lower Shale [16] [39] | Shell fragments, partial plastron & leg scutes. [16] [39] | A nanhsiungchelyid. | |
Bothremydidae genus & species indeterminate | Indeterminate | West Texas [16] | Lower Shale [16] | Isolated peripheral bones (TMM 44068-1, 42452-8); costal bones (TMM 44064-6). [16] | A bothremydid, compatible with Chupacabrachelys . | |
Chupacabrachelys | C. complexus | Big Bend, Texas. [40] | Base of the Upper Shale [40] | A complete skull, and a nearly complete skeleton. [40] | A bothremydid. | |
cf. Denazinemys | cf. D. sp. | West Texas [16] | Lower Shale [16] | Shell elements. [16] | A baenid. | |
cf. Helopanoplia | Big Bend National Park, Texas. [39] | Upper Shale [39] | Shell fragments. [39] | A softshell turtle. | ||
cf. Hoplochelys | Big Bend National Park, Texas. [39] | Upper Shale [39] | Shell fragments. [39] | A kinosternoid. | ||
Terlinguachelys | T. fischbecki | Big Bend National Park, Texas [41] | Rattlesnake Mountain Sandstone [41] | A large, incomplete specimen. [41] | A protostegid. | |
Testudines indeterminate [8] | Morphotype 1 [8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | A shell fragment. | ||
Morphotype 2 [8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | A shell fragment. | |||
Morphotype 3 [8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | A shell fragment. | |||
Morphotype 4 [8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | A shell fragment. | |||
Trionychidae | Gen. et. sp. indet. | Big Bend National Park, Texas. [39] | Upper Shale & Lower Shale [16] [39] | Costal bone (TMM 44068-4) & shell fragments. [39] | A softshell turtle. | |
Lepidosaurs of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Anguidae | Gen. et. sp. indet. | Texas. [42] | Isolated osteoderms & partial right frontal. [42] | An anguid. | ||
Apsgnathus | A. triptodon | Brewster County, Texas. [42] | Jaw elements. [42] | A scincomorph. | ||
Catactegenys | C. solaster | Brewster County, Texas. [42] | Jaw elements & teeth. [42] | A night lizard. | ||
Dryadissector | D. shilleri | West Texas. [28] | Middle Abajo Shale | Numerous isolated teeth. [28] | A varanoid. | |
Mosasauridae | Indeterminate | Ten Bits Ranch. [8] | Rattlesnake Mountain sandstone member. [8] | 1 partial vertebra. [8] | A mosasaur. | |
Odaxosaurus | O. piger | Brewster County, Texas. [42] | Jaw elements. [42] | An anguid. | ||
cf. Parasaniwa | cf. P. wyomingensis | Texas. [42] | Jaw fragments. [42] | A platynotan. | ||
Platynota | Gen. et. sp. indet. | Texas. [42] | Dorsal vertebra (TMM 43057-332). [42] | A platynotan. | ||
?Scincidae | Gen. et. sp. indet. | Brewster County, Texas. [42] | Jaw elements. [42] | A possible skink. | ||
cf. Scincomorpha | Gen. et. sp. indet. | Brewster County, Texas. [42] | Jaw elements. [42] | A scincomorph. | ||
Serpentes | Gen. et. sp. indet. | Texas. [42] | Partial left dentary & right maxilla. [42] | A snake. | ||
cf. Xenosauridae | Texas. [42] | Osteoderms & maxillae. [42] | A knob-scaled lizard. | |||
Mammals of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Alphadon [43] | A. perexiguus | Brewster County, Texas | Upper Shale | A metatherian. | ||
Paleomolops [43] | P. langstoni | Brewster County, Texas | Upper Shale | A tribosphenidan; "cannot be confidently allied with either marsupials or eutherians". [43] | ||
A diversity of bony fish, comprising both marine (Ten Bits/Rattlesnake Mountain locality) and freshwater (Lowerverse/Lower Shale locality) taxa, is known. [44] Approximately 75 whole and broken fragments of coprolites are known from the Rattlesnake Mountain sandstone member, presumably from bony fish. [8]
Bony fish of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Acanthomorpha indet. | family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An acanthomorph. | |
Acanthopterygii indet. | order indet. family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An acanthopterygian. | |
Acanthopterygian fin spine morph 1 | Fin spines | |||||
Acanthopterygian fin spine morph 2 | ||||||
Acanthopterygian fin spine morph 3 | ||||||
Acanthopterygian fin spine morph 4 | ||||||
Euacanthopterygian fin spine morph 1 | ||||||
Euacanthopterygian fin spine morph 2 | ||||||
Albula | A. sp. | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Over 390 complete and fragmentary teeth. [45] | A bonefish. | |
Lowerverse, west Texas [44] | Lower Shale | |||||
Amiidae indet. | genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An amiid. | |
? Anomoeodus | ?A. sp. | Lowerverse, west Texas [44] | Lower Shale | Isolated tooth crown | A pycnodont. Likely transported out of a marine habitat. | |
Atractosteus | A. sp. | Lowerverse, west Texas [44] | Lower Shale | Teeth, jaw fragments, vertebral centra, scales. | A gar. | |
Clupeiformes indet. | family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | A clupeiform. | |
cf. Cyclurus | cf. C. sp. | Lowerverse, west Texas [44] | Lower Shale | 5 fragmentary tooth plates with teeth. | An amiid. | |
Ellimmichthyiformes indet. | family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An ellimmichthyiform. | |
Elopiformes indet. | family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra | An elopiform. | |
?Enchodus | ?E. sp. | Ten Bits ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 2 abraded teeth. [45] | An aulopiform. | |
Lowerverse, west Texas [44] | Lower Shale | 4 isolated teeth. [44] | ||||
Eotexachara | E. malateres | Lowerverse, west Texas [46] | Lower Shale | Dentaries. [46] | A characiform. | |
?Gonorynchiformes indet. | family indet. genus et sp. indet. | Lowerverse, west Texas [46] | Lower Shale | Cranial bone, vertebral centrum, basibranchial. | A gonorynchiform. | |
Hiodontidae indet. | genus et sp. indet. | Lowerverse, west Texas [46] | Lower Shale | Vertebral centra. | A hiodontiform. | |
Lepidotes | ?L. sp. | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Approximately 109 complete and fragmentary teeth. [45] | A lepidotid. | |
cf. Melvius | cf. M. sp. | Lowerverse, west Texas [16] [44] | Lower Shale [16] | Vertebrae, 13 isolated teeth. [16] [44] | An amiid. | |
Micropycnodon | M. sp. | Lowerverse, west Texas [44] | Lower Shale | 4 isolated teeth | A pycnodont. Likely transported out of a marine habitat. | |
Osteichthyes indet. | Indeterminate species A | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | 11 complete teeth. | A bony fish. | |
Indeterminate species B | 8 complete and fragmentary teeth. | |||||
Indeterminate species C | 1 complete tooth. | |||||
Indeterminate species D | 1 complete and 1 partial tooth. | |||||
Indeterminate | Approximately 475 complete and fragmentary teleost centra. [45] | |||||
Tooth morph 1 | Lowerverse, west Texas [44] | Lower Shale | Teeth | |||
Tooth morph 2 | ||||||
Centrum morph 1 | Vertebral centra. | |||||
Centrum morph 2 | ||||||
Centrum morph 3 | ||||||
Centrum morph 4 | ||||||
Ostariophysi indet. | order indet. family indet. genus et sp. indet. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An ostariophysian. | |
Paralbula | P. cf. casei | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Approximately 900 complete and fragmentary teeth and tooth caps. [45] | A bonefish. Lowerverse specimens likely transported out of a marine habitat. | |
Lowerverse, west Texas [44] | Lower Shale | |||||
Primuluchara | P. laramidensis | Lowerverse, west Texas [46] | Lower Shale | Dentaries. [46] | A characiform. | |
Stephanodus | ?S. sp. | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 21 whole and fragmentary specimens. [45] | A pycnodont. | |
? Wilsonichthys | ?W. sp. | Lowerverse, west Texas [44] | Lower Shale | Vertebral centra. | An osteoglossiform. | |
Cartilaginous fish of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Brachyrhyzodus | B. wichitaensis | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 11 complete teeth. [45] | ||
Cantioscyllium | C. aff. meyeri | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 1 anterior tooth and 5 lateral teeth. [8] | A nurse shark. | |
Chiloscyllium | C. aff. greeni | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] | Approximately 90 abraded and fragmentary teeth. [45] | A bamboo shark. | |
Chondrichthyes [8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | 23 placoid scales and 69 dermal scales. | 4 morphotypes of placoid scales (A to D) present. | ||
Columbusia | C. sp. | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 20 complete & fragmentary teeth. [45] | A wobbegong. | |
Cretalamna | C. appendiculata | Ten Bits Ranch, west Texas. [8] | Rattlesnake Mountain sandstone member. [8] | 3 fragmentary teeth. [8] | Reassigned to C. cf. C. sarcoportheta. [45] | |
C. cf. C. sarcoportheta | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 3 incomplete teeth & fragments of additional teeth. [45] | Originally reported as C. appendiculata. | ||
Cretorectolobus [8] | C. olsoni | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | "20 complete and fragmentary teeth". | A carpet shark. | |
Hybodontidae genus & species indeterminate | Indeterminate | West Texas [16] | Lower Shale [16] | Fragment of a dorsal fin spine (TMM 42536-10). [16] | A hybodont. | |
Hybodus [8] | H. sp. | Ten Bits Ranch. | Rattlesnake Mountain sandstone member. [8] | 2 specimens, one complete and one partial tooth. | A hybodont. | |
Igdabatis | I. indicus? | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | A single incomplete tooth. [8] [45] | A myliobatid. | |
Ischyrhiza | I. cf. avonicola | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 5 complete and fragmentary rostral teeth. [45] | A sawskate. | |
I. mira | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 11 fragmentary rostral teeth and 230 whole and fragmentary oral teeth. [45] | A sawskate. | ||
Lonchidion | L. selachos | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 4 complete and 5 fragmentary teeth. [8] [45] | A hybodont. | |
Meristodon | M. sp. | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 2 teeth. [45] | A hybodont. | |
Myliobatiformes | Incertae sedis | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 3 complete specimens. [45] | ||
Protoplatyrhina | P. renae | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] | 70 complete and fragmentary teeth. [45] | A hypsobatid. | |
Ptychotrygon | P. agujaensis | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Over 690 complete and fragmentary teeth. [45] | A sawskate. | |
P. triangularis | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Over 170 complete and fragmentary teeth. [45] | A sawskate. | ||
P. aff. cuspidata | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | 7 whole and fragmentary teeth. [45] | A sawskate. | ||
P. sp. | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | One complete tooth (TMM 46018-71). [45] | A sawskate. | ||
Rhinobatos | R. casieri | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | "14 complete and fragmentary specimens". | A guitarfish. | |
R. sp. | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 18 complete & fragmentary specimens. [45] | A guitarfish. | ||
Rhombodus | R. levis | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Over 22 complete and fragmentary teeth. [45] | A rajiforme. | |
Scapanorhynchus | S. texanus | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | Over 800 complete and fragmentary teeth. [45] | A mitsukurinid. | |
Sclerorhynchidae | Morphotype 1 | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [8] | One fragmentary rostral spine (TMM 46018-59). [45] | A sawskate. | |
Morphotype 2 | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [8] | One fragmentary rostral spine (TMM 46018-60). [45] | A sawskate. | ||
Morphotype 3 | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [8] | 3 fragmentary rostral teeth. [45] | A sawskate. | ||
Morphotype 4 | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [8] | 27 fragmentary rostral spines. [45] | A sawskate. | ||
Serratolamna | S. cf. S. caraibaea | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | About 34 teeth. [45] | A mackerel shark. | |
Squalicorax | S. kaupi | Ten Bits Ranch, west Texas. [8] [45] | Rattlesnake Mountain sandstone member. [8] [45] | "26 complete and fragmentary anterior and lateral teeth". | An anacoracid. | |
S. aff. S. lindstromi | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 4 specimens. [45] | An anacoracid. | ||
S. pristodontus | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | 2 specimens. | An anacoracid. | ||
S. aff. S. yangaensis | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 4 complete & several fragmentary teeth. [45] | An anacoracid. | ||
Squatina | S. hassei | Ten Bits Ranch, west Texas. [8] | Rattlesnake Mountain sandstone member. [8] | A single complete specimen. | An angelshark. | |
S. sp. | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | A complete tooth & 2 fragmentary teeth. [45] | An angelshark. | ||
Texatrygon | T. cf. T. copei | Ten Bits Ranch, west Texas. [45] | Rattlesnake Mountain sandstone member. [45] | 4 complete & fragmentary teeth. [45] | A sawskate formerly reported as T. hooveri. | |
T. hooveri | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member. [8] | 4 complete and fragmentary specimens. [8] | Reassigned to T. cf. T. copei. |
Ammonites of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Abundance | Notes | Images |
Baculites | B. mclearni |
| ||||
Hoplitoplacenticeras | H. plasticum | Rattlesnake Mountain Sandstone | ||||
Pachydiscus | P. paulsoni | Rattlesnake Mountain Sandstone | ||||
Plants of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Sabal [2] | S. bigbendense | Big Bend National Park, Texas. | Upper Shale | 4 seeds. | A palm. It has the largest seeds of any fossil Sabal. [2] | |
S. bracknellense | Big Bend National Park, Texas. | Upper Shale | 5 seeds. | A palm. The seeds are indistinguishable from those of Eocene S. bracknellense, and so were assigned to that species. However, it is likely these Aguja palms would be found to represent distinct species if other parts of the plants were available for comparison. [2] | ||
Color key
| Notes Uncertain or tentative taxa are in small text; |
Gryposaurus was a genus of duckbilled dinosaur that lived about 80 to 75 million years ago, in the Late Cretaceous of North America. Named species of Gryposaurus are known from the Dinosaur Park Formation in Alberta, Canada, and two formations in the United States: the Lower Two Medicine Formation in Montana and the Kaiparowits Formation of Utah. A possible additional species from the Javelina Formation in Texas may extend the temporal range of the genus to 66 million years ago.
Kritosaurus is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard", but is often mistranslated as "noble lizard" in reference to the presumed "Roman nose".
Alamosaurus is a genus of titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. It is the only known titanosaur to have inhabited North America after the nearly 30-million year absence of sauropods from the North American fossil record and probably represents an immigrant from South America.
The Lance (Creek) Formation is a division of Late Cretaceous rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age, and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta.
The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between 82.4 Ma and 74.4 Ma, during Campanian time. It crops out to the east of the Rocky Mountain Overthrust Belt, and the western portion of this formation is folded and faulted while the eastern part, which thins out into the Sweetgrass Arch, is mostly undeformed plains. Below the formation are the nearshore deposits of the Virgelle Sandstone, and above it is the marine Bearpaw Shale. Throughout the Campanian, the Two Medicine Formation was deposited between the western shoreline of the Late Cretaceous Interior Seaway and the eastward advancing margin of the Cordilleran Overthrust Belt. The Two Medicine Formation is mostly sandstone, deposited by rivers and deltas.
The Kirtland Formation is a sedimentary geological formation.
Agujaceratops is a genus of horned dinosaur from the Late Cretaceous (Campanian) of west Texas. It is a chasmosaurine (long-frilled) ceratopsian. Two species are known, Agujaceratops mariscalensis, and A. mavericus.
The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Judith River Group. It dates to the Late Cretaceous, between 79 and 75.3 million years ago, corresponding to the "Judithian" land vertebrate age. It was laid down during the same time period as portions of the Two Medicine Formation of Montana and the Oldman Formation of Alberta. It is an historically important formation, explored by early American paleontologists such as Edward Drinker Cope, who named several dinosaurs from scrappy remains found here on his 1876 expedition. Modern work has found nearly complete skeletons of the hadrosaurid Brachylophosaurus.
The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.
The Fruitland Formation is a geologic formation found in the San Juan Basin in the states of New Mexico and Colorado, in the United States of America. It contains fossils dating it to the Campanian age of the late Cretaceous.
The Moreno Hill Formation is a geological formation in western New Mexico whose strata were deposited in the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation. The age of the formation is dated between approximately 90.9 to 88.6 million years ago based on detrital zircons.
The Javelina Formation is a geological formation in Texas. Dating has shown that the strata date to the Maastrichtian stage of the Late Cretaceous, approximately 70 to 66.5 million years old. The middle part of the formation has been dated to about 69 million years ago plus or minus 1 million years and the top situated near the Cretaceous–Paleogene boundary, dated to 66 Ma ago. Dinosaur remains are among the fossils that have been recovered from the formation.
The Almond Formation is a geological formation of Late Cretaceous age in Wyoming. It was deposited in marsh, deltaic, lagoonal, estuarine, and shallow marine environments along the western shore of the Western Interior Seaway. It consists primarily of fine- to medium-grained sandstone, siltstone, shale, and coal. Fossils from the Almond Formation include remains of dinosaurs and plants.
Angulomastacator is a genus of duck-billed dinosaur from the Campanian-age Aguja Formation of Big Bend National Park, Texas. It is known from a single specimen, TMM 43681–1, a partial left maxilla. This bone is curved down approximately 45° at its anterior end, with the tooth row bent to fit, unlike any other hadrosaur. The unusual characteristics of the maxilla, which have not been reported from elsewhere, supports the hypothesis that the dinosaurs of the Aguja Formation were endemic forms. It was discovered in the upper shale member of the Aguja Formation, among plant, bone, and clam fragments in a bed interpreted as the deposits of a small tributary channel. This bed is just below rocks of the overlying Javelina Formation. Volcanic rocks at about the same level have been dated to 76.9 ± 1.2 million years ago.
Texacephale is a possibly dubious genus of pachycephalosaurid dinosaur from the Campanian stage of the Late Cretaceous. Its fossils come from the Aguja Formation of Big Bend National Park, in Texas, and were described in 2010 by Longrich, Sankey and Tanke. The generic name means Texas + "head" in reference to its place of discovery, and the specific name langstoni honors Wann Langston. It may be a synonym of Stegoceras.
Chupacabrachelys is an extinct genus of bothremydid turtle which existed in western Texas, United States during the late Cretaceous period. Its fossils were discovered in Aguja Formation in the Big Bend region, and its type example is one of the most complete bothremydid specimens known. It was first named by Thomas M. Lehman and Steven L. Wick in 2010 and the type species is Chupacabrachelys complexus. Chupacabrachelys was named after the cryptid of the same name from Mexican folklore.
Bravoceratops is a genus of large chasmosaurine ceratopsid dinosaur that lived approximately 70 million years ago, and is known from the Late Cretaceous Javelina Formation in what is now Texas, United States.
Leptorhynchos is an extinct genus of caenagnathid theropod from the Late Cretaceous of what is now the US state of Texas, although it has been suggested to also exist in Alberta and South Dakota. The type species is L. gaddisi, and it is currently the only widely accepted valid species. The generic name of Leptorhynchos comes from the Greek "leptos" meaning "small" and "rhynchos" meaning "beak". The specific epithet is in honor of the Gaddis family, who owned the land on which the holotype was discovered.
Menefeeceratops is a genus of ceratopsid dinosaur from the Menefee Formation in New Mexico, United States. It is potentially the oldest known member of the ceratopsids, as well as the centrosaurine subfamily, related to animals including Yehuecauhceratops and Crittendenceratops. The type and only species is Menefeeceratops sealeyi, known from a partial, non-articulated skeleton.