Aguja Formation

Last updated
Aguja Formation
Stratigraphic range: Lower to Middle Campanian
~81.5–76.9  Ma
O
S
D
C
P
T
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K
Pg
N
Aguja Formation by Nick Longrich.jpg
Outcrops of the Aguja Formation, Big Bend National Park, Texas
Type Geological formation
Unit of Tornillo Group
Sub-unitsUpper Shale, Terlingua Creek Sandstone, Rattlesnake Mt. Sandstone, Lower Shale & Basal Sandstone Members
Underlies Javelina Formation
Overlies Pen Formation
Lithology
Primary Sandstone, conglomerate, claystone
Other Mudstone, shale, limestone
Location
Coordinates 29°18′N103°30′W / 29.3°N 103.5°W / 29.3; -103.5
Approximate paleocoordinates 35°48′N77°00′W / 35.8°N 77.0°W / 35.8; -77.0
RegionFlag of Texas.svg  Texas
Flag of Chihuahua.svg  Chihuahua
Flag of Coahuila.svg  Coahuila
CountryFlag of the United States.svg  USA
Flag of Mexico.svg  Mexico
Usa edcp relief location map.png
Lightgreen pog.svg
Aguja Formation (the United States)
Relief map of Texas.png
Lightgreen pog.svg
Aguja Formation (Texas)
Paleogeography of the Campanian North america 75mya.png
Paleogeography of the Campanian

The Aguja Formation is a geological formation in North America, exposed in Texas, United States and Chihuahua and Coahuila in Mexico, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation. [1] Fossil palms have also been unearthed here. [2]

Contents

Age

The ages of the Aguja Formation and its primary fossil-bearing unit, the Upper Shale, are not well understood. Due to the presence of the ammonite Baculites mclearni, which only occurs from 80.67 - 80.21 Ma, in the underlying Rattlesnake Mountain Sandstone and the Terlingua Creek Sandstone, it is likely that the Upper Shale was younger than 80.2 Ma. [3] A radiometric date of 76.9 Ma was recovered in the Upper Shale, making it likely the formation wasn't younger than 76.9 Ma. [3] The contact with the overlying Javelina Formation has been estimated at about 70 Ma ago [4] but also as recently as 68.5 million years ago. [5] This is unlikely, however, due to the presence of Bravoceratops, more primitive than an unnamed chasmosaurine from the De-na-zin Member of the Kirtland Formation, in the lowermost section of the formation. [6] The age of the Basal Sandstone is constrained by the presence of Scaphites hippocrepis III in the overlying Pen Formation which has been dated as old as 81.53 Ma. [3] [7]

Paleofauna

A duckbilled dinosaur (Hadrosauridae) caudal vertebra from the Aguja Formation Hadrosaur bone in the Aguja Formation by Nick Longrich.jpg
A duckbilled dinosaur (Hadrosauridae) caudal vertebra from the Aguja Formation

Reptiles

2 fragmentary caudal vertebrae of indeterminate reptiles are known from the Rattlesnake Mountain sandstone member. [8]

Pseudosuchians

Crurotarsans of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Crocodilia IndeterminateTen Bits Ranch, west Texas. [8] Rattlesnake Mountain sandstone member. [8] Teeth. [8] A crocodilian.
Deinosuchus [9] D. riograndensisTexas, [9] Chihuahua, and Coahuila [10] Osteoderm and mandible fragmentA giant alligatoroid.
Deinosuchus riograndensis.png
cf. D. sp.West Texas [11] Lower Shale [11] Cervical osteoderm (TMM 44068-2). [11] An alligatoroid.
Deinosuchus illustration Andrey Atuchin.jpg
Goniopholididae New genus & speciesWest Texas [11] Lower Shale [11] Partial skull & skeleton along with isolated osteoderms & teeth. [11] May pertain to Denazinosuchus or a related taxon.
Phobosuchus [9] P. riograndensis [9] Texas, [9] Chihuahua, and CoahuilaReclassified as a Deinosuchus species

Ornithischians

Ornithischians of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Agujaceratops A. mariscalensisTexas [12] Low Upper Shale [13] Formerly considered a species of Chasmosaurus [12]
Agujaceratops BW.jpg
A. mavericusWest Texas [14] A chasmosaurine.
Angulomastacator A. daviesiTexas [15] Upper Shale [15] A left maxilla. [15] A lambeosaurine.
Life reconstruction of Angulomastacator daviesi.png
Ankylosauridae IndeterminateTexas [16] Osteoderms, vertebrae & limb elements. [16] Remains of an ankylosaurid, possibly represents Euoplocephalus sp. [16]
Aquilarhinus [17] A. palimentusTexas [18] Lower Shale [18] Formerly referred to Kritosaurus and Gryposaurus. [19]
Aquilarhinus palimentus.png
Ceratopsidae genus & species indeterminateIndeterminateWest Texas [11] Lower Shale [11] Ilia, sacral vertebra & sacral ribs. [11] May represent Agujaceratops , but undiagnostic.
Chasmosaurus [20] C. mariscalensis [21] Texas [21] [Twelve] disarticulated skull (sic), postcrania, juvenile." [22] Considered by paleontologists Lucas, Sullivan, and Hunt to be distinct enough from the Chasmosaurus type species, C. belli to warrant being split off to a new genus, Agujaceratops. [12]
Edmontonia E. sp [23] Texas [16] Osteoderm (TVP 45866-2) & skull (AMNH 3076). [16] A nodosaurid.
Edmontonia dinosaur.png
cf. Euoplocephalus [21] E. sp [21] Texas [16] Upper Shale [16] Osteoderms, sacrum & vertebra (TL-05-14). [16] An ankylosaurid.
Euoplocephalus BW.jpg
Kritosaurus [20] Indeterminate [20] Chihuahua [24]
Kritosaurus BW.jpg
Malefica M. deckertiTexas [25] Upper Shale [25] A partial left maxilla. [25] Formerly referred to Kritosaurus. [25]
Nodosauridae genus & species indeterminateIndeterminateWest Texas [11] Lower Shale [11] Isolated & associated osteoderms. [11] Very similar to osteoderms of Invictarx .
Panoplosaurus P. mirusTexas [16] Osteoderm (TMM 45605-4). [16] A nodosaurid.
Panoplosaurus 055.JPG
Texacephale T. langstoniTexas [13] Low Upper Shale [13] Two frontoparietal domes. [13] A pachycephalosaur.
Texacephale langstoni skull dome by Nick Longrich.png
Yehuecauhceratops Y. mudei [26] Coahuila [27] A centrosaurine.
Yehuecauhceratops.jpg

Theropods

Indeterminate ornithomimid remains are known from the Upper Aguja Formation. [28] Indeterminate tyrannosaurid fossils are known from the Upper Aguja Formation of Texas and Mexico. [29]

Theropods of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Avialae incertae sedis West Texas [30] Lower Shale [30] 4 teeth (TMM 45947-349, 350, 351). [30] A bird.
Chirostenotes C. sp.Texas [31] Femur & manual ungual. [31] A caenagnathid
Volant Chirostenotes.jpg
Dromaeosauridae genus & species indeterminateIndeterminateWest Texas [11] Lower Shale [11] Pedal phalanges (TMM 45909-2, TMM 44066-4). [11] A dromaeosaur.
cf. Dromaeosaurus [32] Indeterminate [32] Texas [21]
Dromaeosaurus Restoration.png
Leptorhynchos L. gaddisiTexas [31] Dentaries, caudal vertebra & limb elements. [31] A caenagnathid
Leptorhynchos by Nick Longrich.jpg
Ornithomimidae New genus & speciesWest Texas [11] Lower Shale [11] Vertebrae, ischium & limb elements. [11] An unnamed species referred to informally as the 'Aguja ornithomimid'.
cf. Paronychodon West Texas [30] Lower Shale [30] 2 incomplete teeth (TMM 45947-362). [30] A maniraptoran.
Paronychodon.png
Richardoestesia R. cf. gilmoreiTexas [33] Low Upper Shale [33] Fragment of a small tooth. [33] A coelurosaur.
Ojo Alamo cf. Richardoestesia sp profile reconstruction.jpg
R. isosceles Big Bend National Park, Texas [33] Low Upper Shale [33] Teeth. [33] A coelurosaur.
Saurornitholestes S. cf. langstoniTexas [30] [33] Low Upper Shale & Lower Shale [30] [33] Teeth. [30] [33] A dromaeosaur.
Saurornitholestes digging Burrows wahweap.jpg
Theropoda incertae sedis Morphotype AWest Texas [30] Lower Shale [30] 8 teeth. [30] Unserrated, recurved teeth.
Morphotype BWest Texas [30] Lower Shale [30] 5 tooth crowns. [30] Bi-serrated teeth.
Morphotype CWest Texas [30] Lower Shale [30] 7 teeth. [30] Finely serrated, distal plication only.
Morphotype DWest Texas [30] Lower Shale [30] 5 teeth. [30] Short, coarsely serrated distal plication.
Morphotype EWest Texas [30] Lower Shale [30] A tooth. [30] Strongly recurved, serrated distal plication.
Morphotype FWest Texas [30] Lower Shale [30] 2 tooth fragments. [30] Medium size, finely serrated plications.
cf. Troodon [34] Indeterminate [34] Texas [21]
Troodon (cropped).jpg
Tyrannosauridae IndeterminateWest Texas [30] [35] Upper Shale [35] & Lower Shale [11] [30] Isolated teeth & a handful of non-dental specimens. [30] [35] A relatively small & gracile tyrannosaurid. [35]

Lepidosaurs

Lepidosaurs of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Anguidae Gen. et. sp. indet.Texas. [36] Isolated osteoderms & partial right frontal. [36] An anguid.
Apsgnathus A. triptodon Brewster County, Texas. [36] Jaw elements. [36] A scincomorph.
Catactegenys C. solaster Brewster County, Texas. [36] Jaw elements & teeth. [36] A night lizard.
Dryadissector D. shilleriWest Texas. [30] Lower Shale. [30] Numerous isolated teeth. [30] A varanoid.
Mosasauridae IndeterminateTen Bits Ranch. [8] Rattlesnake Mountain sandstone member. [8] 1 partial vetebra. [8] A mosasaur.
Odaxosaurus O. piger Brewster County, Texas. [36] Jaw elements. [36] An anguid.
cf. Parasaniwa cf. P. wyomingensisTexas. [36] Jaw fragments. [36] A platynotan.
Platynota Gen. et. sp. indet.Texas. [36] Dorsal vertebra (TMM 43057-332). [36] A platynotan.
?Scincidae Gen. et. sp. indet. Brewster County, Texas. [36] Jaw elements. [36] A possible skink.
cf. Scincomorpha Gen. et. sp. indet. Brewster County, Texas. [36] Jaw elements. [36] A scincomorph.
Serpentes Gen. et. sp. indet.Texas. [36] Partial left dentary & right maxilla. [36] A snake.
cf. Xenosauridae Texas. [36] Osteoderms & maxillae. [36] A knob-scaled lizard.

Turtles

Testudines of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Adocus A. sp. Big Bend National Park, Texas. [37] Upper Shale [37] Shell fragments. [37] An adocid.
Adocus shell illustration.png
Baenidae Big Bend National Park, Texas. [37] Upper Shale [37] Shell fragments. [37] 2 morphotypes present (A & B).
Basilemys B. sp. Big Bend National Park, Texas. [37] Upper Shale & Lower Shale [11] [37] Shell fragments, partial plastron & leg scutes. [11] [37] A nanhsiungchelyid.
Basilemys.jpg
Bothremydidae genus & species indeterminateIndeterminateWest Texas [11] Lower Shale [11] Isolated peripheral bones (TMM 44068-1, 42452-8); costal bones (TMM 44064-6). [11] A bothremydid, compatible with Chupacabrachelys .
Chupacabrachelys C. complexus Big Bend, Texas. [38] Base of the Upper Shale [38] A complete skull, and a nearly complete skeleton. [38] A bothremydid.
Chupacabrachelys cropped.png
cf. Denazinemys cf. D. sp.West Texas [11] Lower Shale [11] Shell elements. [11] A baenid.
cf. Helopanoplia Big Bend National Park, Texas. [37] Upper Shale [37] Shell fragments. [37] A softshell turtle.
cf. Hoplochelys Big Bend National Park, Texas. [37] Upper Shale [37] Shell fragments. [37] A kinosternoid.
Terlinguachelys T. fischbecki Big Bend National Park, Texas [39] Rattlesnake Mountain Sandstone [39] A large, incomplete specimen. [39] A protostegid.
Testudines indeterminate [8] Morphotype 1 [8] Ten Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] A shell fragment.
Morphotype 2 [8] Ten Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] A shell fragment.
Morphotype 3 [8] Ten Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] A shell fragment.
Morphotype 4 [8] Ten Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] A shell fragment.
Trionychidae Gen. et. sp. indet. Big Bend National Park, Texas. [37] Upper Shale & Lower Shale [11] [37] Costal bone (TMM 44068-4) & shell fragments. [37] A softshell turtle.

Bony Fish

Approximately 75 whole and broken fragments of coprolites are known from the Rattlesnake Mountain sandstone member, presumably from bony fish. [8]

Bony fish of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Albula A. sp.Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Over 390 complete and fragmentary teeth. [40] A bonefish.
Bonefish Albula vulpes.jpg
?Enchodus ?E. sp.Ten Bits ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 2 abraded teeth. [40] An aulopiform.
Enchodus petrosus.png
Eotexachara E. malateresWest Texas [41] Dentaries. [41] A characiform.
Lepidotes ?L. sp.Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Approximately 109 complete and fragmentary teeth. [40] A lepidotid.
Melvius M. sp.West Texas [11] Lower Shale [11] Vertebrae. [11] An amiid.
Osteichthyes Indeterminate species ATen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 11 complete teeth.
Indeterminate species BTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 8 complete and fragmentary teeth.
Indeterminate species CTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 1 complete tooth.
Indeterminate species DTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 1 complete and 1 partial tooth.
IndeterminateTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] Approximately 475 complete and fragmentary teleost centra. [40]
Paralbula P. caseiTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Approximately 900 complete and fragmentary teeth and tooth caps. [40]
Primuluchara P. laramidensisWest Texas [41] Dentaries. [41] A characiform.
Stephanodus ?S. sp.Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 21 whole and fragmentary specimens. [40] A pycnodont.

Cartilaginous fish

Cartilaginous fish of the Aguja Formation
GenusSpeciesLocationMemberMaterialNotesImages
Brachyrhyzodus B. wichitaensisTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 11 complete teeth. [40]
Cantioscyllium C. aff. meyeriTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 1 anterior tooth and 5 lateral teeth. [8] A nurse shark.
Chiloscyllium C. aff. greeniTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] Approximately 90 abraded and fragmentary teeth. [40] A bamboo shark.
Chondrichthyes [8] Ten Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 23 placoid scales and 69 dermal scales.4 morphotypes of placoid scales (A to D) present.
Columbusia C. sp.Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 20 complete & fragmentary teeth. [40] A wobbegong.
Cretalamna C. appendiculataTen Bits Ranch, west Texas. [8] Rattlesnake Mountain sandstone member. [8] 3 fragmentary teeth. [8] Reassigned to C. cf. C. sarcoportheta. [40]
C. cf. C. sarcoporthetaTen Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 3 incomplete teeth & fragments of additional teeth. [40] Originally reported as C. appendiculata.
Cretalamna reconstruction.png
Cretorectolobus [8] C. olsoniTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] "20 complete and fragmentary teeth".A carpet shark.
Hybodontidae genus & species indeterminateIndeterminateWest Texas [11] Lower Shale [11] Fragment of a dorsal fin spine (TMM 42536-10). [11] A hybodont.
Hybodus [8] H. sp.Ten Bits Ranch.Rattlesnake Mountain sandstone member. [8] 2 specimens, one complete and one partial tooth.A hybodont.
Hybodus hauffianus.png
Igdabatis I. indicus?Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] A single incomplete tooth. [8] [40] A myliobatid.
Ischyrhiza I. cf. avonicolaTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 5 complete and fragmentary rostral teeth. [40] A sawskate.
I. miraTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 11 fragmentary rostral teeth and 230 whole and fragmentary oral teeth. [40] A sawskate.
Ischyrhiza mira.jpg
Lonchidion L. selachosTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 4 complete and 5 fragmentary teeth. [8] [40] A hybodont.
Meristodon M. sp.Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 2 teeth. [40] A hybodont.
Myliobatiformes Incertae sedis Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 3 complete specimens. [40]
Protoplatyrhina P. renaeTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] 70 complete and fragmentary teeth. [40] A hypsobatid.
Ptychotrygon P. agujaensisTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Over 690 complete and fragmentary teeth. [40] A sawskate.
P. triangularisTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Over 170 complete and fragmentary teeth. [40] A sawskate.
P. aff. cuspidataTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] 7 whole and fragmentary teeth. [40] A sawskate.
P. sp.Ten Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] One complete tooth (TMM 46018-71). [40] A sawskate.
Ptychotrygon sp.jpg
Rhinobatos R. casieriTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] "14 complete and fragmentary specimens".A guitarfish.
R. sp.Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 18 complete & fragmentary specimens. [40] A guitarfish.
Rhombodus R. levisTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Over 22 complete and fragmentary teeth. [40] A rajiforme.
Scapanorhynchus S. texanusTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] Over 800 complete and fragmentary teeth. [40] A mitsukurinid.
Scapanorhynchus lewisii.jpg
Sclerorhynchidae Morphotype 1Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [8] One fragmentary rostral spine (TMM 46018-59). [40] A sawskate.
Morphotype 2Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [8] One fragmentary rostral spine (TMM 46018-60). [40] A sawskate.
Morphotype 3Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [8] 3 fragmentary rostral teeth. [40] A sawskate.
Morphotype 4Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [8] 27 fragmentary rostral spines. [40] A sawskate.
Serratolamna S. cf. S. caraibaeaTen Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] About 34 teeth. [40] A mackerel shark.
Squalicorax S. kaupiTen Bits Ranch, west Texas. [8] [40] Rattlesnake Mountain sandstone member. [8] [40] "26 complete and fragmentary anterior and lateral teeth".An anacoracid.
MenuhaSqualicoraxKaupi.jpg
S. aff. S. lindstromiTen Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 4 specimens. [40] An anacoracid.
S. pristodontusTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 2 specimens.An anacoracid.
Squalicorax pristodontus Agassiz, 1843 1.jpg
S. aff. S. yangaensisTen Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 4 complete & several fragmentary teeth. [40] An anacoracid.
Squatina S. hasseiTen Bits Ranch, west Texas. [8] Rattlesnake Mountain sandstone member. [8] A single complete specimen.An angelshark.
S. sp.Ten Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] A complete tooth & 2 fragmentary teeth. [40] An angelshark.
Texatrygon T. cf. T. copeiTen Bits Ranch, west Texas. [40] Rattlesnake Mountain sandstone member. [40] 4 complete & fragmentary teeth. [40] A sawskate formerly reported as T. hooveri.
T. hooveriTen Bits Ranch, west Texas.Rattlesnake Mountain sandstone member. [8] 4 complete and fragmentary specimens. [8] Reassigned to T. cf. T. copei.

Invertebrates

Ammonites

Ammonites of the Aguja Formation
GenusSpeciesLocationMemberAbundanceNotesImages
Baculites B. mclearni
  • Rattlesnake Mountain Sandstone
  • Terlingua Creek Sandstone
Hoplitoplacenticeras H. plasticumRattlesnake Mountain Sandstone
Pachydiscus P. paulsoniRattlesnake Mountain Sandstone
Pachydiscus neubergicus.jpg
Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

Related Research Articles

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<span class="mw-page-title-main">Javelina Formation</span> Geological formation in Texas, USA

The Javelina Formation is a geological formation in Texas. Dating has shown that the strata date to the Maastrichtian stage of the Late Cretaceous, approximately 70 to 66.5 million years old. The middle part of the formation has been dated to about 69 million years ago plus or minus 1 million years and the top situated near the Cretaceous–Paleogene boundary, dated to 66 Ma ago. Dinosaur remains are among the fossils that have been recovered from the formation.

The El Picacho Formation is a geological formation in Texas, United States, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation. The paleosols found here are rich in clay, calcite, and rhizoliths which show that during the Cretaceous period, this fossil formation, just like the neighboring Javelina Formation and Aguja Formation, was a fluvial flood plain.

<i>Angulomastacator</i> Extinct genus of dinosaur

Angulomastacator is a genus of duck-billed dinosaur from the Campanian-age Aguja Formation of Big Bend National Park, Texas. It is known from a single specimen, TMM 43681–1, a partial left maxilla. This bone is curved down approximately 45° at its anterior end, with the tooth row bent to fit, unlike any other hadrosaur. The unusual characteristics of the maxilla, which have not been reported from elsewhere, supports the hypothesis that the dinosaurs of the Aguja Formation were endemic forms. It was discovered in the upper shale member of the Aguja Formation, among plant, bone, and clam fragments in a bed interpreted as the deposits of a small tributary channel. This bed is just below rocks of the overlying Javelina Formation. Volcanic rocks at about the same level have been dated to 76.9 ± 1.2 million years ago.

<i>Chupacabrachelys</i> Extinct genus of turtles

Chupacabrachelys is an extinct genus of bothremydid turtle which existed in western Texas, United States during the late Cretaceous period. Its fossils were discovered in Aguja Formation in the Big Bend region, and its type example is one of the most complete bothremydid specimens known. It was first named by Thomas M. Lehman and Steven L. Wick in 2010 and the type species is Chupacabrachelys complexus. Chupacabrachelys was named after the cryptid of the same name from Mexican folklore.

<i>Yehuecauhceratops</i> Extinct genus of dinosaurs

Yehuecauhceratops is a genus of horned centrosaurine ceratopsid dinosaur from the Late Cretaceous of Coahuila, Mexico. It contains a single species, Y. mudei, described from two partial specimens by Rivera-Sylva et al. in 2016 and formally named by Rivera-Sylva et al. in 2017. It was a small centrosaurine with a body length of 3 metres (9.8 ft), making it smaller than Agujaceratops and Coahuilaceratops, the other two ceratopsids in its environment; the three may have been ecologically segregated. A ridge bearing a single roughened projection near the bottom of the squamosal bone, which probably supported a small horn, allows Yehuecauhceratops to be distinguished from other centrosaurines. Its affinities to nasutoceratopsin centrosaurines, such as Avaceratops and Nasutoceratops, are supported by various morphological similarities to the former.

<i>Leptorhynchos gaddisi</i> Extinct species of dinosaur

Leptorhynchos is an extinct genus of caenagnathid theropod from the Late Cretaceous of what is now the US state of Texas, although it has been suggested to also exist in Alberta and South Dakota. The type species is L. gaddisi, and it is currently the only widely accepted valid species. The generic name of Leptorhynchos comes from the Greek "leptos" meaning "small" and "rhynchos" meaning "beak". The specific epithet is in honor of the Gaddis family, who owned the land on which the holotype was discovered.

<i>Menefeeceratops</i> Extinct genus of dinosaurs

Menefeeceratops is a genus of ceratopsid dinosaur from the Menefee Formation in New Mexico, United States. It is potentially the oldest known member of the ceratopsids, as well as the centrosaurine subfamily, related to animals including Yehuecauhceratops and Crittendenceratops. The type and only species is Menefeeceratops sealeyi, known from a partial, non-articulated skeleton.

References

  1. Weishampel et al., 2004, "Dinosaur distribution (Late Cretaceous, North America).", pp.574-588
  2. Manchester, Steven R.; Lehman, Thomas M.; Wheeler, Elisabeth A. (July 2010). "Fossil Palms (Arecaceae, Coryphoideae) Associated with Juvenile Herbivorous Dinosaurs in the Upper Cretaceous Aguja Formation, Big Bend National Park, Texas". International Journal of Plant Sciences. 171 (6): 679–689. doi:10.1086/653688. ISSN   1058-5893. S2CID   84762968.
  3. 1 2 3 Fowler, Denver Warwick (2017-11-22). "Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America". PLOS ONE. 12 (11): e0188426. Bibcode:2017PLoSO..1288426F. doi: 10.1371/journal.pone.0188426 . ISSN   1932-6203. PMC   5699823 . PMID   29166406.
  4. Woodward, H. N. (2005). Bone histology of the sauropod dinosaur Alamosaurus sanjuanensis from the Javelina Formation, Big Bend National Park, Texas.
  5. Sankey, J. (2010). Faunal composition and significance of high–diversity, mixed bonebeds containing Agujaceratops mariscalensis and other dinosaurs, Aguja Formation (Upper Cretaceous), Big Bend, Texas. In New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium (pp. 520-537).
  6. Fowler, Denver W.; Fowler, Elizabeth A. Freedman (2020-06-05). "Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico". PeerJ. 8: e9251. doi: 10.7717/peerj.9251 . ISSN   2167-8359. PMC   7278894 . PMID   32547873.
  7. Prieto-Márquez, Albert; Wagner, Jonathan R.; Lehman, Thomas (2020-03-18). "An unusual 'shovel-billed' dinosaur with trophic specializations from the early Campanian of Trans-Pecos Texas, and the ancestral hadrosaurian crest" (PDF). Journal of Systematic Palaeontology. 18 (6): 461–498. doi:10.1080/14772019.2019.1625078. ISSN   1477-2019. S2CID   202018197.
  8. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 Schubert, Joseph (May 2013). Elasmobranch and osteichthyan fauna of the Rattlesnake Mountain Sandstone, Aguja Formation (Upper Cretaceous; Campanian), West Texas (Thesis thesis).
  9. 1 2 3 4 5 "Aguja Formation (Upper Shale Member), Big Bend Region, Texas," in Sullivan and Lucas (2006). Page 16.
  10. "The first mandible fragment of Deinosuchus (Eusuchia: Alligatoroidea) discovered in Coahuila, Mexico". ResearchGate. Retrieved 2016-03-27.
  11. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 Lehman, Thomas M.; Wick, Steven L.; Brink, Alyson A.; Shiller, Thomas A. (2019). "Stratigraphy and vertebrate fauna of the lower shale member of the Aguja Formation (lower Campanian) in West Texas". Cretaceous Research. 99: 291–314. Bibcode:2019CrRes..99..291L. doi:10.1016/j.cretres.2019.02.028. ISSN   0195-6671. S2CID   135044927.
  12. 1 2 3 Lucas, Spencer G.; Sullivan, Robert M.; Hunt, Adrian P. (January 2006). "Re-evaluation of Pentaceratops and Chasmosaurus (Ornithischia: Ceratopsidae) in the Upper Cretaceous of the Western Interior". New Mexico Museum of Natural History and Science Bulletin. 35: 367–370.
  13. 1 2 3 4 Longrich, N. R.; Sankey, J.; Tanke, D. (2010). "Texacephale langstoni, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA". Cretaceous Research. 31 (2): 274. Bibcode:2010CrRes..31..274L. doi:10.1016/j.cretres.2009.12.002.
  14. Lehman, Thomas M.; Wick, Steven L.; Barnes, Kenneth R. (2017-08-03). "New specimens of horned dinosaurs from the Aguja Formation of West Texas, and a revision of Agujaceratops". Journal of Systematic Palaeontology. 15 (8): 641–674. doi:10.1080/14772019.2016.1210683. ISSN   1477-2019. S2CID   88907183.
  15. 1 2 3 Wagner, Jonathan R.; Lehman, Thomas M. (2009-06-12). "An enigmatic new lambeosaurine hadrosaur (Reptilia: Dinosauria) from the Upper Shale member of the Campanian Aguja Formation of Trans-Pecos Texas". Journal of Vertebrate Paleontology. 29 (2): 605–611. Bibcode:2009JVPal..29..605W. doi:10.1671/039.029.0208. ISSN   0272-4634. S2CID   128555861.
  16. 1 2 3 4 5 6 7 8 9 10 West, Bryanna (2020). Campanian-Maastrictian Ankylosaurs of West Texas (PDF) (Thesis). Texas Tech University.
  17. Listed as ?Gryposaurus sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
  18. 1 2 Prieto-Márquez, Albert; Wagner, Jonathan R.; Lehman, Thomas (2020-03-18). "An unusual 'shovel-billed' dinosaur with trophic specializations from the early Campanian of Trans-Pecos Texas, and the ancestral hadrosaurian crest". Journal of Systematic Palaeontology. 18 (6): 461–498. doi:10.1080/14772019.2019.1625078. ISSN   1477-2019. S2CID   202018197.
  19. Prieto-Márquez, Albert (2014-02-17). "Skeletal morphology of Kritosaurus navajovius (Dinosauria: Hadrosauridae) from the Late Cretaceous of the North American south-west, with an evaluation of the phylogenetic systematics and biogeography of Kritosaurini". Journal of Systematic Palaeontology. 12 (2): 133–175. doi:10.1080/14772019.2013.770417. ISSN   1477-2019. S2CID   84942579.
  20. 1 2 3 "Dinosaur distribution (Texas and Chihuahua)." Weishampel, et al. (2004). Page 582 and 588.
  21. 1 2 3 4 5 6 "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
  22. "Table 23.1," in Weishampel, et al. (2004). Page 496.
  23. Listed as Edmontonia cf. rugosidens in "Dinosaur distribution (Late Cretaceous; North America; Texas).", Weishampel et al., 2004, p.582
  24. "Dinosaur distribution (Late Cretaceous; North America; Chihuahua)." Weishampel, et al. (2004). Page 588.
  25. 1 2 3 4 Prieto-Márquez, Albert; Wagner, Jonathan R. (2022-11-10). "A new 'duck-billed' dinosaur (Ornithischia: Hadrosauridae) from the upper Campanian of Texas points to a greater diversity of early hadrosaurid offshoots". Cretaceous Research. 143: 105416. doi:10.1016/j.cretres.2022.105416. ISSN   0195-6671. S2CID   253470207.
  26. Rivera-Sylva, H.E.; Hendrick, B.P.; Dodson, P. (2016). "A Centrosaurine (Dinosauria: Ceratopsia) from the Aguja Formation (Late Campanian) of Northern Coahuila, Mexico". PLOS ONE. 11 (4): e0150529. Bibcode:2016PLoSO..1150529R. doi: 10.1371/journal.pone.0150529 . PMC   4830452 . PMID   27073969.
  27. Rivera-Sylva, H.E.; Frey, E.; Stinnesbeck, W.; Guzman-Gutirrez, J.R.; Gonzalez-Gonzalez (2017). "Mexican ceratopsids: Considerations on their diversity and evolution". Journal of South American Earth Sciences. doi:10.1016/j.jsames.2017.01.008.
  28. Sullivan, R.M., and Lucas, S.G. 2006. "The Kirtlandian land-vertebrate "age" – faunal composition, temporal position and biostratigraphic correlation in the nonmarine Upper Cretaceous of western North America [ permanent dead link ]." New Mexico Museum of Natural History and Science, Bulletin 35:7-29.
  29. Mortimer, M (2004). "Tyrannosauroidea". The Theropod Database. Archived from the original on 2013-09-29. Retrieved 2007-08-21.
  30. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 Wick, Steven L.; Lehman, Thomas M.; Brink, Alyson A. (2015). "A theropod tooth assemblage from the lower Aguja Formation (early Campanian) of West Texas, and the roles of small theropod and varanoid lizard mesopredators in a tropical predator guild". Palaeogeography, Palaeoclimatology, Palaeoecology. 418: 229–244. Bibcode:2015PPP...418..229W. doi:10.1016/j.palaeo.2014.11.018. ISSN   0031-0182.
  31. 1 2 3 4 Longrich, Nicholas R.; Barnes, Ken; Clark, Scott; Millar, Larry (April 2013). "Caenagnathidae from the Upper Campanian Aguja Formation of West Texas, and a Revision of the Caenagnathinae". Bulletin of the Peabody Museum of Natural History. 54 (1): 23–49. doi:10.3374/014.054.0102. ISSN   0079-032X. S2CID   128444961.
  32. 1 2 Listed as cf. Dromaeosaurus sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
  33. 1 2 3 4 5 6 7 8 9 Sankey, Julia T. (2001). "Late Campanian Southern Dinosaurs, Aguja Formation, Big Bend, Texas". Journal of Paleontology. 75 (1): 208–215. doi:10.1666/0022-3360(2001)075<0208:LCSDAF>2.0.CO;2. ISSN   0022-3360. JSTOR   1306931.
  34. 1 2 Listed as cf. Troodon sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
  35. 1 2 3 4 Lehman, Thomas M.; Wick, Steven L. (September 2012). "Tyrannosauroid dinosaurs from the Aguja Formation (Upper Cretaceous) of Big Bend National Park, Texas". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 471–485. doi:10.1017/S1755691013000261. ISSN   1755-6910. S2CID   129232391.
  36. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 Nydam, Randall L.; Rowe, Timothy B.; Cifelli, Richard L. (2013). "Lizards and Snakes of the Terlingua Local Fauna (late Campanian), Aguja Formation, Texas, with Comments on the Distribution of Paracontemporaneous Squamates Throughout the Western Interior of North America". Journal of Vertebrate Paleontology. 33 (5): 1081–1099. Bibcode:2013JVPal..33.1081N. doi:10.1080/02724634.2013.760467. ISSN   0272-4634. JSTOR   42568627. S2CID   86519841.
  37. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Sankey, Julia (January 2006). "Turtles of the upper Aguja Formation (late Campanian), Big Bend National Park, Texas". New Mexico Museum of Natural History and Science Bulletin. 35: 235–243.
  38. 1 2 3 Lehman, Thomas M.; Wick, Steven L. (2010). "Chupacabrachelys complexus, N. Gen. N. Sp. (testudines: Botheremydidae), from the Aguja Formation (campanian) of West Texas". Journal of Vertebrate Paleontology. 30 (6): 1709–1725. Bibcode:2010JVPal..30.1709L. doi:10.1080/02724634.2010.520782. ISSN   0272-4634. JSTOR   25790797. S2CID   129303574.
  39. 1 2 3 Lehman, Thomas M.; Tomlinson, Susan L. (November 2004). "Terlinguachelys fischbecki, a new genus and species of sea turtle (Chelonioidea: Protostegidae) from the Upper Cretaceous of Texas". Journal of Paleontology. 78 (6): 1163–1178. doi:10.1666/0022-3360(2004)078<1163:TFANGA>2.0.CO;2. ISSN   0022-3360. S2CID   54755373.
  40. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87 88 89 90 91 92 93 94 95 96 Schubert, Joseph A.; Wick, Steven L.; Lehman, Thomas M. (January 2017). "An Upper Cretaceous (middle Campanian) marine chondrichthyan and osteichthyan fauna from the Rattlesnake Mountain sandstone member of the Aguja Formation in West Texas". Cretaceous Research. 69: 6–33. Bibcode:2017CrRes..69....6S. doi:10.1016/j.cretres.2016.08.008. S2CID   133098369.
  41. 1 2 3 4 Wick, Steven L. (2021-12-01). "New early Campanian characiform fishes (Otophysi: Characiformes) from West Texas support a South American origin for known Late Cretaceous characiforms from North America". Cretaceous Research. 128: 104993. Bibcode:2021CrRes.12804993W. doi:10.1016/j.cretres.2021.104993. ISSN   0195-6671.

Bibliography