Cyclurus Temporal range: Campanian occurrence Possible | |
---|---|
Fossil of C. kehreri | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Clade: | Halecomorphi |
Order: | Amiiformes |
Family: | Amiidae |
Subfamily: | Amiinae |
Genus: | † Cyclurus Agassiz, 1844 |
Type species | |
†Cyclurus valenciennesi Agassiz, 1844 | |
Species | |
See text | |
Synonyms | |
Cyclurus (Ancient Greek for "rounded tail") is an extinct genus of freshwater amiid ray-finned fish known from the Late Cretaceous to the Early Oligocene across much of the Northern Hemisphere. [1] It is thought to be the closest relative of the extant bowfins in the genus Amia , although species of Cyclurus were significantly smaller in size compared to Amia. [2]
Remains of Cyclurus are first known from western North America during the Maastrichtian stage of the Cretaceous with the species C. fragosus, with the genus having presumably diverged from Amia shortly before. Potential earlier remains are known from the Campanian. [1] [2] Alongside Amia, Cyclurus survived the Cretaceous-Paleogene extinction event in a refugium in western North America. [2] Shortly afterwards, it saw a dramatic range expansion over the Paleogene, colonizing Europe and Asia by the Late Paleocene and becoming particularly successful and speciose in Europe. By the Late Eocene, it was one of the last two surviving halecomorph genera, alongside Amia. However, alongside most Amia species aside from those in eastern North America, it became extinct during the Oligocene. [2] [3]
The following species are known: [3]
The species †C. "lacus" (Russell, 1928) (=†Stylomyleodon lacusRussell, 1928) from the middle Paleocene of Alberta, †C. "chinzhalensis" (Sytchevskaya, 1986) (=†Amia chinzhalensisSytchevskaya, 1986) from the mid-late Eocene of Kazakhstan, and †C. "russelli" (Janot, 1966) (=†"Amia" russelliJanot, 1966) from the late Paleocene of France are considered nomina dubia due to insufficiently diagnostic remains. The Geiseltal specimens of C. kehreri differ morphologically from the Messel ones, and may represent a distinct species. [3] Indeterminate Cyclurus remains are known from the middle Eocene of Jilin, China, the late Paleocene Ravenscrag Formation of Saskatchewan & the Maastrichtian Horseshoe Canyon Formation of Alberta, Canada, and possibly the Campanian-aged Aguja Formation of Texas, USA. [8] [9] [10] [11]
C. kehreri is the most common fossil fish known from the Messel pit, and in fact the most abundant fossil amiid in the world, with thousands of specimens recovered. Fossils of C. kehreri are known representing different growth stages. Almost no specimens of C. kehreri are known with fish bones in their stomach (a specimen with percoid bones in its stomach is thought to have likely scavenged on them), suggesting that unlike other amiids, Cyclurus likely fed on small invertebrates instead of other fish. However, more recently, two C. kehreri specimens have been discovered with specimens of the bat Palaeochiropteryx caught in their mouths, suggesting that they either opportunistically attacked the dying bats or attempted scavenging on them, although these attempts were unsuccessful. [12]
Contrasting with C. kehreri's abundance, C. gurleyi is one of the rarest fishes from the Green River Formation, with only 8 specimens known from the hundreds of thousands of fossil fishes excavated. [3]
The Amiiformes order of fish has only two extant species, the bowfins: Amia calva and Amia ocellicauda, the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. The order first appeared in the Triassic, and the extinct members include both marine and freshwater species, many of which are morphologically disparate from bowfins, such as the caturids.
The Amiidae are a family of basal ray-finned fishes. The bowfin and the eyespot bowfin are the only two species to survive today, although additional species in all four subfamilies of Amiidae are known from Jurassic, Cretaceous, and Eocene fossils.
Podocnemididae is a family of pleurodire (side-necked) turtles, once widely distributed. Most of its 41 genera and 57 species are now extinct. Seven of its eight surviving species are native to South America: the genus Peltocephalus, with two species, only one of which is extant ; and the genus Podocnemis, with six living species of South American side-necked river turtles and four extinct. There is also one genus native to Madagascar: Erymnochelys, the Madagascan big-headed turtle, whose single species E. madagascariensis.
The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.
Madtsoiidae is an extinct family of mostly Gondwanan snakes with a fossil record extending from early Cenomanian to late Pleistocene strata located in South America, Africa, India, Australia and Southern Europe. Madtsoiidae include very primitive snakes, which like extant boas and pythons would likely dispatch their prey by constriction. Genera include some of the longest snakes known such as Vasuki, measuring at least 11–15 metres (36–49 ft) long, and the Australian Wonambi and Yurlunggur. As a grouping of basal forms the composition and even the validity of Madtsoiidae is in a state of flux as new pertinent finds are described, with more recent evidence suggesting that it is paraphyletic as previously defined.
Coriops is an extinct genus of freshwater osteoglossomorph fish, possibly a hiodontiform, with a single species known from the Late Cretaceous of western North America.
Bothremydidae is an extinct family of side-necked turtles (Pleurodira) known from the Cretaceous and Cenozoic. They are closely related to Podocnemididae, and are amongst the most widely distributed pleurodire groups, with their fossils having been found in Africa, India, the Middle East, Europe, North America and South America. Bothremydids were aquatic turtles with a high morphological diversity, indicative of generalist, molluscivorous, piscivorous and possibly herbivorous grazing diets, with some probably capable of suction feeding. Unlike modern pleurodires, which are exclusively freshwater, bothremydids inhabited freshwater, marine and coastal environments. Their marine habits allowed bothremydids to disperse across oceanic barriers into Europe and North America during the early Late Cretaceous (Cenomanian). The youngest records of the group are indeterminate remains from Saudi Arabia and Oman, dating to the Miocene.
Madtsoia is an extinct genus of madtsoiid snakes. It is known from the Eocene of Argentina, the Paleocene of Brazil, the Late Cretaceous (Maastrichtian) of India, and the Late Cretaceous (Maastrichtian) of Madagascar. The type species was the largest with an estimated length of 9–10 m (30–33 ft), and the other three species were smaller. A 5.1 m (17 ft) long M. madagascariensis would have weighed 50 kg (110 lb), but an isolated specimen suggests that this species reached 8 m (26 ft) in maximum length.
Bathysoma is an extinct genus of marine lampriform ray-finned fish from the early-mid Paleocene. It contains a single species, B. lutkeni from Sweden. Its fossils are common in exposures of the Danian København Limestone Formation at Limhamns kalkbrott, one of the largest quarries in northern Europe. A single specimen is also known from an erratic boulder from the Selandian Lellinge Greensand Formation of southern Sweden.
Calamopleurus is a prehistoric genus of marine holostean ray-finned fish from the Early Cretaceous of South America and northern Africa. It was a relative of the modern bowfin, with both belonging to the family Amiidae.
Ampheristus is an extinct genus of prehistoric marine ray-finned fish. It was a basal or stem member of the family Ophidiidae, which contains modern cusk-eels. Fossils are known from worldwide from the Late Cretaceous to the late Paleogene, making it a rather successful survivor of the Cretaceous-Paleogene extinction event.
Dapalis is an extinct genus of prehistoric glassfish known from the Late Cretaceous to the Early Miocene. It is known from both freshwater and marine habitats of India, Australia, New Zealand, and much of mainland Europe.
Cylindracanthus is an extinct, enigmatic genus of marine ray-finned fish with fossils known throughout North America, Europe, Asia and Africa from the Late Cretaceous to the late Eocene, with potential Oligocene records and a possible Miocene record also known. It is exclusively known from its distinctive partial remains, which are long cylindrical bony spines that are usually considered rostrum fragments, as well as some associated teeth. These spines are abundant & widespread throughout this timespan, and are useful indicators of a nearshore marine environment, but the taxonomic identity of the fish is still highly uncertain and debated.
The Ravenscrag Formation is a stratigraphic unit of early Paleocene age in the Western Canada Sedimentary Basin. It was named for the settlement of Ravenscrag, Saskatchewan, and was first described from outcrops at Ravenscrag Butte near the Frenchman River by N.B. Davis in 1918.
Melvius is a genus of vidalamiin amiid fish from the Late Cretaceous. The type species, Melvius thomasi, was described by Bryant in 1987 from Hell Creek Formation. A second species Melvius chauliodous, was named and described by Hall and Wolburg in 1989 from Kirtland Formation, and it is now considered to be one of the index taxa of the Kirtlandian land-vertebrate age. Both species of Melvius were very large at its size. A vertebral remain of M. thomasi would belongs to fish with standard length of 161 cm (5.28 ft), and there are some specimens exceeds height of that vertebra. Total length of this species would be at least 193–205 cm (6.33–6.73 ft). However, M. thomasi would be a “dwarf” compared to M. chauliodous, a specimen of M. chauliodous with abdominal centra which is 6.57 cm (2.59 in) wide would indicate standard length over 2 m (6.6 ft), and there is even larger abdominal centra which is 7.3 cm (2.9 in) wide.
Tingmiatornis is a genus of flighted and possibly diving ornithurine dinosaur from the High Arctic of Canada. The genus contains a single species, T. arctica, described in 2016, which lived during the Turonian epoch of the Cretaceous.
Amia, commonly called bowfin, is a genus of ray-finned fish related to gars in the infraclass Holostei. They are regarded as taxonomic relicts, being the sole surviving species of the order Amiiformes and clade Halecomorphi, which dates from the Triassic to the Eocene, persisting to the present. There are two living species in Amia, Amia calva and Amia ocellicauda, and a number of extinct species which have been described from the fossil record.
Meiolaniformes is an extinct clade of stem-group turtles, defined as all taxa more closely related to Meiolania than to Cryptodira and Pleurodira. It is known from the Early Cretaceous to the Holocene of Australia, Oceania and South America. Some Eurasian taxa have been suggested to be part of the group, but this is disputed.
Amia? hesperia is an extinct species of bony fish in the bowfin family, Amiidae. The species is known from fossils found in the early Eocene deposits of northern Washington state in the United States and southeastern British Columbia. The species is one of eight fish species identified in the Eocene Okanagan Highlands paleofauna.
Maliamia is an extinct genus of amiid ray-finned fish from the Early Eocene, known from fragmentary remains found in the Tamaguélelt Formation of Mali. It was described in 1989, based on fossils recovered by three separate expeditions in 1975, 1979–80, and 1981. The type species is Maliamia gigas, named in reference to its large size.
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