Wuliuan | ||||||||||||||||||||||||||||||||||||||
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Chronology | ||||||||||||||||||||||||||||||||||||||
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Etymology | ||||||||||||||||||||||||||||||||||||||
Name formality | Formal | |||||||||||||||||||||||||||||||||||||
Name ratified | 2018 [2] | |||||||||||||||||||||||||||||||||||||
Former name(s) | Cambrian Stage 5 | |||||||||||||||||||||||||||||||||||||
Usage information | ||||||||||||||||||||||||||||||||||||||
Celestial body | Earth | |||||||||||||||||||||||||||||||||||||
Regional usage | Global (ICS) | |||||||||||||||||||||||||||||||||||||
Time scale(s) used | ICS Time Scale | |||||||||||||||||||||||||||||||||||||
Definition | ||||||||||||||||||||||||||||||||||||||
Chronological unit | Age | |||||||||||||||||||||||||||||||||||||
Stratigraphic unit | Stage | |||||||||||||||||||||||||||||||||||||
First proposed by | Zhao et al., 2018 [3] | |||||||||||||||||||||||||||||||||||||
Time span formality | Formal | |||||||||||||||||||||||||||||||||||||
Lower boundary definition | FAD of Oryctocephalus indicus . | |||||||||||||||||||||||||||||||||||||
Lower boundary GSSP | Wuliu-Zengjiayan, Guizhou, China 26°04′51″N108°24′50″E / 26.0807°N 108.4138°E | |||||||||||||||||||||||||||||||||||||
Lower GSSP ratified | 2018 [2] | |||||||||||||||||||||||||||||||||||||
Upper boundary definition | FAD of the Trilobite Ptychagnostus atavus | |||||||||||||||||||||||||||||||||||||
Upper boundary GSSP | Drumian section, Wheeler Shale, Utah, U.S.A. 39°30′42″N112°59′29″W / 39.5117°N 112.9915°W | |||||||||||||||||||||||||||||||||||||
Upper GSSP ratified | 2006 [4] |
The Wuliuan stage is the fifth stage of the Cambrian, and the first stage of the Miaolingian Series of the Cambrian. It was formally defined by the International Commission on Stratigraphy in 2018. [5] Its base is defined by the first appearance of the trilobite species Oryctocephalus indicus ; it ends with the beginning of the Drumian Stage, marked by the first appearance of the trilobite Ptychagnostus atavus around 504.5 million years ago. [6]
The 'golden spike' that formally defines the base of the age is driven into the Wuliu-Zengjiayan (乌溜-曾家崖) section of the Kaili formation, near Balang Village in the Miaoling Mountains, Guizhou, China. [7]
Three sections were discussed as GSSP candidates: the Wuliu-Zengjiayan section near Balang in Guizhou province (China), a section on Split Mountain in Nevada (USA) [8] and the "Molodo river section" along the Molodo river (Sakha Republic, Russia). [9] The Wuliu-Zengjiayan section is an outcrop of the Kaili Formation in the Wuliu quarry. The first candidate for the beginning of the Wuliuan was the trilobite Oryctocephalus indicus , the second candidate was the trilobite Ovatoryctocara granulata . [10]
The Wuliu-Zengjiayan section was chosen as the formal base in 2018, with the first appearance of Oryctocephalus indicus being chosen as the defining marker for the GSSP. [2]
The base of Wuliuan stage (and, accordingly, the entire Miaolingian Series) is characterized by the first major extinction of trilobites, known as the Olenellid Biomere boundary. This event is linked by a sudden negative carbonate carbon excursion. [11]
Benthic graptolites have reached a considerable diversity im the Wuliuan. The most common graptolite genus of this age is Sphenoecium , whose robust colonies were found all over the world. [12] Numerous panarthropods, including trilobites, agnostoids, hurdiids and bradoriids, are known from Wuliuan deposits. [13]
The Furongian is the fourth and final epoch and series of the Cambrian. It lasted from 497 to 485.4 million years ago. It succeeds the Miaolingian series of the Cambrian and precedes the Lower Ordovician Tremadocian Stage. It is subdivided into three stages: the Paibian, Jiangshanian and the unnamed 10th stage of the Cambrian.
The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite species Undulograptus austrodentatus around 467.3 million years ago. It lasted for about 8.9 million years until the beginning of the Sandbian around 458.4 million years ago. This stage of the Ordovician was marked by the beginning of the Andean-Saharan glaciation.
The Paibian is the lowest stage of Furongian series of the Cambrian. It follows the Guzhangian and is succeeded by the Jiangshanian Stage. The base is defined as the first appearance of the trilobite Glyptagnostus reticulatus around 497 million years ago. The top, or the base of the Jiangshanian is defined as the first appearance of the trilobite Agnostotes orientalis around 494 million years ago.
The Kaili Formation(凯里組) is a stratigraphic formation which was deposited during the Lower and Middle Cambrian. The formation is approximately 200 metres (660 ft) thick and was named after the city Kaili in the Guizhou province of southwest China.
The Terreneuvian is the lowermost and oldest series of the Cambrian geological system. Its base is defined by the first appearance datum of the trace fossil Treptichnus pedum around 538.8 million years ago. Its top is defined as the first appearance of trilobites in the stratigraphic record around 521 million years ago. This series' name was formally accepted by the International Commission on Stratigraphy in 2007.
The Fortunian age marks the beginning of the Phanerozoic Eon, the Paleozoic Era, and the Cambrian Period. It is the first of the two stages of the Terreneuvian series. Its base is defined as the first appearance of the trace fossil Treptichnus pedum 538.8 million years ago. The top of the Fortunian which is the base of the Stage 2 of the Cambrian has not been formally defined yet, but will correspond to the appearance of an Archeocyatha species or "Small shelly fossils" approximately 529 million years ago.
The Jiangshanian is the middle stage of the Furongian series. It follows the Paibian Stage and is succeeded by the still unnamed Stage 10 of the Cambrian. The base is defined as the first appearance of the trilobite Agnostotes orientalis which is estimated to be 494 million years ago. The Jiangshanian lasted until approximately 489.5 million years ago.
The Guzhangian is an uppermost stage of the Miaolingian Series of the Cambrian. It follows the Drumian Stage and precedes the Paibian Stage of the Furongian Series. The base is defined as the first appearance of the trilobite Lejopyge laevigata around 500.5 million years ago. The Guzhangian-Paibian boundary is marked by the first appearance of the trilobite Glyptagnostus reticulatus around 497 million years ago.
The Drumian is a stage of the Miaolingian Series of the Cambrian. It succeeds the Wuliuan and precedes the Guzhangian. The base is defined as the first appearance of the trilobite Ptychagnostus atavus around 504.5 million years ago. The top is defined as the first appearance of another trilobite Lejopyge laevigata around 500.5 million years ago.
Oryctocephalus indicus is a species of corynexochid trilobite from the Cambrian. Its first appearance is proposed for the lower boundary of the Wuliuan, which corresponds to the beginning of the Miaolingian. The species was first described by the British paleontologist Frederick Richard Cowper Reed in 1910 as Zacanthoides indicus. It was transferred to the genus Oryctocephalus by the American paleontologist Charles Elmer Resser in 1938.
Ptychagnostus atavus is a species of agnostid trilobite. It was originally described by Swedish paleontologist Sven Axel Tullberg as Agnostus atavus in 1880. It is used in biostratigraphy as an index fossil. Its first appearance at the GSSP section in the Wheeler Shale of Utah is defined as the beginning of the Drumian Age of the Miaolingian.
Stage 10 of the Cambrian is the still unnamed third and final stage of the Furongian series. It follows the Jiangshanian and precedes the Ordovician Tremadocian Stage. The proposed lower boundary is the first appearance of the trilobite Lotagnostus americanus around 489.5 million years ago, but other fossils are also being discussed. The upper boundary is defined as the appearance of the conodont Iapetognathus fluctivagus which marks the beginning of the Tremadocian and is radiometrically dated as 485.4 million years ago.
Lejopyge laevigata is a species of agnostid trilobite belonging to the genus Lejopyge. It existed during the Guzhangian to the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy.
The Miaolingian is the third Series of the Cambrian Period, and was formally named in 2018. It lasted from about 509 to 497 million years ago and is divided in ascending order into 3 stages: the Wuliuan, Drumian, and Guzhangian. The Miaolingian is preceded by the unnamed Cambrian Series 2 and succeeded by the Furongian series.
Glyptagnostus reticulatus is a species of agnostid trilobite belonging to the genus Glyptagnostus. It existed during the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy. It was characterized by an unusual net-like pattern of furrows on both the cephalon and the pygidium.
Cambrian Stage 4 is the still unnamed fourth stage of the Cambrian and the upper stage of Cambrian Series 2. It follows Cambrian Stage 3 and lies below the Wuliuan. The lower boundary has not been formally defined by the International Commission on Stratigraphy. One proposal is the first appearance of two trilobite genera, Olenellus or Redlichia. Another proposal is the first appearance of the trilobite species Arthricocephalus chauveaui. Both proposals will set the lower boundary close to 514 million years ago. The upper boundary corresponds to the beginning of the Wuliuan.
Agnostotes orientalis is a species of agnostid trilobite belonging to the genus Agnostotes. It existed during the Jiangshanian Age of the Cambrian. It is an important index fossil in biostratigraphy.
Cambrian Series 2 is the unnamed 2nd series of the Cambrian. It lies above the Terreneuvian series and below the Miaolingian. Series 2 has not been formally defined by the International Commission on Stratigraphy, lacking a precise lower boundary and subdivision into stages. The proposed lower boundary is the first appearance of trilobites which is estimated to be around 521 million years ago.
Ctenocystoidea is an extinct clade of echinoderms, which lived during the Cambrian and Ordovician periods. Unlike other echinoderms, ctenocystoids had bilateral symmetry, or were only very slightly asymmetrical. They are believed to be one of the earliest-diverging branches of echinoderms, with their bilateral symmetry a trait shared with other deuterostomes. Ctenocystoids were once classified in the taxon Homalozoa, also known as Carpoidea, alongside cinctans, solutes, and stylophorans. Homalozoa is now recognized as a polyphyletic group of echinoderms without radial symmetry. Ctenocystoids were geographically widespread during the Middle Cambrian, with one species surviving into the Late Ordovician.
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