Rostroconchia Temporal range: | |
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Technophorus sharpei | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Subphylum: | Conchifera |
Class: | † Rostroconchia Pojeta et al., 1972 [2] |
The Rostroconchia is a class of extinct molluscs dating from the early Cambrian to the Late Permian. They were initially thought to be bivalves, but were later given their own class. They have a single shell in their larval stage, and the adult typically has a single, pseudo-bivalved shell enclosing the mantle and muscular foot. The anterior part of the shell probably pointed downward and had a gap from which the foot could probably emerge. Rostroconchs probably lived a sedentary semi-infaunal lifestyle. There were probably more than 1,000 species of members of this class.
Approximately 3 dozen genera and an even greater number of species have been described. Generally, rostroconchs are small, less than two centimeters in length, but larger forms, found in United States Devonian limestones, can grow to a length of 15 cm.
Externally, rostroconchs look much like bivalves and rostroconchs probably had an extendable muscular foot, indicated by a prominent anterior gape in the rostroconch's shell. It seems, however, that the internal anatomy and morphology of the foot were closer to that of the scaphopods.
Rostroconchs began their life as a small, bilaterally symmetrical, univalved protoconch planktonic larva. The bilateral shell grew into two valves as the rostroconch entered adulthood. [1] Adult rostroconchs differ from bivalves because they have no functional hinge. Unlike the shell of a bivalve, which was able to move or articulate, the shell layers of a rostroconch — the layers of rigid calcite— continue across the whole dorsal area of the rostroconch. The two valves would have been rigidly fixed in place, and would have to have been broken periodically to allow the rostroconch shell to grow.
The posterior of the shell contains a flattened tube that is called the rostrum. The rostroconch probably burrowed itself into sediment, anterior first, leaving the rostrum above the sediment to be used as a water filtration system.
Heraultipegma is the earliest, very primitive, rostroconch genus dating from the Late Terreneuvian. True Rostroconchs appeared during the Ordovician, heavily competing with the bivalves until their decline in the end-early Ordovician turnover.
Early, primitive rostroconchs such as Ribeiroia had a hinge in which all shell layers covered the dorsal region resulting in a very rigid shell. In Conocardium , a more advanced rostroconch, the outer shell layers do not cross the entire margin, suggesting independent steps towards the bivalve flexible hinge.
Some evidence suggests that the conocardoid rostroconchs were the predecessors to the Scaphopoda.
Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. Shipworms bore into wood, clay, or stone and live inside these substances.
Kimberella is an extinct genus of bilaterian known only from rocks of the Ediacaran period. The slug-like organism fed by scratching the microbial surface on which it dwelt in a manner similar to the gastropods, although its affinity with this group is contentious.
A siphon is an anatomical structure which is part of the body of aquatic molluscs in three classes: Gastropoda, Bivalvia and Cephalopoda.
The tusk shells or tooth shells, technically the Scaphopoda, are members of a class of shelled marine mollusc with worldwide distribution, and are the only class of exclusively infaunal marine molluscs. Shells of species within this class range in length 0.5–18 cm (0.20–7.09 in). Members of the order Dentaliida tend to be larger than those of the order Gadilida.
A valve is each articulating part of the shell of a mollusc or another multi-shelled animal such as brachiopods and some crustaceans. Each part is known as a valve or in the case of chitons, a "plate". Members of two classes of molluscs, the Bivalvia (clams) and the Polyplacophora (chitons), have valves.
A bivalve shell is part of the body, the exoskeleton or shell, of a bivalve mollusk. In life, the shell of this class of mollusks is composed of two hinged parts or valves. Bivalves are very common in essentially all aquatic locales, including saltwater, brackish water, and freshwater. The shells of bivalves commonly wash up on beaches and along the edges of lakes, rivers, and streams. Bivalves by definition possess two shells or valves, a "right valve" and a "left valve", that are joined by a ligament. The two valves usually articulate with one another using structures known as "teeth" which are situated along the hinge line. In many bivalve shells, the two valves are symmetrical along the hinge line—when truly symmetrical, such an animal is said to be equivalved; if the valves vary from each other in size or shape, inequivalved. If symmetrical front-to-back, the valves are said to be equilateral, and are otherwise considered inequilateral.
Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.
The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.
Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.
Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.
Solemya velum, the Atlantic awning clam, is a species of marine bivalve mollusc in the family Solemyidae, the awning clams. This species is found along the eastern coast of North America, from Nova Scotia to Florida and inhabits subtidal sediments with high organic matter (OM) content and low Oxygen, such as salt ponds, salt marshes, and sewage outfalls.
Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.
Watsonella is an extinct genus of mollusc known from early (Terreneuvian) Cambrian strata. It has been hypothesized to be close to the origin of bivalves. It contains a single species, Watsonella crosbyi.
Fabulina fabula, the bean-like tellin, is a species of marine bivalve mollusc in the family Tellinidae. It is found off the coasts of northwest Europe, where it lives buried in sandy sediments.
Tellimya ferruginosa is a species of small marine bivalve mollusc in the family Lasaeidae. It is found on the eastern side of the Atlantic Ocean.
Similodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of eleven genera in the subfamily Praenuculinae. Similodonta is known from Middle Ordovician through Middle Silurian fossils found in Europe and North America. The genus currently contains eight accepted species, Similodonta ceryx, Similodonta collina, Similodonta djupvikensis, Similodonta magna, Similodonta recurva, Similodonta spjeldnaesi, Similodonta wahli and the type species Similodonta similis.
Paterinata is an extinct class of linguliform brachiopods which lived from the lower Cambrian ("Tommotian") to the Upper Ordovician (Hirnantian). It contains the single order Paterinida and the subfamily Paterinoidea. Despite being some of the earliest brachiopods to appear in the fossil record, paterinides stayed as a relatively subdued and low-diversity group even as other brachiopods diversified later in the Cambrian and Ordovician. Paterinides are notable for their high degree of convergent evolution with rhynchonelliform (articulate) brachiopods, which have a similar set of muscles and hinge-adjacent structures.
Trimerellida is an extinct order of craniate brachiopods, containing the sole superfamily Trimerelloidea and the families Adensuidae, Trimerellidae, and Ussuniidae. Trimerellidae was a widespread family of warm-water brachiopods ranging from the Middle Ordovician to the late Silurian (Ludlow). Adensuidae and Ussuniidae are monogeneric families restricted to the Ordovician of Kazakhstan. Most individuals were free-living, though some clustered into large congregations similar to modern oyster reefs.
Ostreoidea is a taxonomic superfamily of bivalve marine mollusc, sometimes simply identified as oysters, containing two families. The ostreoids are characterized in part by the presence of a well developed axial rod. Anal flaps are known to exist within the family Ostreidae but not within the more-primitive Gryphaeidae. The scar from the adductor muscle is simple, with a single, central scar. In the majority, the right valve is less convex than the left.
This article includes a list of general references, but it lacks sufficient corresponding inline citations .(February 2010) |