Patagorhacos Temporal range: Miocene (Colhuehuapian) | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Cariamiformes |
Family: | † Phorusrhacidae |
Genus: | † Patagorhacos Agnolin & Chafrat, 2015 [1] |
Species: | †P. terrificus |
Binomial name | |
†Patagorhacos terrificus Agnolin & Chafrat, 2015 [1] | |
Patagorhacos is an extinct genus of medium-sized phorusrhacid from the early Miocene of Patagonia. Currently only the single species Patagorhacos terrificus is known, which is represented by two highly fragmentary specimen, one belonging to the back of the skull and the other being the distal end of a leg bone. It was described together with the contemporary rheid Opisthodactylus horacioperezi .
Fossil remains of this phorusrhacid have been discovered in Paso Córdoba, a Natural Protection area in the Rio Negro Province of Argentina. The rocks of the area belong to the Chichinales Formation, which dates to the early Miocene (Colhuehuapian in accordance with South American land mammal age (SALMA) classification. [2] Patagorhacos is known from only two specimens. The holotype (MPCN-PV-377) is the distal end of a right quadrate, while the single referred specimen consists of the distal end of an ulna (MPCN-PV-379). Despite the lack in overlap, the ulna was assigned to Patagorhacos for its presence in the same locality and strata as the holotype quadrate and the fact that its size is consistent with the type specimen. [1]
The name Patagorhacos is a combination of Patagonia and rhacos, which was chosen for its use in the name of the type genus of the family, Phorusrhacos . The species name "terrificus" was chosen to represent the "terrific" nature of phorusrhacids. [1]
The quadrate of Patagorhacos is a robust bone belonging to a medium-sized Phorusrhacid. The body is comparably thin and compressed with strongly concave inner and outer margins. The pterygoid condyle meanwhile is strongly convex and very prominent, with a well-developed egg-shaped fossa on its rostrodorsal edge. In proximity to the condyle and separated by a thin bone wall are two more depressions, likely pneumatic fossae. The cotyle that receives the quadratojugal is deep and ovoid in shape and located above the lateral condyle. Towards the distal end of the bone three condyles can be seen, one medial, one lateral and one caudal, each separated through each other by grooves of varying width and depth. [1]
The ulna referred to Patagorhacos is poorly preserved and suffered heavily from erosion. However, it can be determined that the diaphysis was rather thin and the dorsal condyle well developed. When viewed from the side it's notably less prominent than that of Patagornis , Paraphysornis and Mesembriornis . [1]
Patagorhacos was found in the Patagonian Chichinales Formation, which is mostly known for its fossil mammals. The local fauna includes astrapotheres, the litoptern Cramauchenia , the notoungulates Colpodon (a leontiniid) and Cochilius volvens (an interatheriid), rodents ( Willidewu esteparius and Australoprocta ), xenarthrans and borhyaenoids. [3] Bird remains from the formation are comparatively poor. A part of a tibiotarsus has previously been classified as an undetermined species of psilopterine phorusrhacid, a relative of Patagorhacos. Other birds include an undetermined wading bird and Opisthodactylus horacioperezi , a species of rhea. During the Miocene the area likely consisted of open but wooded environment with temperate climate and a proximity to freshwater. [1]
Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous flightless birds that were among the largest apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 53 to 0.1 million years (Ma) ago.
Velocisaurus is a genus of noasaurid theropod dinosaur from the Late Cretaceous period of Argentina.
Epachthosaurus was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America.
Osteodontornis is an extinct seabird genus. It contains a single named species, Osteodontornis orri, which was described quite exactly one century after the first species of the Pelagornithidae was. O. orri was named after the naturalist Ellison Orr (1857-1951).
Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.
Brontornis is an extinct genus of giant bird that inhabited Argentina during the Early to Middle Miocene. Its taxonomic position is highly controversial, with authors alternatively considering it to be a cariamiform, typically a phorusrhacid or an anserimorph.
Cramauchenia is an extinct genus of litoptern South American ungulate. Cramauchenia was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus Macrauchenia. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian, as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan. In 1981 Soria made C. insolita a junior synonym of C. normalis. A specimen of C. normalis was described in 2010 from Cabeza Blanca in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA.
Paraphysornis is an extinct genus of giant flightless terror birds that inhabited Brazil during Late Oligocene or Early Miocene epochs. Although not the tallest phorusrhacid, Paraphysornis was a notably robust bird, having short and robust tarsal bones not suited for pursuit hunting.
Opisthodactylus is an extinct genus of rhea from the Early to Middle Miocene Santa Cruz and Chichinales Formations and the Late Miocene (Montehermosan) Andalhuala Formation of Argentina. Three species are described: the type species, O. patagonicus, O. kirchneri and O. horacioperezi. The species O. kirchneri was described in 2017 by Noriega et al. Fossils of O. horacioperezi were found together with fossils of Patagorhacos terrificus.
Kelenken is a genus of phorusrhacid, an extinct group of large, predatory birds, which lived in what is now Argentina in the middle Miocene about 15 million years ago. The only known specimen was discovered by high school student Guillermo Aguirre-Zabala in Comallo, in the region of Patagonia, and was made the holotype of the new genus and species Kelenken guillermoi in 2007. The genus name references a spirit in Tehuelche mythology, and the specific name honors the discoverer. The holotype consists of one of the most complete skulls known of a large phorusrhacid, as well as a tarsometatarsus lower leg bone and a phalanx toe bone. The discovery of Kelenken clarified the anatomy of large phorusrhacids, as these were previously much less well known. The closest living relatives of the phorusrhacids are the seriemas. Kelenken was found to belong in the subfamily Phorusrhacinae, along with for example Devincenzia.
Devincenzia is an extinct genus of giant flightless predatory birds in the family Phorusrhacidae or "terror birds" that lived during the Early Miocene (Deseadan) Fray Bentos Formation of Uruguay and Late Miocene (Huayquerian) Ituzaingó Formation to Early Pliocene (Montehermosan) of Argentina. The type species D. pozzi was formerly known as Onactornis pozzi. It stood about 2.5 metres (8.2 ft) tall, making it one of the largest Phorusrhacids and carnivorous birds known.
Patagornis is a genus of extinct flightless predatory birds of the family Phorusrhacidae. Known as "terror birds", these lived in what is now Argentina during the Early and Middle Miocene; the Santa Cruz Formation in Patagonia contains numerous specimens. Patagornis was an agile, medium sized Patagornithine and was likely a pursuit predator.
Psilopterus is an extinct genus of phorusrhacid from the Middle Oligocene to possibly the Late Pleistocene of Argentina and Uruguay. Compared to other phorusrhacids, members of the genus are both relatively gracile and diminutive, and include the smallest known species of terror bird: with the head raised P. bachmanni was 70–80 centimeters (2.3–2.6 ft) in height and weighed about 5 kilograms (11 lb), while the largest members of the genus were only about 8 kilograms (18 lb). The birds resemble the modern cariama, except with a heavier build and considerably smaller wings. The strong morphological similarity between the claws of the predatory cariama and Psilopterus, both of which are sharp, curved, and laterally compressed, may indicate they were used to strike prey. Tonni and Tambussi also suggested that, in contrast to the other, larger terror birds, Psilopterus could use their claws to climb trees, and could even fly, but this has been rejected in more recent literature. Fossil finds in Uruguay indicate the genus may have survived until 96,040 ± 6,300 years ago, millions of years after the larger phorusrhacids became extinct.
Aristonectes is an extinct genus of large elasmosaurid plesiosaurs that lived during the Maastrichtian stage of the Late Cretaceous. Two species are known, A. parvidens and A. quiriquensis, whose fossil remains were discovered in what are now Patagonia and Antarctica. Throughout the 20th century, Aristonectes was a difficult animal for scientists to analyze due to poor fossil preparation, its relationships to other genera were uncertain. After subsequent revisions and discoveries carried out from the beginning of the 21st century, Aristonectes is now recognised as the type genus of the subfamily Aristonectinae, a lineage of elasmosaurids characterized by an enlarged skull and a reduced length of the neck.
Dryornis, also called the Argentinian vulture, is an extinct genus of cathartid, known from Argentina. The genus contains two species, D. pampeanus and D. hatcheri.
Rhea mesopotamica is an extinct species of bird in the genus Rhea, whose living species are known as suris, rhea, or choiques. It lived in the Southern Cone of South America.
Annakacygna is a genus of flightless marine swan from the Miocene of Japan. Named in 2022, Annakacygna displays a series of unique adaptations setting it apart from any other known swan, including a filter feeding lifestyle, a highly mobile tail and wings that likely formed a cradle for their hatchlings in a fashion similar to modern mute swans. Additionally, it may have used both wings and tail as a form of display. All of these traits combined have led the researchers working on it to dub it "the ultimate bird". Two species are known, A. hajimei, which was approximately the size of a black swan, and A. yoshiiensis which exceeded the mute swan in both size and weight. The describing authors proposed the vernacular name Annaka short-winged swan for the genus.
The Cerro Bandera Formation is a geological formation in Neuquén Province, Argentina, in northern Patagonia, which dates to the Early Miocene, around 21 to 17.5 million years ago. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Cerro Bandera Formation include a variety of turtles and birds, also includes many mammals such as South American native ungulates as well as armadillos, and caviomorph rodents.
The Chichinales Formation is a geological formation in Río Negro Province, Argentina which dates from the Late Oligocene to the Early Miocene, around 23 to 17.5 million years ago. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Chichinales Formation, including a variety of turtles and birds, also includes many mammals such as South American native ungulates as well as armadillos, and caviomorph rodents.
Yatenavis is an extinct genus of enantiornithine bird from the Late Cretaceous Chorrillo Formation of Santa Cruz Province, Argentina. The genus contains a single species, Y. ieujensis, known from a partial humerus.