Bathornithidae

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Bathornithidae
Temporal range: Late Eocene to Early Miocene 37–20  Ma
Bathornis grallator restoration.jpg
Depiction of Bathornis grallator.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Cariamiformes
Family: Bathornithidae
J. Cracraft, 1968
Genera

Bathornithidae is an extinct family of birds from the Eocene to Miocene of North America. Part of Cariamiformes, they are related to the still extant seriemas and the extinct Phorusrhacidae. They were likely similar in habits, being terrestrial, long-legged predators, some of which attained massive sizes.

It has been suggested that most, if not all, North American Paleogene cariamiforme fossils are part of this group. [1] Storrs Olson also referred the European Elaphrocnemus to this clade, [2] though it has since been rejected. [3] Conversely, some analysis have instead recovered them as a polyphyletic group, with Bathornis and kin being sister taxa to phorusrhacids while Paracrax is rendered closer to modern seriemas, [4] though this assessment is heavily debated. [5]

The most recent consensus is that Bathornithidae is relegated exclusively to Bathornis , as a clade of Cariamiformes outside of a clade including seriemas and phorusrhacids, as well as a possible European specimen. Paracrax and Eutreptornis are understood to be odd taxa whose cariamiform affinities are not fully resolved. [6] A 2024 study however finds Bathornis as closer to seriemas than phorusrhacids were. [7]

Biology

Though some forms like Paracrax wetmorei might have been capable of flight, most taxa were flightless, [8] constituting examples of flightless birds in mammal dominated environments. Paracrax gigantea, Paracrax antiqua and the larger Bathornis species in particular might have occupied macropredatory niches akin to that of phorusrhacids, the former and latter reaching heights of over 2 m (6 ft 7 in)

Bathornis proper appears to have favoured wetland environments. It was a highly diverse genus, spanning a wide variety of species at various sizes, from the Eocene to the Miocene. [9]

Related Research Articles

<span class="mw-page-title-main">Gruiformes</span> Order of birds

The Gruiformes are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like".

<span class="mw-page-title-main">Seriema</span> Family of birds

The seriemas are the sole living members of the small bird family Cariamidae, which is also the only surviving lineage of the order Cariamiformes. Once believed to be related to cranes, they have been placed near the falcons, parrots and passerines, as well as the extinct Phorusrhacidae. The seriemas are large, long-legged territorial birds that range from 70–90 cm (28–35 in) in length. They live in grasslands, savanna, dry woodland and open forests of Brazil, Bolivia, Argentina, Paraguay and Uruguay. There are two species of seriemas, the red-legged seriema and the black-legged seriema. Names for these birds in the Tupian languages are variously spelled as siriema, sariama, and çariama, and mean "crested".

<span class="mw-page-title-main">Phorusrhacidae</span> Extinct family of flightless birds

Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous, mostly flightless birds that were among the largest apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 53 to 0.1 million years (Ma) ago, and perhaps even up to 21,600 ± 1,000 years ago.

<span class="mw-page-title-main">Plotopteridae</span> Extinct family of sea birds

Plotopteridae is an extinct family of flightless seabirds with uncertain placement, generally considered as member of order Suliformes. They exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant penguins. That they lived in the North Pacific, the other side of the world from the penguins, has led to them being described at times as the Northern Hemisphere's penguins, though they were not closely related. More recent studies have shown, however, that the shoulder-girdle, forelimb and sternum of plotopterids differ significantly from those of penguins, so comparisons in terms of function may not be entirely accurate. Plotopterids are regarded as closely related to Anhingidae (darters) and Phalacrocoracidae (cormorants). On the other hand, there is a theory that this group may have a common ancestor with penguins due to the similarity of forelimb and brain morphology. However, the endocast morphology of stem group Sphenisciformes differs from both Plotopteridae and modern penguins.

<i>Andalgalornis</i> Extinct genus of birds

Andalgalornis is a genus of flightless predatory birds of the extinct family Phorusrhacidae that lived in Argentina. The type and only species is A. steulleti.

<i>Strigogyps</i> Extinct genus of birds in the familie Ameghinornithidae

Strigogyps is an extinct genus of prehistoric bird from the Middle Eocene to Early Oligocene of France and Germany. It was probably around the size of a large chicken or a guan, weighing not quite 1 kilogram (2.2 lb). Apparently, as indicated by the ratio of lengths of wing to leg bones, S. sapea was flightless. Its legs were not adapted to running, so it seems to have had a walking lifestyle similar to trumpeters. Unlike other Cariamiformes, which appear to have been mostly carnivorous, Strigogyps specimens suggest a herbivorous diet.

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<i>Palaeotis</i> Extinct genus of birds

Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard, and gave it its consequent name. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes.

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1987.

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1976.

<i>Devincenzia</i> Extinct genus of birds

Devincenzia is an extinct genus of giant flightless predatory birds in the family Phorusrhacidae or "terror birds" that lived during the Early Miocene (Deseadan) Fray Bentos Formation of Uruguay, Late Miocene (Huayquerian) Ituzaingó Formation, Early Pliocene (Montehermosan) of Argentina, and possibly the Early Pleistocene Raigón Formation of Uruguay. The type species D. pozzi was formerly known as Onactornis pozzi. The largest possible specimen weighed up to 350 kilograms (770 lb), making it one of the largest phorusrhacids and carnivorous birds known.

<i>Patagornis</i> Extinct genus of birds

Patagornis is a genus of extinct flightless predatory birds of the family Phorusrhacidae. Known as "terror birds", these lived in what is now Argentina during the Early and Middle Miocene; the Santa Cruz Formation in Patagonia contains numerous specimens. Patagornis was an agile, medium sized Patagornithine and was likely a pursuit predator.

Cunampaia is a dubious genus of extinct mesoeucrocodylian. Fossils have been found from the Divisadero Largo Formation of Mendoza Province, Argentina, and date back to the Divisaderan to Tinguirirican regional South American Land Mammal Age of the Late Eocene epoch.

<span class="mw-page-title-main">Cariamiformes</span> Order of birds

Cariamiformes is an order of primarily flightless birds that has existed for over 60 million years. The group includes the family Cariamidae (seriemas) and the extinct families Phorusrhacidae, Bathornithidae, Idiornithidae and Ameghinornithidae. Though traditionally considered a suborder within Gruiformes, both morphological and genetic studies show that it belongs to a separate group of birds, Australaves, whose other living members are Falconidae, Psittaciformes and Passeriformes.

Paracrax is a genus of extinct North American flightless birds, possibly related to modern seriemas and the extinct terror birds. Part of Bathornithidae, it is a specialised member of this group, being cursorial carnivores much like their South American cousins, some species attaining massive sizes.

Eutreptornis is a genus of extinct possible cariamiforme bird from the Late Eocene of Utah. It is traditionally considered to be a bathornithid, though a combination of the relative incompleteness of the material alongside some differences from other bathornithids have raised some suspicions about this affiliation.

<i>Bathornis</i> Extinct genus of birds

Bathornis is an extinct lineage of birds related to modern day seriemas, that lived in North America about 37–20 million years ago. Like the closely related and also extinct phorusrhacids, it was a flightless predator, occupying predatory niches in environments classically considered to be dominated by mammals. It was a highly diverse and successful genus, spanning a large number of species that occurred from the Priabonian Eocene to the Burdigalian Miocene epochs.

Elaphrocnemus is a genus of extinct bird from the Eocene and Oligocene periods of Europe. Part of Cariamiformes, its closest living relatives are seriemas, though it differs significantly from them, being a better flyer.

<span class="mw-page-title-main">Eogruidae</span> Extinct family of birds

Eogruidae is a family of large, flightless birds that inhabited Asia from the Eocene to Pliocene epochs. Related to modern ostriches, it was formerly thought to be related to cranes, limpkins and trumpeters and that the similarities with ostriches were due to similar speciations to cursoriality, with both groups showing reduced numbers of toes to two in some taxa. It has been suggested that competition from true ostriches has caused the extinction of these birds, though this has never been formally tested and several ostrich taxa do occur in the late Cenozoic of Asia and some species do occur in areas where ostrich fossils have also been found. It has been suggested that the family is paraphyletic, with Ergilornithidae more closely related to modern ostriches than to Eogrus or Sonogrus.

Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds. Recent research shows that these birds may actually be palaeognaths related to ostriches.

References

  1. Gerald Mayr (2009). Paleogene Fossil Birds
  2. Olson, Storrs L. (1985): Section X.A.I.b. The Tangle of the Bathornithidae. In: Farner, D.; King, J. & Parkes, K. (eds.): Avian Biology 8: 146–150. Academic Press, New York.
  3. Gerald Mayr (2009). Paleogene Fossil Birds
  4. Federico L. Agnolin (2009). "Sistemática y Filogenia de las Aves Fororracoideas (Gruiformes, Cariamae)" (PDF). Fundación de Historia Natural Felix de Azara: 1–79.
  5. Mayr, G., & Noriega, J. I. A well-preserved partial skeleton of the poorly known early Miocene seriema Noriegavis santacrucensis (Aves, Cariamidae). Acta palaeontologica Polonica, 60(3):589-598.
  6. Mayr, G. (2016). "Osteology and phylogenetic affinities of the middle Eocene North American Bathornis grallator—one of the best represented, albeit least known Paleogene cariamiform birds (seriemas and allies)". Journal of Paleontology. 90 (2): 357–374. Bibcode:2016JPal...90..357M. doi:10.1017/jpa.2016.45. S2CID   88936361.
  7. Thomas W. LaBarge, Jacob D. Gardner and Chris L. Organ, The evolution and ecology of gigantism in terror birds (Aves, Phorusrhacidae), Published:24 April 2024https://doi.org/10.1098/rspb.2024.0235
  8. Cracraft, J. (1968). "A review of the Bathornithidae (Aves, Gruiformes), with remarks on the relationships of the suborder Cariamae". American Museum Novitates (2326): 1–46. hdl:2246/2536.
  9. Benton, R. C.; Terry, D. O.; Evanoff, E.; McDonald, H. G. (25 May 2015). The White River Badlands: Geology and Paleontology. Indiana University Press. ISBN   978-0-253-01608-9.