Salamanca Formation

Last updated
Salamanca Formation
Stratigraphic range: latest Maastrichtian-Danian
(Tiupampan-Riochican)
~66.0–57  Ma
Unit of Río Chico Group
Underlies Peñas Coloradas Formation
(Río Chico Group)
Overlies Lago Colhué Huapí Formation
(Chubut Group)
Lithology
Primary Sandstone, mudstone
Other Limestone
Location
CountryFlag of Argentina.svg  Argentina
Extent Golfo San Jorge Basin
Type section
Region Chubut and Santa Cruz Provinces
Country Argentina
Map of Punta Peligro.png
Location of the Salamanca Formation within Argentina

The Salamanca Formation is a geologic formation in the Golfo San Jorge Basin of central Patagonia that yields well-preserved, well-dated fossils from the early Paleocene. Studies of these fossils are providing new data on plant and animal diversity following the end-Cretaceous extinction event. [1] [2]

Contents

The Salamanca Formation crops out in the San Jorge Basin in southern Chubut and northern Santa Cruz provinces, Argentina, overlying the Cretaceous Chubut Group and is part of the Paleocene and Eocene Río Chico Group. [3] [4] [5] [6] The formation yields abundant plant remains [7] [8] [9] [10] [11] [12] as well as fossils of invertebrates, marine macrofaunas, [13] reptiles, [14] [15] [16] [17] [18] and mammals. [2] [19] [20] [21] [22] [23] [24] [4] [3] The formation consists primarily of estuarine to shallow marine deposits. [25] [26]

Palynological analysis of the Salamanca Formation shows low floral diversity after the end-Cretaceous mass extinction, followed by a rapid recovery. 50% of all pollen types are angiosperms, whereas gymnosperms accounted for only ~13% of total richness; however, Classopollis pollen, representing the extinct conifer family Cheirolepidiaceae, is the most abundant palynomorph. [3] Wood assemblages from the Salamanca Formation are dominated by conifers, but the presence of fossil angiosperm woods indicate that they were also part of the canopy. [8] [9] The co-occurrence of palms, dicot woods with indistinct growth rings, and alligatorids, [15] [16] [17] [18] indicates temperature remained above freezing year-round. The results of leaf physiognomic analyses indicate that the climate in the San Jorge Basin during the early Paleocene was warm subtropical. [27] [28] [29] [30]

Paleoflora

NameSpeciesLocalityMaterialNotesImage
Agathis [31] A. immortalisLeaves, twigs, pollen cones, ovuliferous complexes, pollen, seedsA conifer belonging to Araucariaceae. Oldest known species of Agathis [31]
Akania [10] A. sp.Palacio de los LorosA flowering plant of the family Akaniaceae
Azolla [32] A. keujaPalacio de los LorosA fern belonging to the family Salviniaceae
Banaraphyllum [10] B. ovatumPalacio de los LorosA flowering plant
Cissites [10] C. patagonicumPalacio de los LorosA flowering plant belonging to Vitaceae, related to the modern genus Cissus
Classopollis [3] C. sp.Palacio de los LorosPollen belonging to the extinct conifer family Cheirolepidiaceae
Cunoniantha [33] C. bicarpellataA flowering plant belonging to the family Cunoniaceae
Dryophyllum [10] D. australisPalacio de los LorosA flowering plant
Fagophyllum [10] F. duseniPalacio de los LorosLeaves, related to Fagaceae
Kirketapel [34] K. salamanquensisA scale-leaved conifer belonging to the family Podocarpaceae.
Lacinipetalum [35] L. spectabilumPalacio de los LorosA flowering plant belonging to the family Cunoniaceae
Laurophyllum [10] L. chubutensis

L. piatnitzkyi

Palacio de los LorosA flowering plant belonging to the family Lauraceae
Lygodium [10] Sp.A fern belonging to the Schizaeales, another indeterminate distinct fern is also present
Myrica [10] M. premiraPalacio de los LorosA flowering plant belonging to the family Myricaceae
Nothofagus [10] sp.Leaves
Notiantha [1] N. grandensisRancho GrandeFlowersA flowering plant belonging to the family Rhamnaceae
Palaeophytocrene [36] P. gaEstancia Las VioletasA fruitA floweing plant belonging to the family Icacinaceae
Paracacioxylon [8] P frenguelliiPalacio de los LorosWoodA flowering plant belonging to the family Mimosoideae
Paranymphaea [10] P aristolochiaformisPalacio de los LorosA flowering plant belonging to the family Nymphaeaceae
Sterculia [10] [37] S. acuminatalobaPalacio de los LorosA flowering plant belonging to the family Malvaceae
Stephania [38] S. psittacaPalacio de los LorosA flowering plant belonging to the family Menispermaceae
Suessenia [1] S. grandensisRancho GrandeLeavesA flowering plant belonging to the family Rhamnaceae
Wilkinsoniphyllum [38] W. menispermoides Palacio de los LorosLeavesA flowering plant belonging to the family Menispermaceae
Tripylocarpa [39] T. aestuariaSeedA flowering plant belonging to the palm tribe Cocoseae
Volkheimerites [40] V. labyrinthusFlowering plant pollen of uncertain placement
Fabaceae [10]
Sapindaceae [10]
Urticaceae [10]
Rosaceae [10]
Podocarpaceae [10] Leaves and cones

Paleofauna

Fish

NameSpeciesLocalityMaterialNotesImage
Xiphactinus [41] X. sp.A member of the Ichthyodectiformes.
Xiphactinus audax.png

Amphibians

NameSpeciesLocalityMaterialNotesImage
Calyptocephalella [42] C. sabrosaPunta PeligroA frog belonging to the family Calyptocephalellidae.
Gigantobatrachus [43] G. casamiquelaiPunta PeligroA frog

Reptiles

Testudines

NameSpeciesLocalityMaterialNotesImage
Hydromedusa [44] H. casamayorensisPunta PeligroA member of the family Chelidae.
Najadochelys [45] N. patagonicaPunta Peligro
Peligrochelys [46] [47] P. walshaePunta PeligroA member of Meiolaniformes
Salamanchelys [48] S. palaeocenicaPunta PeligroA member of the family Chelidae.
Trionyx T. argentinaPunta PeligroA member of the family Trionychidae
Yaminuechelys [45] Y. maiorPunta PeligroA member of the family Chelidae.

Crocodylomorphs

NameSpeciesLocalityMaterialNotesImage
Eocaiman [15] [16] E. palaeocenicusPunta PeligroA caimanine alligatorid.
Necrosuchus [18] N. ionensisEstancia Las Violetas
Necrosuchus ionensis.jpg
Protocaiman [17] P. peligrensisPunta Peligro
Protocaiman skull.jpg

Lepidosaurs

NameSpeciesLocalityMaterialNotesImage
Kawasphenodon [49] K. peligrensisPunta PeligroA sphenodontian

Mammals

Monotremes

NameSpeciesLocalityMaterialNotesImage
Monotrematum [50] [51] M. sudamericanumPunta PeligroTeeth from the upper and lower jaws [52] A monotreme

Allotheres

NameSpeciesLocalityMaterialNotesImage
Sudamerica [53] S. ameghinoiPunta PeligroA gondwanathere
Sudamerica jaw.svg

Meridiolestidans

NameSpeciesLocalityMaterialNotesImage
Peligrotherium [2] [54] P. tropicalisPunta PeligroA member of the clade Mesungulatoidea.
Peligrotherium.jpg

Meridiungulates

NameSpeciesLocalityMaterialNotesImage
Escribania [21] [23] E. chubutensis

E. talonicuspsis

Punta PeligroA member of the family Didolodontidae.
Escribania chubutensis.jpg
Raulvaccia [21] R. peligrensisPunta PeligroA member of the family Didolodontidae.
Requisia [24] R. vidmariPunta PeligroA member of the family Notonychopidae.

Metatherians

NameSpeciesLocalityMaterialNotesImage
Derorhynchus [55] D. aff. minutusPunta PeligroA metatherian of uncertain affinities
Didelphopsis [55] Indeterminate
?Sparassodonta [55] IndeterminateAstragalus
Polydolopimorphia [55] IndeterminateTwo taxa, one of which is a member of Bonapartheriidae
Metatheria [55] IndeterminateRight petrosal (MPEF-PV 2235)

Invertebrates

After [56] unless otherwise noted

NameSpeciesMaterialNotesImage
Acesta A bivalve belonging to Limidae
Arca A bivalve belonging to Arcidae
? Astarte A bivalve belonging to Astartidae
Aporrhaidae IndeterminateGastropod
Austrophaera A bivalve
Barbatia A bivalve belonging to Arcidae
Bathytormus A bivalve belonging to Crassatellidae
Buccinidae Four distinct generaGastropod
Cancellariidae IndeterminateA gastropod
Calyptraeidae IndeterminateGastropod
Cardites A bivalve belonging to Carditidae
Cavoscala A gastropod belonging to Epitoniidae
Claibornicardia A bivalve belonging to Carditidae
Costacopluma [57] C. salamancaA crab belonging to the family Retroplumidae
Crenilabium A gastropod belonging to Acteonidae
Cubitostrea A bivalve belonging to Ostreidae
Cucullaea Bivalve
Darwinices A gastropod belonging to Naticidae
? Dosinia A bivalve belonging to Veneridae
Fusinus A gastropod belonging to Fasciolariidae
Fyfea A gastropod
Globisininae IndeterminateGastropods belonging to the family Naticidae
Glycymerita A bivalve belonging to Glycymerididae
Gryphaostrea A bivalve
Gyroscala A gastropod belonging to Epitoniidae
Harpidae Nov. Gen.A gastropod
Heteroterma A gastropod belonging to Tudiclidae
?Larma A bivalve belonging to Veneridae
Ledina A bivalve belonging to Nuculanidae
Leionucula A bivalve belonging to Nuculidae
Lucinidae IndeterminateBivalve
Marwickia A bivalve belonging to Veneridae
Microfulgur A gastropod belonging to Fasciolariidae
Neilo A bivalve belonging to Malletiidae
Nucula A bivalve belonging to Nuculidae
Panopea A bivalve belonging to Hiatellidae
Patellogastropoda IndeterminateGastropod
Perissodonta A gastropod belonging to Struthiolariidae
Phacoides A bivalve belonging to Lucinidae
Pinna A bivalve belonging to Pinnidae
Priscaphander A gastropod belonging to Scaphandridae
Priscophicus A gastropod belonging to Ficidae
Pseudomaura A gastropod
Pteromyrtea A bivalve belonging to Lucinidae
Pseudotylostoma A gastropod
Pycnodonte A bivalve belonging to Gryphaeidae
Ringiculidae IndeterminateA gastropod
Rocalaria A gastropod belonging to Fasciolariidae
Rotundicardia A bivalve belonging to Carditidae
Sacella A bivalve belonging to Nuculanidae
Spineilo A bivalve belonging to Malletiidae
Struthioptera A gastropod belonging to Aporrhaidae
Sulcobuccinum A gastropod belonging to Pseudolividae
Taioma A gastropod belonging to Fasciolariidae
Talochlamys A bivalve belonging to Pectinidae
Tejonia A gastropod belonging to Naticidae
Titomaya [58] T. longobucca & T. chalcedonicaA bivalve belonging to Venerida
Tornatellaea A gastropod belonging to Acteonidae
Turritellidae Nov. gen.A gastropod
Venericardia A bivalve belonging to Carditidae
Volutidae IndeterminateGastropod
Yoldia A bivalve belonging to Yoldiidae
? Zemacies A gastropod belonging to Borsoniidae

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