Laventan

The Laventan (Spanish : Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age. ^{[1] [2]}

Etymology

The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group. ^[2]

Formations

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Locations of Laventan formations

Formation bold is type	Country	Basin	Notes
Honda Group	Colombia	Upper Magdalena Valley	[3]
Honda Group	Bolivia	Quebrada Honda Basin	[4]
Aisol Formation	Argentina	San Rafael Block	[5]
Bahía Inglesa Formation	Chile	Caldera Basin	[6]
Capadare Formation	Venezuela	Falcón Basin	[7]
Choquecota Formation	Bolivia	Altiplano Basin	[8]
Cura-Mallín Group	Chile	Cura-Mallín Basin	[9]
Ipururo Formation	Peru	Ucayali Basin	[10]
Paraná Formation	Argentina	Paraná Basin	[11]
Pebas Formation	Brazil   Colombia   Ecuador   Peru	Amazon Basin	[10] [12]
Pisco Formation	Peru	Pisco Basin	[13]
Santa Inés Formation	Venezuela	Eastern Venezuela Basin	[14]
Sincelejo Formation	Colombia	Sinú-San Jacinto Basin	[15]
Socorro Formation	Venezuela	Falcón Basin	[16]
Urumaco Formation	Venezuela	Falcón Basin	[17]

Fossil content

Group	Fossils	Formation	Notes
Mammals	lesser bulldog bat (Noctilio albiventris), LaVal's disk-winged bat (Thyroptera lavali), Spix's disk-winged bat (Thyroptera tricolor), Acarechimys minutissimus , Anachlysictis gracilis , Anadasypus hondanus , Aotus dindensis , Boreostemma acostae , B. gigantea , Brievabradys laventensis , Cebupithecia sarmientoi , Dukecynus magnus , Eodolichotis elachys , E. maddeni , Glossotheriopsis pascuali , Granastrapotherium snorki , Hilarcotherium castanedaii , Hondadelphys fieldsi , Hondathentes cazador , Huilabradys magdaleniensis , Huilatherium pluriplicatum , Kiotomops lopezi , Lagonimico conclucatus , Lycopsis longirostrus , Megadolodus molariformis , Micodon kiotensis , Micoureus laventicus , Microscleromys cribiphilus , M. paradoxalis , Microsteiromys jacobsi , Miocallicebus villaviejai , Miocochilius anomopodus , Mohanamico hershkovitzi , Mormopterus colombiensis , Nanoastegotherium prostatum , Neoglyptatelus originalis , Neonematherium flabellatum , Neoreomys huilensis , Neotamandua borealis , Notonycteris magdalenensis , N. sucharadeus , Nuciruptor rubricae , Pachybiotherium minor , Palynephyllum antimaster , Patasola magdalenae , Pedrolypeutes praecursor , Pericotoxodon platignathus , Pithiculites chenche , Potamops mascahehenes , Potamosiren magdalenensis , Prodolichotis guerreroi , P. pridiana , Prolicaphrium sanalfolsensis , Prothoatherium colombianus , Pseudoprepotherium confusum , Rhodanodolichotis antepridiana , R. vucetichae , Ricardomys longidens , Saimiri annectens , S. fieldsi , Scleromys colombianus , S. schurmanni , Scirrotherium hondaensis , Stirtonia tatacoensis , S. victoriae , Thylamys colombianus , Thylamys minutus , Thyroptera robusta , Villarroelia totoyoi , Xenastrapotherium kraglievichi , Cochilius sp. , Diclidurus sp. , Eumops sp. , Hapalops sp. , Olenopsis sp. , Theosodon sp. , ?Steiromys sp. , Dolichotinae sp., ?Echimyidae sp., Megalonychidae sp., Megatheriinae sp., Nothrotheriinae sp., Dasypodidae sp., Leontiniidae sp., Proterotheriidae indet., Toxodontidae sp.	Honda Group Colombia	[18] [19] [20] [21] [22] [23] [24] [25] [26] [27] [28] [29] [30] [31] [32] [33] [34] [35] [36] [37] [38] [39] [40] [41] [42] [43] [44] [45] [46] [47] [48] [49] [50] [51] [52] [53] [54] [55] [56] [57] [58] [59] [60] [61] [62] [63] [64] [65] [66] [67] [68] [69] [70] [71] [72] [73] [74] [75] [76] [77] [78] [79] [80]
	Acyon myctoderos , Australogale leptognathus , Chimeralestes ambiguus , Guiomys unica , Hapalops angustipalatus , Hemihegetotherium trilobus , Hiskatherium saintandrei , Hondalagus altiplanensis , Lakukullus anatisrostratus , Mesoprocta hypsodus , Miocochilius federicoi , Palaeothentes relictus , P. serratus , Propalaehoplophorus andinus , Quebradahondomys potosiensis , Acarechimys sp. , cf. Paratrigodon sp. , Prolagostomus sp. , cf. Prozaedyus sp. , Lagostominae indet., Mesotheriinae indet., Borhyaenidae indet., Caenolestidae indet., Capromyidae indet., Caviidae indet., Chinchillidae indet., Echimyidae indet., Euphractinae indet., Macraucheniidae indet., Octodontidae indet., Proterotheriidae indet., ? Xenastrapotherium indet.	Honda Group Bolivia	[81] [82] [83] [84] [85]
	Nesodon taweretus , cf. Astrapotherium sp. , cf. Theosodon sp. , Lomaphorini indet., Mylodontinae indet., Propalaehoplophorinae indet., Toxodontidae indet.	Aisol	[86] [87]
	Microtypotherium choquecotense , Prozaedyus sp. , Chinchillidae indet., Glyptodontidae indet., Hegetotheriidae indet.	Choquecota	[88]
	Heteropsomyinae (aff. Acarechimys) sp. , Dasyproctidae (aff. Alloiomys) sp. , Prolagostomus sp. , Caviomorpha indet., Dasypodidae indet., Typotheria indet.	Cura-Mallín	[89]
	Drytomomys cf. aequatorialis , Granastrapotherium cf. snorki , Miocochilius anomopodus , Neoglyptatelus originalis , Parapropalaehoplophorus septentrionalis , Pericotoxodon cf. platignathus , Potamarchus murinus , cf. Theosodon sp. , Acarechimys sp. , Boreostemma sp. , Megathericulus sp. , cf. Tetramerorhinus sp. , Urumacotherium sp. , Xenastrapotherium sp. , Borhyaenoidea indet., Delphinida indet., Glyptodontidae indet., Macraucheniidae indet., Megalonychidae indet., Mylodontidae indet., Octodontoidea indet., Potamarchinae indet., Uruguaytheriinae indet.	Ipururo	[90] [91] [92] [93] [94] [95]
	Pontistes rectifrons , Prionodelphis rovereti , Properiptychus argentinus , Protautoga longidens , cf. Balaenoptera sp. , Chrysophrys sp. , Dioplotherium sp. , Balaenidae indet., Physeteridae indet., Sciaenidae indet., ?Sparidae indet.	Paraná	[96]
	Dinomyidae indet., ? Octodontoidea indet.	Pebas	[97]
	Boreostemma venezolensis	Santa Inés	[98]
	Neoglyptatelus sincelejanus	Sincelejo	[15] [99]
	Mirandabradys socorrensis , Megatheriidae indet.	Socorro	[100] [101]
	cf. Olenopsis sp. , Mylodontidae indet.	Urumaco	[102]
Birds	Aramus paludigrus , Galbula hylochoreutes , Hoazinoides magdalenae	Honda Group Colombia	[103] [104]
	Psilopteridae indet.	Aisol	[87]
	Pygoscelis calderensis , Spheniscus megaramphus , S. urbinai , Pelagornis sp. , aff. Thalassarche sp.	Bahía Inglesa	[105]
	Pelagornis cf. chilensis	Capadare	[106]
Reptiles & amphibians	cane toad (Bufo marinus), Chelus colombiana , Chelonoidis hesterna , Colombophis portai , Dracaena colombiana , ?Eunectes stirtoni , Podocnemis medemi , Gryposuchus colombianus , Langstonia huilensis , Purussaurus neivensis , Balanerodus logimus , Charactosuchus fieldsi , Mourasuchus atopus , Eocaiman sp. , Gavialis sp. , Sebecus sp. , Alligatoridae indet.	Honda Group Colombia	[107] [108] [109] [110] [111] [112] [113] [114] [115] [116] [117] [118] [119] [120] [121]
	Crocodylia sp.	Aisol	[87]
	Bairdemys thalassica	Capadare	[122]
	Barinasuchus arveloi	Ipururo	[90]
	Caiman wannlangstoni , Gnatusuchus pebasensis , Gryposuchus pachakamue , Purussaurus neivensis , Kuttanacaiman iquitosensis , Mourasuchus atopus , Paleosuchus sp. , Podocnemis sp. , Gavialoidea indet.	Pebas	[123] [97] [124]
	Colombophis spinosus , Ikanogavialis gameroi , Caiman sp. , Mourasuchus sp. , Purussaurus sp. , Thecachampsa sp. , Alethinophidia indet., Boinae indet.	Socorro	[100] [125] [126]
	Globidentosuchus cf. brachyrostris , Caiman sp. , Mourasuchus sp. , Purussaurus sp.	Urumaco	[127]
Fishes & crustaceans	Colossoma macropomum , Lepidosiren paradoxa , cf. Acanthicus , Arapaima sp. , Brachyplatystoma cf. B. vaillanti , Brachyplatystoma promagdalena , Sylviocarcinus piriformis , cf. Corydoras sp. , Hoplias sp. , cf. Hoplosternum sp. , Hydrolycus sp. , Phractocephalus hemioliopterus , Serrasalmus sp. , Pygocentrus sp. , or Pristobrycon sp. ( cf. Myletes sp. ), Ariidae indet., Characidae cf. Tetragonopterinae, Cichlidae indet., Doradidae indet., Loricariidae indet., Potamotrygonidae indet.	Honda Group Colombia	[128] [129] [130] [131]
	Megalodon, Carcharias acutissima , Carcharodon hastalis , Galeocerdo aduncus , Hemipristis serra , Carcharhinus sp. , Heterodontus sp. , Squalus sp. , Squatina sp. , Dasyatidae indet., Myliobatoidea indet., Potamotrygonidae indet.	Paraná	[96]
	cf. Hydrolycus sp. , Leporinus sp. , Potamotrygon sp. , Pristis sp. , Anostomidae indet.	Pebas	[97]
	Megalodon, Aetobatus arcuatus , Hemipristis serra , Mugil cephalus , Carcharhinus sp. , Citharichthys sp. , Galeocerdo sp. , Rhinoptera sp. , Sphyrna sp. , Symphurus sp.	Socorro	[132]
Insects	Macroteleia yaguarum , Sycorax peruensis	Pebas	[133]
Flora	Astroniumxylon parabalansae , Piptadenioxylon paraexcelsa , Solanumxylon paranensis	Paraná	[134]

Correlations

The Laventan (13.8 to 11.8 Ma) correlates with:

• NALMA
  • latest Barstovian (15.97-13.65 Ma) ^[135]
  • early Clarendonian (13.65-10.3 Ma) ^[136]
• Californian ages
  • Luisian (15.5-13.5 Ma)
• ELMA - Astaracian (15.97-11.608 Ma) ^[137]
• CPS ^[138]
  • Badenian (13.65-12.7 Ma) ^[139]
  • Sarmatian (12.7-11.608 Ma) ^[140]
• New Zealand stratigraphy - Southland epoch (15.9-10.92 Ma) ^[141]
  • Lillburnian (15.1-12.98 Ma) ^[142]
  • early Waiauan (12.98-10.92 Ma) ^[143]
• Australian ages
  • Bairnsdalian (15.0-10.5 Ma)
• Japanese ages
  • Tozawan (15.97-13.5 Ma)
• NMAC - Tunggurian (13.65-11.1 Ma) ^[144]

Laventan correlations in South America

Formation	Honda	Honda	Aisol	Cura-Mallín	Pisco	Ipururo	Pebas	Capadare	Urumaco	Inés	Paraná	Map
Basin	VSM	Honda	San Rafael	Caldera	Pisco	Ucayali	Amazon	Falcón		Venezuela	Paraná	Laventan (South America)
Country	Colombia	Bolivia	Argentina	Chile	Peru			Venezuela			Argentina
Boreostemma
Hapalops
Miocochilius
Theosodon
Xenastrapotherium
Mylodontidae
Sparassodonta
Primates
Rodents
Birds
Terror birds
Reptiles
megalodon
Flora
Insects
Environments	Fluvial			Fluvio-deltaic		Fluvio-lacustrine		Fluvio-deltaic			Fluvial	Laventan volcanoclastics Laventan fauna Laventan flora
Volcanic	Yes

References

1. Paleo Database: Laventan
2. Madden et al., 1997
3. Villarroel et al., 1996, p.54
4. Marshall & Sempere, 1991, p.635
5. Forasiepi et al., 2011, p.149
6. Le Roux et al., 2016, p.46
7. Penín & Villarroel, 2005, p.138
8. Marshall & Sempere, 1991, p.636
9. Utgé et al., 2009
10. Antoine et al., 2016, p.56
11. Martín Pérez, 2013, p.51
12. Wesselingh et al., 2006, p.304
13. Brand et al., 2011
14. Simpson, 1947, p.2
15. Villarroel & Clavijo, 2005, p.348
16. Linares, 2004, p.5
17. Linares, 2004, p.16
18. Croft, 2007, p.299
19. Croft, 2007, p.300
20. Croft, 2007, p.301
21. Croft, 2007, p.302
22. Croft, 2007, p.303
23. Meldrum & Kay, 1997
24. Organ & Lemelin, 2011
25. Rosenberger et al., 1991
26. Setoguchi & Rosenberger, 1988
27. Setoguchi, 1985
28. Takai et al., 2001
29. Tejedor, 2013, p.22
30. Defler, 2004, p.32
31. Cebupithecia sarmientoi at Fossilworks.org
32. Miocallicebus villaviejai at Fossilworks.org
33. Neosaimiri annectens at Fossilworks.org
34. Stirtonia tatacoensis at Fossilworks.org
35. Stirtonia victoriae at Fossilworks.org
36. Vallejo Pareja et al., 2015
37. Pardo Jaramillo, 2010
38. Granastrapotherium snorki, Sebecus sp. at Fossilworks.org
39. Villarroel, 2000, p.118
40. Kay & Madden, 1997, p.171
41. Neonematherium flabellatum at Fossilworks.org
42. Brievabradys laventensis at Fossilworks.org
43. Huilabradys magdaleniensis at Fossilworks.org
44. Czaplewski et al., 2003, p.278
45. Neotamandua borealis at Fossilworks.org
46. Pseudoprepotherium confusum at Fossilworks.org
47. Anadasypus hondanus at Fossilworks.org
48. Boreostemma acostae at Fossilworks.org
49. Boreostemma gigantea at Fossilworks.org
50. Neoglyptatelus originalis at Fossilworks.org
51. Pedrolypeutes praecursor at Fossilworks.org
52. Scirrotherium hondaensis at Fossilworks.org
53. Prolicaphrium sanalfolsensis at Fossilworks.org
54. Prothoatherium colombianus at Fossilworks.org
55. Duke Locality 33 at Fossilworks.org
56. Villarroelia totoyoi at Fossilworks.org
57. Villarroel, 1997
58. Villarroel et al., 1996, p.63
59. Hoffstetter, 1971, p.40
60. Suárez et al., 2015
61. Marshall, 1976
62. Hondadelphys fieldsi at Fossilworks.org
63. Pachybiotherium minor at Fossilworks.org
64. Thylamys colombianus at Fossilworks.org
65. Thylamys minutus at Fossilworks.org
66. Hondathentes cazador at Fossilworks.org
67. Pithiculites chenche at Fossilworks.org
68. Neoreomys huilensis at Fossilworks.org
69. Rhodanodolichotis antepridiana at Fossilworks.org
70. Scleromys colombianus at Fossilworks.org
71. Scleromys schurmanni at Fossilworks.org
72. Prodolichotis guerreroi at Fossilworks.org
73. UCMP V4936 at Fossilworks.org
74. Eodolichotis elachys at Fossilworks.org
75. Eodolichotis maddeni at Fossilworks.org
76. Rhodanodolichotis vucetichae at Fossilworks.org
77. Takai et al., 1991
78. Kiotomops lopezi at Fossilworks.org
79. Potamops mascahehenes at Fossilworks.org
80. Thyroptera robusta at Fossilworks.org
81. Engelman et al., 2016
82. Pujos et al., 2014
83. Hondalagus at Fossilworks.org
84. Quebrada Honda at Fossilworks.org
85. Engelman et al., 2018
86. Forasiepi et al., 2015
87. Cerro Aisol at Fossilworks.org
88. Choquecota at Fossilworks.org
89. Cerro los Pinos at Fossilworks.org
90. IN-DTC-20 at Fossilworks.org
91. IN-DTC-28 at Fossilworks.org
92. IN-DTC-32 at Fossilworks.org
93. Río Sepa at Fossilworks.org
94. SEP 005 at Fossilworks.org
95. IN 008 at Fossilworks.org
96. Pueblo Brugo to Diamante at Fossilworks.org
97. CTA-45 at Fossilworks.org
98. Banks of the Rio Guere at Fossilworks.org
99. Calle Fria-Segovia at Fossilworks.org
100. South East of 'Cerro los Chivatos' at Fossilworks.org
101. Mirandabradys Urumaco 01 at Fossilworks.org
102. Biozona 1 Loc 44-FU at Fossilworks.org
103. Rasmussen, 1997
104. Miller, 1953
105. Los Dedos at Fossilworks.org
106. Zumbador Cave at Fossilworks.org
107. Estes & Wassersug, 1963
108. Gryposuchus colombianus at Fossilworks.org
109. Sebecus huiensis at Fossilworks.org
110. Dracaena colombiana at Fossilworks.org
111. Purussaurus neivensis at Fossilworks.org
112. Charactosuchus fieldsi at Fossilworks.org
113. Mourasuchus atopus at Fossilworks.org
114. UCMP Locality V4524 at Fossilworks.org
115. Balanerodus logimus at Fossilworks.org
116. Eocaiman at Fossilworks.org
117. Cadena et al., 2008, p.1206
118. De la Fuente et al., 2013, p.102
119. Geochelone hesterna at Fossilworks.org
120. Hsiou et al., 2010
121. (in Spanish) Fauna de La Venta - serpientes
122. Ferreira et al., 2015
123. Salas Gismondi et al., 2015
124. IQ114 at Fossilworks.org
125. Quebrada Honda at Fossilworks.org
126. East of Capirote, road to Quebrada Honda at Fossilworks.org
127. Sur Quebrada Bejucal at Fossilworks.org
128. (in Spanish) Fauna de La Venta - flora, cangrejos y peces
129. Kay & Madden, 1997, p.185
130. Lundberg & Chernoff, 1992, p.5
131. Brachyplatystoma promagdalena at Fossilworks.org
132. Socorro at Fossilworks.org
133. Amazonian amber at Fossilworks.org
134. Toma Vieja at Fossilworks.org
135. Barstovian at Fossilworks.org
136. Clarendonian at Fossilworks.org
137. Astaracian at Fossilworks.org
138. Central Paratethys Stages at Fossilworks.org
139. Badenian at Fossilworks.org
140. Sarmatian] at Fossilworks.org
141. Southland epoch at Fossilworks.org
142. Lillburnian at Fossilworks.org
143. Waiauan at Fossilworks.org
144. Tunggurian at Fossilworks.org

Bibliography

Laventan

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Honda Group, Colombia

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• Takai, Masanaru; Federico Anaya; Hisashi Suzuki; Nobuo Shigehara, and Takeshi Setoguchi. 2001. A New Platyrrhine from the Middle Miocene of La Venta, Colombia, and the Phyletic Position of Callicebinae. Anthropological Science, Tokyo 109(4). 289. .
• Takai, Masanaru; Takeshi Setoguchi; Carlos Villarroel A.; Alberto Cadena G., and Nobuo Shigehara. 1991. A New Miocene Molossid Bat from La Venta, Colombia, South America. Memoirs of the Faculty of Science, Kyoto University. Series of Geology and Mineralogy 56. 1–9. Accessed 2017-05-04.
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• Villarroel A., Carlos S. 2000. Un nuevo Mylodontinae (Xenarthra, Tardigrada) en la Fauna de La Venta, Mioceno de Colombia: el estado actual de la Familia Orophodontidae. Revista de la Academia Colombiana de Ciencias Exactas Físicas y Naturales 24. 117–127. Accessed 2017-05-04.
• Villarroel A., Carlos S. 1997. La Estructura de la Dentición Caduca de Huilatherium pluriplicatum, Leontiniidae (Notoungulata) del Mioceno de Colombia. Geología Colombiana 22. 139–149. Accessed 2017-05-04.
• Villarroel A., Carlos; Takeshi Setoguchi; Jorge Brieva, and Carlos Macia. 1996. Geology of the La Tatacoa "Desert" (Huila, Colombia): Precisions on the Stratigraphy of the Honda Group, the Evolution of the "Pata High" and the Presence of the La Venta Fauna. Memoirs of the Faculty of Science, Kyoto University. Series of Geology and Mineralogy 58. 41–66. Accessed 2017-05-04.
• Wheeler, Brandon. 2010. Community ecology of the Middle Miocene primates of La Venta, Colombia: the relationship between ecological diversity, divergence time, and phylogenetic richness. Primates 51(2). 131–138. .

Honda Group, Bolivia

• Croft, Darin A. 2007. The Middle Miocene (Laventan) Quebrada Honda Fauna, southern Bolivia and a description of its Notoungulates. Palaeontology 50. 277–303. .
• Engelman, Russell K.; Federico Anaya, and Darin A. Croft. 2018. Australogale leptognathus, gen. et sp. nov., a second species of small sparassodont (Mammalia: Metatheria) from the middle Miocene locality of Quebrada Honda, Bolivia. Journal of Mammalian Evolution in press. _. Accessed 2018-10-01.
• Engelman, Russell K.; Federico Anaya, and Darin A. Croft. 2016. New palaeothentid marsupials (Paucituberculata) from the middle Miocene of Quebrada Honda, Bolivia, and their implications for the palaeoecology, decline and extinction of the Palaeothentoidea. Journal of Systematic Palaeontology 15(10). 787–820. Accessed 2019-02-12.
• Pujos, François; Gerardo De Iuliis; Bernardino Mamani Quispe, and Ruben Andrade Flores. 2014. Lakukullus anatisrostratus, gen. et sp. nov., a new massive nothrotheriid sloth (Xenarthra, Pilosa) from the middle Miocene of Bolivia. Journal of Vertebrate Paleontology 34(5). 1243–1248. Accessed 2019-02-12.

Aisol Formation

• Forasiepi, Analía M.; Esperanza Cerdeño; Mariano Bond; Gabriela I. Schmidt; Maximiliano Naipauer; Fiona R. Straehl; Agustín G. Martinelli; Alberto C. Garrido, and Mark D. Schmitz and James L. Crowley. 2015. New toxodontid (Notoungulata) from the Early Miocene of Mendoza, Argentina. Paläontologische Zeitschrift 89(3). 611–634. Accessed 2019-02-13.
• Forasiepi, Analía M.; Agustín G. Martinelli; Marcelo S. De la Fuente; Sergio Dieguez, and Mariano Bond. 2011. Paleontology and stratigraphy of the Aisol Formation (Neogene), San Rafael, Mendoza , 135–154. Cenozoic Geology of the Central Andes of Argentina, SCS Publisher, Salta. Accessed 2017-10-01.

Bahía Inglesa Formation

• Achurra, L.E.; J.P. Lacassie; J.P. Le Roux; C. Marquadt; M. Belmar; J. Ruiz Del Solar, and S.E. Ishman. 2009. Manganese nodules in the Miocene Bahía Inglesa Formation, north-central Chile: Petrography, geochemistry, genesis and palaeoceanographic significance. Sedimentary Geology 217(1). 128–139. Accessed 2017-08-15.
• Achurra, L.; C. Marquadt; J.P. Lacassie, and J.P. Le Roux. 2003. [ Nódulos de Mn de la Formación Bahía Inglesa: geoquímica, génesis e implicancias paleoambientales], 1–10. X Congreso Geológico Chileno. Accessed 2017-08-15.
• Acosta Hospitaleche, Carolina, and Jhoann Canto. 2005. Primer registro de cráneos asignados a Palaeospheniscus (Aves, Spheniscidae) procedentes de la Formación Bahía Inglesa (Mioceno Medio-Tardío), Chile. Revista Chilena de Historia Natural 78. 489–495. Accessed 2017-08-15.
• Benson, Richard D. 2015. A new species of penguin from the late Miocene of Chile, with comments on the stratigraphic range of Palaeospheniscus. Scientific Publications of the Science Museum of Minnesota 8(4). 5–22. Accessed 2019-02-13.
• Chávez Hoffmeister, Martín Felipe. 2008. La ornitofauna de la Formación Bahía Inglesa, Caldera, Chile (MSc. thesis) , 1–165. Universidad Austral de Chile. Accessed 2017-08-15.
• Chávez, Martín F. 2007. Sobre la presencia de Paraptenodytes y Palaeospheniscus (Aves: Sphenisciformes) en la Formación Bahía Inglesa, Chile. Revista Chilena de Historia Natural 80(2). 255–259. Accessed 2017-08-15.
• Chávez, Martín. 2007. Spheniscus megaramphus Stucchi et al. 2003 (Aves: Sphenisciformes) en la Formación Bahía Inglesa, Chile. Boletín de la Sociedad Geológica del Perú 102. 101–107. Accessed 2017-08-15.
• Le Roux, Jacobus P.; Luciano Achurra; Álvaro Henríquez; Catalina Carreño; Huber Rivera; Mario E. Suárez; Scott E. Ishman; Nicholas D. Pyenson, and Carolina S. Gutstein. 2016. Oroclinal bending of the Juan Fernández Ridge suggested by geohistory analysis of the Bahía Inglesa Formation, north-central Chile. Sedimentary Geology 333. 32–49. Accessed 2017-08-15.
• Le Roux, Jacobus P.; Luciano Achurra; Álvaro Henríquez; Catalina Carreño, and Huber Rivera. 2015. [ Lithostratigraphy, depositional environments and tectonic setting of the Bahía Inglesa Formation west of Copiapó, north-central Chile], 821–824. XIV Congreso Geológico Chileno. Accessed 2017-08-15.
• Oyanadel Urbina, Pablo; Jorge Carrillo Briceño; Jaime Villafaña; Victor Castelleto; Cristian Varas; Alex Alballay, and Marcelo Rivadeneira. 2015. [ Nuevo registro de familias de peces óseos en Formación Bahía Inglesa], 1–4. XIV Congreso Geológico Chileno. Accessed 2017-08-15.
• Staig, Felipe; Sebastián Hernández; Patricio López; Jaime A. Villafaña; Cristian Varas; Luis Patricio Soto, and Jorge D. Carrillo Briceño. 2015. Late Neogene Elasmobranch fauna from the Coquimbo Formation, Chile. Revista Brasileira de Paleontologia 18(2). 261–272. Accessed 2017-08-15.
• Suárez, Mario E. 2011. Línea de base paleontológica proyecto Puerto Santa Fe , 1–29. Comuna de Caldera Región de Atacama. Accessed 2017-08-15.
• Valenzuela Toro, Ana, and Carolina S. Gutstein. s.a. [ Mamíferos marinos fósiles (excepto Cetacea) de Formación Bahía Inglesa: Registro, implicancias paleoambientales y perspectivas de estudio]. . .. 770–772. Accessed 2017-08-15.

Capadare Formation

• Ferreira, Gabriel S.; Ascanio D. Rincón; Andrés Solórzano, and Max C. Langer. 2015. The last marine pelomedusoids (Testudines: Pleurodira): a new species of Bairdemys and the paleoecology of Stereogenyina. PeerJ 3. e1063. Accessed 2019-02-13.
• Penín Cendón, José Antonio, and Vanessa Ilia Villarroel Estaba. 2005. Integracrón y actualización geológica del área de Jacura y Capadare, Estado Falcón , 1–199. Universidad Central de Venezuela. Accessed 2017-08-15.

Choquecota and Mauri Formations

• Marshall, Larry G., and Thierry Sempere. 1991. The Eocene to Pleistocene vertebrates of Bolivia and their stratigraphic context: A review. Revista técnica de YPFB 12. 631–652. Accessed 2017-08-15.

Cura-Mallín Group

• Flynn, John J.; Reynaldo Charrier; Darin A. Croft; Phillip B. Gans; Trystan M. Herriott; Jill A. Wertheim, and André R. Wyss. 2008. Chronologic implications of new Miocene mammals from the Cura-Mallín and Trapa Trapa formations, Laguna del Laja area, south central Chile. Journal of South American Earth Sciences 26(4). 412–423. Accessed 2017-10-20.
• Shockey, Bruce J.; John J. Flynn; Darin A. Croft; Phillip Gans, and André R. Wyss. 2012. New leontiniid Notoungulata (Mammalia) from Chile and Argentina : comparative anatomy, character analysis, and phylogenetic hypotheses. American Museum Novitates (3737). 1–64. Accessed 2019-02-15.
• Utgé, Silvana; Andrés Folguera; Vanesa Litvak, and Víctor A. Ramos. 2009. Geología del sector norte de la Cuenca de Cura Mallín en las Lagunas de Epulaufquen, Neuquén. Revista de la Asociación Geológica Argentina 64(2). 231–248. Accessed 2017-10-01.

Gran Bajo del Gualicho Formation

• Reichler, Valeria A. 2010. Estratigrafía y paleontología del Cenozoico marino del Gran Bajo y Salinas del Gualicho, Argentina, y descripción de 17 especies nuevas. Andean Geology 31. 177–219. Accessed 2018-09-11.

Ipururo Formation

• Antoine, Pierre-Olivier; M. Alejandra Abello; Sylvain Adnet; Ali J. Altamirano Sierra; Patrice Baby; Guillaume Billet; Myriam Boivin; Ysabel Calderón, and Adriana Candela, Jules Chabain, Fernando Corfu, Darin A. Croft, Morgan Ganerød, Carlos Jaramillo, Sebastian Klaus, Laurent Marivaux, Rosa E. Navarrete, Maëva J. Orliac, Francisco Parra, María Encarnación Pérez, François Pujos, Jean-Claude Rage, Anthony Ravel, Céline Robinet, Martin Roddaz, Julia Victoria Tejada Lara, Jorge Vélez Juarbe, Frank P. Wesselingh, Rodolfo Salas Gismondi. 2016. A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru. Gondwana Research 31. 30–59. Accessed 2017-08-15.

Paraná Formation

• Cione, Alberto Luis; Daniel Alfredo Cabrera, and María Julia Barla. 2012. Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic. Geobios 45(2). 167–172. Accessed 2017-10-22.
• Martín Pérez, Leandro. 2013. Sistemática, tafonomía y paleoecología de los invertebrados de la Formación Paraná (Mioceno), Provincia de Entre Ríos, Argentina (PhD thesis) , 1–398. Universidad Nacional de La Plata. Accessed 2017-08-15.

Pebas Formation

• Antoine, Pierre-Olivier; M. Alejandra Abello; Sylvain Adnet; Ali J. Altamirano Sierra; Patrice Baby; Guillaume Billet; Myriam Boivin; Ysabel Calderón, and Adriana Candela, Jules Chabain, Fernando Corfu, Darin A. Croft, Morgan Ganerød, Carlos Jaramillo, Sebastian Klaus, Laurent Marivaux, Rosa E. Navarrete, Maëva J. Orliac, Francisco Parra, María Encarnación Pérez, François Pujos, Jean-Claude Rage, Anthony Ravel, Céline Robinet, Martin Roddaz, Julia Victoria Tejada Lara, Jorge Vélez Juarbe, Frank P. Wesselingh, Rodolfo Salas Gismondi. 2016. A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru. Gondwana Research 31. 30–59. Accessed 2017-08-15.
• Wesselingh, F.P.; M.C. Hoorn; J. Guerrero; M.E. Räsänen; L. Romero Pittmann, and J. Salo. 2006. The stratigraphy and regional structure of Miocene deposits in western Amazonia (Peru, Colombia and Brazil), with implications for late Neogene landscape evolution. Scripta Geologica 133. 291–322. Accessed 2017-08-15.

Pisco Formation

• Altamirano Sierra, Alí J. 2013. Primer registro de pelicano (Aves: Pelecanidae) para el Mioceno tardio de la formacion Pisco, Peru. Bulletin de l'Institut français d'études andines 42. 1–12. Accessed 2017-09-04.
• Báez Gómez, Diego A. 2006. Estudio paleoambiental de la formación Pisco:: Localidad Ocucaje. Revista del Instituto de Investigaciones FIGMMG 9. 64–69. Accessed 2017-09-04.
• Bianucci, Giovanni; Claudio Di Celma; Mario Urbina, and Olivier Lambert. 2016. New beaked whales from the late Miocene of Peru and evidence for convergent evolution in stem and crown Ziphiidae (Cetacea, Odontoceti). PeerJ 4. e2479. Accessed 2019-02-12.
• Brand, Leonard; Mario Urbina; Arthur Chadwick; Thomas J. DeVries, and Raul Esperante. 2011. A high resolution stratigraphic framework for the remarkable fossil cetacean assemblage of the Miocene/Pliocene Pisco Formation, Peru. Journal of South American Earth Sciences 31(4). 414–425. .
• Collareta, Alberto; Olivier Lambert; Christian De Muizon; Mario Urbina, and Giovanni Bianucci. 2017. Koristocetus pescei gen. et sp. nov., a diminutive sperm whale (Cetacea: Odontoceti: Kogiidae) from the late Miocene of Peru. Fossil Record 20(2). 259–278. Accessed 2019-02-12.
• Lambert, Olivier; Giovanni Bianucci; Mario Urbina, and Jonathan H. Geisler. 2017. A new inioid (Cetacea, Odontoceti, Delphinida) from the Miocene of Peru and the origin of modern dolphin and porpoise families. Zoological Journal of the Linnean Society 179(4). 919–946. .
• Marx, Felix G.; Olivier Lambert, and Christian De Muizon. 2017. A new Miocene baleen whale from Peru deciphers the dawn of cetotheriids. Royal Society Open Science 4(9). 170560. Accessed 2019-02-12.
• Marx, Felix G., and Naoki Kohno. 2016. A new Miocene baleen whale from the Peruvian desert. Royal Society Open Science 3(10). 160542. Accessed 2019-02-12.
• Poma Porras, Orlando; Edgard Horna Santillán, and Raúl Esperante. 2009. Baleen Fósil (Cetacea: mysticeti) en Sedimentos de la Cuenca Marina del Neógeno en la Formación Pisco, al Sur del Perú. Revista de Investigación Universitaria 1. 84–97. Accessed 2017-09-04.
• Ramassamy, Benjamin; Olivier Lambert; Alberto Collareta; Mario Urbina, and Giovanni Bianucci. 2018. Description of the skeleton of the fossil beaked whale Messapicetus gregarius: searching potential proxies for deep-diving abilities. Fossil Record 21(1). 11–32. Accessed 2019-02-11.
• Solís Mundaca, Flavio Alejandro. 2018. Bioestratigrafía e implicancias paleoceanográficas de las diatomeas de la sección Cerro Caucato, Formación Pisco, Ica, Peru (MSc. thesis) , 1–158. Universidad Peruana Cayetano Heredia. Accessed 2018-09-10.
• Stucchi, Marcelo; Steven D. Emslie; Rafael M. Varas Malca, and Mario Urbina Schmitt. 2015a. A new late Miocene condor (Aves, Cathartidae) from Peru and the origin of South American condors. Journal of Vertebrate Paleontology 35(5). e972507. Accessed 2019-02-13.
• Stucchi, Marcelo; Rafael M. Varas Malca, and Mario Urbina Schmitt. 2015b. New Miocene sulid birds from Peru and considerations on their Neogene fossil record in the Eastern Pacific Ocean. Acta Palaeontologica Polonica 61(2). 417–427. Accessed 2019-02-13.
• Stucchi, M. 2007. Los pingüinos de la Formación Pisco (Neógeno), Perú. 4th European Meeting on the Palaeontology and Stratigraphy of Latin America, Cuadernos del Museo Geominero 8. 367–373. Accessed 2017-09-04.

Santa Inés Formation

• Simpson, George Gaylord. 1947. A Miocene glyptodont from Venezuela. American Museum Novitates (1368). 1–10. .

Sincelejo Formation

• Villarroel A., Carlos, and Javier Clavijo. 2005. Los mamíferos fósiles y las edades de las sedimentitas continentales del Neógeno de la Costa Caribe Colombiana. Revista de la Academia Colombiana de Ciencias Exactas Físicas y Naturales 29. 345–356. Accessed 2019-02-04.

Socorro and Urumaco Formations

• Carrillo Briceño, Jorge D.; Erin Maxwell; Órangel A. Aguilera; Rodolfo Sánchez, and Marcelo R. Sánchez Villagra. 2015. Sawfishes and Other Elasmobranch Assemblages from the Mio-Pliocene of the South Caribbean (Urumaco Sequence, Northwestern Venezuela). PLoS ONE 10(10). e0139230. .
• Linares, Omar J. 2004. Bioestratigrafía de la fauna de mamíferos de las formaciones Socorro, Urumaco y Codore (Mioceno Medio-Plioceno Temprano) de la región de Urumaco, Falcón, Venezuela. Paleobiología Neotropical 1. 1–26. Accessed 2017-08-15.
• Quijano Ballesteros, Jhon Richard. 2005. Estudio magnetoestratigráfico en la sección de El Mamón (miembro medio de la Formación Urumaco, Estado Falcón) (MSc. thesis) , 1–92. Universidad Simón Bolívar. Accessed 2017-10-27.
• Rincón, Ascanio D.; Andrés Solórzano; H. Gregory McDonald, and Marisol Montellano Ballesteros. 2018. Two new megalonychid sloths (Mammalia: Xenarthra) from the Urumaco Formation (late Miocene), and their phylogenetic affinities. Journal of Systematic Palaeontology 17(5). 409–421. .
• Rincón, Ascanio D.; H. Gregory McDonald; Andrés Solórzano; Mónica Núñez Flores, and Damián Ruiz Ramoni. 2015. A new enigmatic Late Miocene mylodontoid sloth from northern South America. Royal Society Open Science 2(2). 140256. Accessed 2019-02-13.

Further reading

• Croft, Darin A. 2016. Horned Armadillos and Rafting Monkeys: The Fascinating Fossil Mammals of South America , 1–320. Indiana University Press ISBN   9780253020949. Accessed 2017-10-21.
• Fleagle, John G., and Alfred L. Rosenberger. 2013. The Platyrrhine Fossil Record , 1–256. Elsevier ISBN   9781483267074. Accessed 2017-10-21.
• Sánchez Villagra, Marcelo R.; Órangel A. Aguilera, and Alfredo A. Carlini. 2010. Urumaco and Venezuelan Paleontology: The Fossil Record of the Northern Neotropics , 1–304. Indiana University Press ISBN   9780253002006. Accessed 2017-10-21.
• Hartwig, W.C., and D.J. Meldrum. 2002. The Primate Fossil Record - Miocene platyrrhines of the northern Neotropics , 175–188. Cambridge University Press, ISBN   978-0-521-08141-2. Accessed 2017-09-24.
• Kay, Richard F.; Richard H. Madden; John J. Flynn, and Richard L. Cifelli. 1997. Vertebrate Paleontology in the Neotropics: The Miocene Fauna of La Venta, Colombia , 1–608. Smithsonian Institution Scholarly Press ISBN   9781935623854. Accessed 2017-10-21.
• Simpson, George Gaylord. 1996. Splendid Isolation: The Curious History of South American Mammals , 1–275. UMI ISBN   9780783733111. Accessed 2017-10-21.