Granastrapotherium

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Granastrapotherium
Temporal range: Middle Miocene (Laventan)
~13.8–11.8  Ma
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Granastrapotherium skull 2.jpg
Reconstructed skull of G. snorki
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Astrapotheria
Family: Astrapotheriidae
Subfamily: Uruguaytheriinae
Genus: Granastrapotherium
Johnson & Madden, 1997
Species:
G. snorki
Binomial name
Granastrapotherium snorki
Johnson & Madden, 1997

Granastrapotherium is an extinct genus of ungulate mammals, described from remains found in rocks of the Honda Group in the Tatacoa Desert, [1] in the Colombian departments of Huila and Tolima, at the Miocene fossil site La Venta. The only species formally recognized is Granastrapotherium snorki (from Spanish, gran, "great"; Astrapotherium , "lightning beast"; and snorkel, breathing tube, in reference to the trunk). Remains found in Bolivia and Peru, [2] seem to belong to Granastrapotherium or a very similar animal.

Contents

Description

Life reconstruction of a male Granastrapotherium snorki. Granastrapotherium reconstruction.jpg
Life reconstruction of a male Granastrapotherium snorki.

This astrapothere differs from its coeval, the uruguaytheriine astrapotheriid Xenastrapotherium by their larger size, between 3 and 4 tonnes, [3] with tusks about one meter long, making it one of the largest representatives of Astrapotheria, only surpassed by some species of Parastrapotherium . [4] Other differences include the presence of only one premolar, the lack of incisors in both jaws and the disposition of the canine tooth, which are very large and horizontal, which reminds much less of those of hippos and more of the tusks of some ancient relatives of elephants (such as Palaeomastodon ), although the defences in elephants and their relatives are not formed by the canines but the incisors. Similarly, the large nostrils appear extremely withdrawn on the skull, so this creature had to have a larger trunk than other astrapotheres. Most likely, like elephants, this animal used its muscular proboscis together with its tusks to cut leaves off trees and shrubs. [5]

Phylogeny

Cladogram based in the phylogenetic analysis published by Vallejo-Pareja et al., 2015, showing the position of Granastrapotherium: [6]

Eoastrapostylops

Trigonostylops

Tetragonostylops

Albertogaudrya

Scaglia

Astraponotus

Maddenia

Comahuetherium

Parastrapotherium

Astrapotheriinae

Astrapotherium

Astrapothericulus

Uruguaytheriinae

Uruguaytherium

Hilarcotherium

Xenastrapotherium

Granastrapotherium

Related Research Articles

<i>Trigonostylops</i>

Trigonostylops is an extinct genus of South American meridiungulatan ungulate, from the Late Paleocene to Late Eocene of South America and Antarctica. It is the only member of the family Trigonostylopidae.

La Venta is a fossil locality located in the modern departments of Tolima and Huila in Colombia. This site is one of the richest Neogene fossil assemblages in South America and represents the best-known Cenozoic fossil site outside of Argentina. It provides a glimpse of what life in the region was like before the main wave of the Great American Interchange.

<span class="mw-page-title-main">Astrapotheria</span> Extinct order of mammals

Astrapotheria is an extinct order of South American and Antarctic hoofed mammals that existed from the late Paleocene to the Middle Miocene, 59 to 11.8 million years ago. Astrapotheres were large, rhinoceros-like animals and have been called one of the most bizarre orders of mammals with an enigmatic evolutionary history.

The Laventan age is a period of geologic time within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.

<i>Astrapotherium</i> Extinct genus of mammals

Astrapotherium is an extinct genus of large astrapotherian ungulate native to South America during the early-middle Miocene. It is the best known member of the group The type species. A. magnus have been found in the Santa Cruz Formation in Argentina. Other fossils have been found in the Deseado, Sarmiento, and Aisol Formations of Argentina and Chile.

<span class="mw-page-title-main">Astrapotheriidae</span> Extinct family of mammals

Astrapotheriidae is an extinct family of herbivorous South American land mammals that lived from the Late Eocene to the Middle Miocene 37.71 to 15.98 million years ago. The most derived of the astrapotherians, they were also the largest and most specialized mammals in the Tertiary of South America. There are two sister taxa: Eoastrapostylopidae and Trigonostylopidae.

<i>Xenastrapotherium</i>

Xenastrapotherium is an extinct genus of astrapothere, a type of hoofed herbivorous mammal, native to South America, which lived in the Middle to Late Miocene period, typically during the Laventan stage. It is a member of the family Astrapotheriidae in the subfamily Uruguaytheriinae, large astrapotheres, equipped with a trunk-like nose and protruding teeth, similar to the elephants, but their tusks were the canine teeth, not the incisors. Xenastrapotherium was a genus widely distributed in northern South America, in contrast to other species of astrapotheres which lived in the area of the Southern Cone of the continent. It differed from other astrapotheres by having two lower incisors on each side of the jaw and the tusks have a pronounced longitudinal curvature, although their general shape and size are probably very similar to Astrapotherium, whose weight would be 900 to 1,500 kilograms, comparable to the current black rhinoceros.

Uruguaytherium is an extinct genus of astrapotherid mammal from the Late Oligocene to Early Miocene of South America. It was named by the Argentinean paleontologist Lucas Kraglievich in 1928, from a fragmentary fossil found in the Fray Bentos Formation of the department of Río Negro in Uruguay, and the type species is U. beaulieui. The related genera Xenastrapotherium and Granastrapotherium, which make up Uruguaytheriinae with Uruguaytherium, are also from South America, although them colonizated the equatorial zone. The holotype specimen of Uruguaytherium is a partial mandible (the left mandibular ramus), with a preserved third molar, or M3.

<i>Huilatherium</i> Extinct genus of mammals

Huilatherium is an extinct genus of leontiniid, a group of hoofed mammals belonging to the order Notoungulata, that comprises other South American ungulate families that evolved in parallel with some mammals of the Northern hemisphere. The leontiinids were a family of herbivorous species comprising medium to large browsers, with relatively short skulls and robust limbs, somewhat similar to their relatives, the best known toxodontids.

<i>Hilarcotherium</i>

Hilarcotherium is an extinct genus of astrapotheriid mammals that lived in South America during the Middle Miocene (Laventan). The type species is H. castanedaii, found in sediments of the La Victoria Formation, part of the Honda Group in the department of Tolima in Colombia. In 2018, Carrillo et al. described a partial skull and mandible of a second species H. miyou from the Castilletes Formation in the Cocinetas Basin of northern Colombia, and estimated the body weight of the animal at 6,465 kilograms (14,253 lb).

<i>Astraponotus</i> Extinct genus of mammals

Astraponotus is an extinct genus of astrapotheriids. It lived during the Middle-Late Eocene and its fossil remains have been found in the Sarmiento Formation of Argentina, South America.

Maddenia is an extinct genus of astrapothere, meridiungulate herbivore mammals characterised by its large tusks and the development of proboscis, endemic of South America. This genus was discovered in an outcrop near to the Lake Colhué Huapi in the place La Cantera, in the Chubut Province, in Argentina, in sediments corresponding to the Sarmiento Formation, that dates of the Late Oligocene.

<i>Miocochilius</i> Extinct genus of notoungulates

Miocochilius is an extinct genus of small notoungulate mammals (typotheres) native to South America. The genus lived during the Middle Miocene epoch. The genus contains two described species, the type species M. anomopodus described in 1953 by Ruben Arthur Stirton and M. federicoi, described and included in the genus by Darin A. Croft.

The Biblián Formation is an Early Miocene geologic formation of the Cuenca Basin in Ecuador. Fossils of Xenastrapotherium aequatorialis have been found in the formation.

Megadolodus is an extinct genus of proterotheriid litopterns.

Tetragonostylops is an extinct genus of mammal, related to Astrapotheria. It lived during the Late Paleocene, and its fossils were discovered in South America.

Huilabradys is an extinct genus of ground sloths of the family Nothrotheriidae that lived in what is now Colombia. Huilabradys was discovered in the strata of the La Tatacoa desert in the Huila department, in the Villavieja Formation, and is part of the so-called La Venta fauna, a fossiliferous location from the mid-Miocene period that has provided a notable paleontological contribution on the Miocene faunas of northern South America. The remains discovered are basically fragments of the jaws and teeth, allowed the identification of this species, whose only species is Huilabradys magdaleniensis, and was classified as a member of the nothrotheriid subfamily Nothrotheriinae, which comprises small to medium-sized species of ground sloths.

Nanoastegotherium is an extinct genus of cingulate, belonging to the family Dasypodidae, which includes the modern nine-banded armadillos. The name of the genus means "small Astegotherium", referring to its small size, smaller than the modern southern long-nosed armadillo, and to its affinities with Astegotherium, with which it forms the tribe Astegotheriini, within the family Dasypodidae. Its type species is Nanoastegotherium prostatum, whose species translates to "earlier" due to its age compared to Astegotherium.

<i>Magdalenabradys</i> Extinct genus of ground sloths

Magdalenabradys is an extinct genus of mylodontid ground sloths that lived during the Middle Miocene and Early Pliocene of what is now Colombia and Venezuela. Fossils have been found in the Villavieja Formation of the Honda Group in Colombia, and the Codore and Urumaco Formations of Venezuela.

References

  1. Mauricio Pardo Jaramillo. Reporte de un nuevo ejemplar de Granastrapotherium snorki en el Valle Superior del Magdalena, Desierto de la Tatacoa, Huila. Colombia Archived 2014-02-02 at the Wayback Machine .. Rev. Acad. Colomb. Cienc. 34 (131): 253-256, 2010. ISSN 0370-3908.
  2. Antoine, R., Salas-Gismondi, P., Baby, M., Benammi, S., Brusset, D., de Franceschi, N., Espurt, C., Goillot, F., Pujos, J., Tejada and M. Urbina. The Middle Miocene (Laventan) Fitzcarrald fauna, Amazonian Peru. 2007. Proceedings of the European Meeting on the Palaeontology and Stratigraphy of Latin America, Cuadernos del Museo Geominero, nº 8. Instituto Geológico y Minero de España, Madrid, 2007.
  3. Johnson, Steven C. and Madden, Richard H. 1997. Uruguaytheriinae Astrapotheres of Tropical South America. Chapter 22 in "Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia". Edited by Richard F. Kay, Richard H. Madden, Richard L. Cifelli, and John J. Flynn. Smithsonian Institution Press. Washington and London.
  4. Kramarz, Alejandro G.; Bond, Mariano (2008). "Revision of Parastrapotherium (Mammalia, Astrapotheria) and other Deseadan astrapotheres of Patagonia". Ameghiniana. 45 (3). Retrieved September 18, 2013.
  5. Johnson, Steven C. & Madden, Richard H. 1997. Uruguaytheriinae Astrapotheres of Tropical South America. Chapter 22 in “Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia”. Edited by Richard F. Kay, Richard H. Madden, Richard L. Cifelli, and John J. Flynn. Smithsonian Institution Press. Washington and London.
  6. M. C. Vallejo-Pareja; J. D. Carrillo; J. W. Moreno-Bernal; M. Pardo-Jaramillo; D. F. Rodriguez-Gonzalez; J. Muñoz-Duran (2015). "Hilarcotherium castanedaii, gen. et sp. nov., a new Miocene astrapothere (Mammalia, Astrapotheriidae) from the Upper Magdalena Valley, Colombia" (PDF). Journal of Vertebrate Paleontology. 35 (2): e903960. Bibcode:2015JVPal..35E3960V. doi:10.1080/02724634.2014.903960. S2CID   130728894.