Description
Its body was slender, with a very short tail and strong forelegs, although much shorter than the hind legs, equipped with long metatarsals.
The skull had a complete dentition, but the second and third incisors, as well as the canines and the first lower premolars, were vestigial. The first upper incisors were very large, and showed a considerable development. The molars had two internal folds separated by a further deep fold, like the Interatheriidae ; dental cementum was present, but to a lesser extent than in Interatherium . The lower teeth were flat on the outside and bilobed on the inside.
The tibia and fibula were not fused, and the forearm and lower leg bones were respectively shorter than the humerus and femur, but the hind legs were much longer than the forelegs. The second and third fingers of the hand were of similar length, and the fourth was smaller ; the fifth finger was small, even smaller than in its relative Pachyrukhos . The hand seems more specialized than in Pachyrukhos, and shares similarities with that of Paedotherium . The metatarsals were elongated, especially the third.
Classification
Propachyrucos was first described in 1897 by Florentino Ameghino, based on a right mandible fragment found in Oligocene terrains of the Sarmiento Formation in Argentina, initially believed by Ameghino to have been Cretaceous. The type species is Propachyrucos smithwoodwardi, but other species have been attributed to the genus, such as P. aequilatus, P. robustus, and P. ameghinorum, the latter species, described in 1945 by George Gaylord Simpson from fossils found in the Chubut Province, is known from a remarkably complete skeleton, permitting to reconstruct the animal appearance with high precision. However, recent researches tend to indicate that all the other species of Propachyrucos are in fact synonymous with the type species. [1]
Propachyrucos was a hegetothere, a group of notoungulates that evolved in isolation on the South American continent, giving rise to forms resembling hares and rabbits. Propachyrucos was a member of Pachyrukhinae, a subfamily of hegetotheres particularly similar to modern Lagomorphs.
Pyrotherium is an extinct genus of South American ungulate, of the order Pyrotheria, that lived in what is now Argentina and Bolivia, during the Late Oligocene. It was named Pyrotherium because the first specimens were excavated from an ancient volcanic ash deposit. Fossils of the genus have been found in the Deseado and Sarmiento Formations of Argentina and the Salla Formation of Bolivia.
Cramauchenia is an extinct genus of litoptern South American ungulate. Cramauchenia was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus Macrauchenia. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian, as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan. In 1981 Soria made C. insolita a junior synonym of C. normalis. A specimen of C. normalis was described in 2010 from Cabeza Blanca in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA.
Interatherium is an extinct genus of interatheriid notoungulate from the Early to Middle Miocene (Colhuehuapian-Mayoan). Fossils have been found in the Santa Cruz, Collón Curá and Sarmiento Formations in Argentina.
The Malargüe Group is a group of geologic formations of the Neuquén Basin of the Mendoza, Neuquén, Río Negro and La Pampa Provinces in northern Patagonia, Argentina. The formations of the Malargüe Group range in age between the middle Campanian to Deseadan, an Oligocene age of the SALMA classification, straddling the Cretaceous–Paleogene boundary, about 79 million to 30 million years in age. The group overlies the older Neuquén Group, separated by an unconformity dated to 79 Ma. The rocks of the Malargüe Group comprise both marine and continental deposits which are over 400 m (1312 ft) thick in total.
Prosotherium is an extinct genus of hegetotheriid notoungulate. It lived during the Late Oligocene, and its fossilized remains were found in South America.
Tremacyllus is an extinct genus of hegetotheriids. It lived from the Late Miocene to the Late Pleistocene and its fossilized remains were discovered in South America.
Proadinotherium is an extinct genus of toxodontid. It lived between the Late Oligocene and the Early Miocene in what is now South America.
Ethegotherium is an extinct genus of Notoungulates, belonging to the suborder Typotheria. It lived from the Lower to the Middle Miocene, and its fossilized remains were discovered in South America. It might be a synonym of the genus Prohegetotherium.
Eohyrax is an extinct genus of Notoungulate, belonging to the suborder Typotheria. It lived during the Middle Eocene, and its remains were discovered in South America.
Argyrohippus is an extinct genus of notoungulate, belonging to the family Notohippidae. It lived from the Late Oligocene to the Early Miocene, and its fossilized remains were found in South America.
Periphragnis is an extinct genus of isotemnid notoungulates that lived from the Middle Eocene to the Early Oligocene in what is now Argentina and Chile.
Asmodeus is an extinct genus of mammal, belonging to the order Notoungulata. It lived during the Late Oligocene, in what is today South America.
Eomorphippus is an extinct genus of notohippid notoungulate that lived from the Late Eocene to the Early Oligocene in what is today South America.
Eurygenium is an extinct genus of notoungulate belonging to the family Notohippidae. It lived during the Late Oligocene in what is today South America.
Morphippus is an extinct genus of notohippid notoungulate that lived during the Middle to Late Oligocene in what is now South America.
Ancylocoelus is an extinct genus of mammal, belonging to the order Notoungulata. It lived during the Late Oligocene, in what is today Argentina, in South America.
Argyrohyrax is an extinct genus of interatheriid notoungulate that lived during the Late Oligocene, of what is now Argentina and Bolivia.
Patriarchus is an extinct genus of interatheriid notoungulates that lived during the Early Miocene in what is now Argentina. Fossils of this genus have been found in the Santa Cruz Formation of Argentina.
Notopithecus is an extinct genus of Notoungulate, belonging to the suborder Typotheria. It lived from the Middle to the Late Eocene and its fossilized remains were discovered in South America.
Hegetotheriopsis is an extinct genus of hegetotheriid notoungulate. It lived from the Late Oligocene to the Early Miocene, and its fossilized remains are found in Argentina.
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