Moqueguahippus | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Notoungulata |
Family: | † Notohippidae |
Genus: | † Moqueguahippus Shockey et al. 2006 |
Species: | †M. glycisma |
Binomial name | |
†Moqueguahippus glycisma Shockey et al., 2006 | |
Moqueguahippus is an extinct genus of notohippid notoungulates that lived during the Late Oligocene of what is now Peru. Fossils of this genus have been found in the Moquegua Formation of Peru, which it was named after. [1] [2]
The genus name, Moqueguahippus, refers to the Moquegua Formation where it was discovered, and -hippus meaning "horse", an epithet used to describe most notohippids. The specific name, glycisma, is derived from the Greek word Glykisma, meaning "cake", in reference to the type locality, Pan de Azucar, meaning "Sugarloaf Mountain". [1]
Moqueguahippus is a relatively large notohippid. Its skull was rather massive and had a short muzzle. The dentition was complete and continuous, lacking a diastema, and the premolars and molars were high-crowned (hypsodont), like in its relative Rhynchippus . The premolars had a well developed cingulum, while the molars were elongated. The shape of the premolars and molars was reminiscent to those of Equidae. Moqueguahippus, like its relatives, may have had a robust body with slender limbs. Its most peculiar distinctive characteristics were the cup-shaped internal cingula and the lack of cingula on the premolars, and a thick layer of cementum. [1] [3]
Moqueguahippus was first described in 2006, based on fossilized remains found in the Moquegua Formation in Peru, in deposits dating to the Late Oligocene. Moqueguahippus was placed within the family Notohippidae, a family of toxodont notoungulates with horse-like teeth, with very high-crowned molars. More recent research, however, suggests that this group is paraphyletic, containing increasingly derived forms, some of them close to the ancestors of "true" Toxodontidae. In 2021, it was found to be a derived notohippid, being the sister taxon of a clade including the Miocene genera Argyrohippus and Notohippus . [4]
The following position of the Toxodontia is based on Martínez et al. 2021, showing the position of Moqueguahippus. [4]
Toxodontia |
| "Isotemnidae" Paranotohippids "Isotemnidae" "Notohippidae" | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Toxodontia is a suborder of the meridiungulate order Notoungulata. Most of the members of the five included families, including the largest notoungulates, share several dental, auditory and tarsal specializations. The group is named after Toxodon, the first example of the group to be discovered by science.
Pyrotheria is an order of extinct meridiungulate mammals. These mastodon-like ungulates include the genera Baguatherium, Carolozittelia, Colombitherium, Griphodon, Propyrotherium, Proticia, and Pyrotherium.
Pyrotheriidae is the only family in the order Pyrotheria, provided one does not include the Paleocene genus, Carodnia. These extinct, mastodon-like ungulates include the genera Baguatherium, Carolozittelia, Griphodon, Propyrotherium, and Pyrotherium. Fossils of the family have been found in Argentina, Brazil, Bolivia and Peru.
Peltephilus, the horned armadillo, is an extinct genus of armadillo xenarthran mammals that first inhabited Argentina during the Oligocene epoch, and became extinct in the Miocene epoch. Notably, the scutes on its head were so developed that they formed horns. Aside from the horned gophers of North America, it is the only known fossorial horned mammal. P. ferox had skull about 11.7 centimetres (4.6 in), and estimated body mass is around 11.07 kilograms (24.4 lb).
Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.
Rhynchippus is an extinct genus of notoungulate mammals from the Late Oligocene of South America. The genus was first described by Florentino Ameghino in 1897 and the type species is R. equinus, with lectotype MACN A 52–31. Fossils of Rhynchippus have been found in the Agua de la Piedra and Sarmiento Formations of Argentina, the Salla and Petaca Formations of Bolivia, the Tremembé Formation of Brazil, and the Moquegua Formation of Peru.
Cramauchenia is an extinct genus of litoptern South American ungulate. Cramauchenia was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus Macrauchenia. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian, as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan. In 1981 Soria made C. insolita a junior synonym of C. normalis. A specimen of C. normalis was described in 2010 from Cabeza Blanca in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA.
Notohippidae is a paraphyletic extinct family of notoungulate mammals from South America. Notohippids are known from the Eocene and Oligocene epochs.
Mesotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pleistocene of South America. Mesotheriids were small to medium-sized herbivorous mammals adapted for digging.
The Deseadan age is a period of geologic time within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Tinguirirican and precedes the Colhuehuapian age.
The Santa Rosa local fauna consists of the animals found in the Paleogene fossil site of Santa Rosa in eastern Peru. The age of the Santa Rosa fauna is difficult to determine, but may be Eocene (Mustersan) or Oligocene (Deseadan).
The Agua de la Piedra Formation is a Late Oligocene geologic formation of the Malargüe Group that crops out in the southernmost Precordillera and northernmost Neuquén Basin in southern Mendoza Province, Argentina.
Trachytherus is an extinct genus of mesotheriid notoungulate that lived from the Late Oligocene to the Early Miocene in what is now South America.
Eurygenium is an extinct genus of notoungulate belonging to the family Notohippidae. It lived during the Late Oligocene in what is today South America.
Morphippus is an extinct genus of notohippid notoungulate that lived during the Middle to Late Oligocene in what is now South America.
Anayatherium is an extinct genus of notoungulate belonging to the family Leontiniidae. It lived during the Late Oligocene, and its fossilized remains were discovered in South America.
Pascualihippus is an extinct genus of notoungulate belonging to the family Notohippidae. It lived during the Late Oligocene, in what is now Bolivia.
The Sarmiento Formation, in older literature described as the Casamayor Formation, is a geological formation in Chubut Province, Argentina, in central Patagonia, which spans around 30 million years from the mid-Eocene to the early Miocene. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Sarmiento Formation, including a variety of birds, crocodilians, turtles and snakes, also includes many mammals such as South American native ungulates as well as armadillos, and caviomorph rodents.
Teushentherium is an extinct genus of notohippid notoungulate from the Oligocene of Argentina. Fossils have been found in the Sarmiento Formation of Argentina.
Rosendo is an extinct genus of notohippid notoungulates that lived during the Early Oligocene in what is now Argentina and Chile. Fossils of this genus have been found in the Sarmiento Formation and the Abanico Formations of Argentina and Chile.