Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined during the Late Neogene, with only the large toxodontids persisting until the end of the Pleistocene, perishing as part of the Quaternary extinction event among with most other large mammals in the Americas. Collagen analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls, including rhinoceroses, tapirs and equines. However their relationships to other South American ungulates are uncertain. Several groups of notoungulates separately evolved ever-growing cheek teeth.
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups conventionally ranked as orders—Astrapotheria, Litopterna, Notoungulata, Pyrotheria, and Xenungulata—as well as the primitive "condylarth" groups Didolodontidae and Kollpaniinae. It has been proposed that some or all of the members of this group form a clade, named Meridiungulata, though the relationships of South American ungulates remain largely unresolved. The two largest groups of South American ungulates, the notoungulates and the litopterns, were the only groups to persist beyond the mid Miocene. Only a few of the largest species of notoungulates and litopterns survived until the end-Pleistocene extinctions.
Pyrotheria is an order of extinct meridiungulate mammals. These mastodon-like ungulates include the genera Baguatherium, Carolozittelia, Colombitherium, Griphodon, Propyrotherium, Proticia, and Pyrotherium.
Trigonostylops is an extinct genus of South American meridiungulatan ungulate, from the Late Paleocene to Late Eocene of South America and Antarctica. It is the only member of the family Trigonostylopidae.
Victorlemoinea is an extinct litoptern genus of the family Sparnotheriodontidae, that lived from the Early to Middle Eocene. Fossils of Victorlemoinea have been found in the Las Flores, Sarmiento and Koluel Kaike Formations of Argentina, the Itaboraí Formation of Brazil and La Meseta Formation, Antarctica.
Carodnia is an extinct genus of South American ungulate known from the Early Eocene of Brazil, Argentina, and Peru. Carodnia is placed in the order Xenungulata together with Etayoa and Notoetayoa.
Astrapotheria is an extinct order of South American and Antarctic hoofed mammals that existed from the late Paleocene to the Middle Miocene, 59 to 11.8 million years ago. Astrapotheres were large, rhinoceros-like animals and have been called one of the most bizarre orders of mammals with an enigmatic evolutionary history.
The Itaboraian age is a period within the Early Eocene geologic time epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age.
The Riochican age is a period of geologic time within the Paleocene and Eocene epochs of the Paleogene, used more specifically within the South American land mammal ages (SALMA). It follows the Peligran and precedes the Itaboraian age.
Etayoa is an ungulate of the family Carodniidae in the order Xenungulata that lived during the Early Eocene in northern South America.
Cocatherium is an extinct genus of marsupial mammals of uncertain family placement, from the earliest Paleocene of South America, predating the Tiupampan South American land mammal age. The genus was described based on a fossil molar that was found in the Danian part of the Cretaceous-Paleogene Lefipán Formation in the Cañadón Asfalto Basin in north-central Patagonia, Argentina. The type species of the genus is C. lefipanum.
The Itaboraí Formation is a highly fossiliferous geologic formation and Lagerstätte of the Itaboraí Basin in Rio de Janeiro, southeastern Brazil. The formation reaching a thickness of 100 metres (330 ft) is the defining unit for the Itaboraian South American land mammal age (SALMA), dating to the Early Eocene, approximately 53 to 50 Ma.
The Río Loro Formation is a geological formation of the Sierras Pampeanas in Tucumán Province Argentina whose strata date back to the Late Paleocene of the Paleogene, or Riochican in the SALMA classification.
Notiolofos is an extinct genus of sparnotheriodontid ungulate from the order Litopterna. The animal lived during the Eocene, in modern-day Antarctica. The genus contains two species, N. arquinotiensis, the type species, and N. regueroi.
Ernestokokenia is an extinct genus of mammal, belonging to the Didolodontidae. It lived during the Early Eocene and the Middle Eocene, and its fossils were discovered in South America.
Lamegoia is an extinct genus of mammals, belonging to the family Didolodontidae. It contains a single species, Lamegoia conodonta, which lived during the Late Paleocene in what is now South America.
Notoetayoa is an extinct genus of mammal, from the order Xenungulata. It contains a single species, Notoetayoa gargantuai, which lived during the Middle Paleocene. Its fossilized remains were discovered in South America.
Rodcania is an extinct genus of mammal, belonging to the order Xenungulata. It contains a single species, Rodcania kakan, which lived during the Paleocene. Its remains were found in South America. The genus name is an anagram of Carodnia.
Sparnotheriodontidae is an enigmatic extinct family of litopterns known primarily from teeth. Sparnotheriodontids are one of two South American native ungulate clades known to have reached Antarctica, the other being astrapotheres.
Sparnotheriodon is an extinct genus of sparnotheriodontid litoptern that lived during the Middle Eocene of what is now Argentina, leaving fossils in the Sarmiento Formation.
