| Scaglia kraglievichorum | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | † Astrapotheria |
| Family: | † Astrapotheriidae |
| Subfamily: | † Astrapotheriinae |
| Genus: | † Scaglia Simpson, 1957 |
| Species: | †S. kraglievichorum |
| Binomial name | |
| †Scaglia kraglievichorum Simpson, 1957 | |
Scaglia is an extinct genus of South American astrapotherid land mammal that lived during the Eocene (Casamayoran to Divisaderan in the SALMA classification). [1]
The genus was named after Argentinian naturalist Galileo Juan Scaglia, [2] and the type species after Argentinian palaeontologist Lucas Kraglievich.
Its type specimen, recovered from the Sarmiento Formation of Argentina, is MMCNT-MdP 207. [3] Like Albertogaudrya , Scaglia was the size of a sheep or a small tapir, hence among the larger mammals in South America at that time. [4]
Cladogram according to Bond et al., 2011, standing out the phylogenetic position of Scaglia: [5]
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups conventionally ranked as orders—Astrapotheria, Litopterna, Notoungulata, Pyrotheria, and Xenungulata—as well as the primitive "condylarth" groups Didolodontidae and Kollpaniinae. It has been proposed that some or all of the members of this group form a clade, named Meridiungulata, though the relationships of South American ungulates remain largely unresolved. The two largest groups of South American ungulates, the notoungulates and the litopterns, were the only groups to persist beyond the mid Miocene. Only a few of the largest species of notoungulates and litopterns survived until the end-Pleistocene extinction event around 12,000 years ago where they became extinct with most other large mammals in the Americas, shortly after the first arrival of humans into the region.
Xenungulata is an order of extinct and primitive South American hoofed mammals that lived from the Late Paleocene to Early Eocene. Fossils of the order are known from deposits in Brazil, Argentina, Peru, and Colombia. The best known member of this enigmatic order is the genus Carodnia, a tapir-like and -sized animal with a gait similar to living African elephants.
Trigonostylops is an extinct genus of South American meridiungulatan ungulate, from the Late Paleocene to Late Eocene of South America and Antarctica. It is the only member of the family Trigonostylopidae.
Cramauchenia is an extinct genus of litoptern South American ungulate. Cramauchenia was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus Macrauchenia. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian, as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan. In 1981 Soria made C. insolita a junior synonym of C. normalis. A specimen of C. normalis was described in 2010 from Cabeza Blanca in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA.
Mesembriornis is a genus of intermediate-sized phorusrhacids that grew up to 1.5 metres (4.9 ft) in height. They represent a well-distinct lineage of terror birds, differing from the massive large groups and the smaller Psilopterinae. In general proportions, they most resembled the Patagornithinae which flourished somewhat earlier, mainly to the south of the range of Mesembriornis. Fossils of the terror bird have been found in Montehermosan deposits of the Monte Hermoso Formation, as well as the Andalgala Formation and Chapadmalal Formation in Argentina.
Astrapotheria is an extinct order of South American and Antarctic hoofed mammals that existed from the late Paleocene to the Middle Miocene, 59 to 11.8 million years ago. Astrapotheres were large, rhinoceros-like animals and have been called one of the most bizarre orders of mammals with an enigmatic evolutionary history.
Remingtonocetus is an extinct genus of early cetacean freshwater aquatic mammals of the family Remingtonocetidae endemic to the coastline of the ancient Tethys Ocean during the Eocene. It was named after naturalist Remington Kellogg.
Patagornis is a genus of extinct flightless predatory birds of the family Phorusrhacidae. Known as "terror birds", these lived in what is now Argentina during the Early and Middle Miocene; the Santa Cruz Formation in Patagonia contains numerous specimens. Patagornis was an agile, medium sized Patagornithine and was likely a pursuit predator.
The Casamayoran age is a period of geologic time within the Early Eocene epoch of the Paleogene, used more specifically within the South American land mammal age (SALMA) classification. It follows the Itaboraian and precedes the Mustersan age.
Astrapotheriidae is an extinct family of herbivorous South American land mammals that lived from the Late Eocene to the Middle Miocene 37.71 to 15.98 million years ago. The most derived of the astrapotherians, they were also the largest and most specialized mammals in the Tertiary of South America. There are two sister taxa: Eoastrapostylopidae and Trigonostylopidae.
Antarctodon is an extinct genus of mammals from the Early Eocene. It is a basal astrapotherian which lived in what is now Seymour Island, Antarctic Peninsula, at that moment still connected to South America where most of the astrapotherians were found. The holotype and only specimen MLP 08-XI-30-1, an isolated right p4 or m1, was found in the Telm 5 Member of the La Meseta Formation in West Antarctica. It was first named by Mariano Bond, Alejandro Kramarz, Ross D. E. MacPhee and Marcelo Reguero in 2011 and the type species is Antarctodon sobrali.
Granastrapotherium is an extinct genus of ungulate mammals, described from remains found in rocks of the Honda Group in the Tatacoa Desert, in the Colombian departments of Huila and Tolima, at the Miocene fossil site La Venta. The only species formally recognized is Granastrapotherium snorki. Remains found in Bolivia and Peru, seem to belong to Granastrapotherium or a very similar animal.
Albertogaudrya is an extinct genus of astrapotherian mammal that lived in present-day Patagonia and Salta, Argentina during the Eocene 48.6 to 37.2 million years ago. Fossils of Albertogaudrya have been found in the Lumbrera and Sarmiento Formations. It is named after French palaeontologist Albert Gaudry.
Hilarcotherium is an extinct genus of astrapotheriid mammals that lived in South America during the Middle Miocene (Laventan). The type species is H. castanedaii, found in sediments of the La Victoria Formation, part of the Honda Group in the department of Tolima in Colombia. In 2018, Carrillo et al. described a partial skull and mandible of a second species H. miyou from the Castilletes Formation in the Cocinetas Basin of northern Colombia, and estimated the body weight of the animal at 6,465 kilograms (14,253 lb).
Astraponotus is an extinct genus of astrapotheriids. It lived during the Middle-Late Eocene and its fossil remains have been found in the Sarmiento Formation of Argentina, South America.
Maddenia is an extinct genus of astrapothere, meridiungulate herbivore mammals characterised by its large tusks and the development of proboscis, endemic of South America. This genus was discovered in an outcrop near to the Lake Colhué Huapi in the place La Cantera, in the Chubut Province, in Argentina, in sediments corresponding to the Sarmiento Formation, that dates of the Late Oligocene.
Dryornis, also called the Argentinian vulture, is an extinct genus of cathartid, known from Argentina. The genus contains two species, D. pampeanus and D. hatcheri.
Sparnotheriodontidae is an enigmatic extinct family of litopterns known primarily from teeth. Sparnotheriodontids are one of two South American native ungulate clades known to have reached Antarctica, the other being astrapotheres.
Tetragonostylops is an extinct genus of mammal, related to Astrapotheria. It lived during the Late Paleocene, and its fossils were discovered in South America.
The Sarmiento Formation, in older literature described as the Casamayor Formation, is a geological formation in Chubut Province, Argentina, in central Patagonia, which spans around 30 million years from the mid-Eocene to the early Miocene. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Sarmiento Formation, including a variety of birds, crocodilians, turtles and snakes, also includes many mammals such as South American native ungulates as well as armadillos, and caviomorph rodents.
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