Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines, rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do.
Ungulates are members of the diverse clade Euungulata which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Artiodactyls are mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine as perissodactyls do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.
Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Artiodactyla, along with species including whales, pigs, deer, cattle, and antelopes.
Notoungulata is an extinct order of ungulates that inhabited South America from the early Paleocene to the end of the Pleistocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined from the late Neogene onwards, with only the large toxodontids persisting until the end of the Pleistocene, perishing as part of the Late Pleistocene extinctions among with most other large mammals in the Americas. Collagen sequence analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls, including rhinoceroses, tapirs and equines as part of the clade Panperissodactyla. However their relationships to other South American ungulates are uncertain. Several groups of notoungulates separately evolved ever-growing cheek teeth.
Dinocerata or Uintatheria, also known as uintatheres, is an extinct order of large herbivorous hoofed mammals with horns and protuberant canine teeth, known from the Paleocene and Eocene of Asia and North America. With body masses ranging up to 4,500 kilograms (9,900 lb) they represent some of the earliest known large mammals.
Coryphodon is an extinct genus of pantodonts of the family Coryphodontidae.
Didolodus is an extinct genus of mammals from Middle Eocene Argentina. It is an ungulate mammal of uncertain affinities, possibly related to Litopterna, though this is uncertain due to the lack of reliable post-cranial remains, and for now remains Meridiungulata incertae sedis. Its remains were found in the Sarmiento Formation of Patagonia.
Trigonostylops is an extinct genus of South American meridiungulatan ungulate, from the Late Paleocene to Late Eocene of South America and Antarctica. It is the only member of the family Trigonostylopidae.
Pyrotherium is an extinct genus of South American ungulate, of the order Pyrotheria, that lived in what is now Argentina and Bolivia, during the Late Oligocene. It was named Pyrotherium because the first specimens were excavated from an ancient volcanic ash deposit. Fossils of the genus have been found in the Deseado and Sarmiento Formations of Argentina and the Salla Formation of Bolivia.
Elomeryx is an extinct genus of artiodactyl ungulate, and is among the earliest known anthracotheres. The genus was extremely widespread, first being found in Asia in the middle Eocene, in Europe during the latest Eocene, and having spread to North America by the early Oligocene. The closest living relatives of the Elomeryx are bovids, suids, and cetaceans.
Amynodontidae is a family of extinct perissodactyls related to true rhinoceroses. They are commonly portrayed as semiaquatic hippo-like rhinos but this description only fits members of the Metamynodontini; other groups of amynodonts like the cadurcodontines had more typical ungulate proportions and convergently evolved a tapir-like proboscis.
Notiolofos is an extinct genus of sparnotheriodontid ungulate from the order Litopterna. The animal lived during the Eocene, in modern-day Antarctica. The genus contains two species, N. arquinotiensis, the type species, and N. regueroi.
Ernestokokenia is an extinct genus of mammal, belonging to the Didolodontidae. It lived during the Early Eocene and the Middle Eocene, and its fossils were discovered in South America.
Saltaodus is an extinct genus of mammals, belonging to the family Didolodontidae. It lived during the Late Eocene, in what is now South America.
Asmithwoodwardia is an extinct genus of mammals, from the order Litopterna. It lived during the Late Paleocene and the Early Eocene, and its fossilized remains were found in South America.
Eohyrax is an extinct genus of Notoungulate, belonging to the suborder Typotheria. It lived during the Middle Eocene, and its remains were discovered in South America.
Rhyphodon is an extinct genus of notoungulate, who lived from the Middle to the Late Eocene in what is today South America.
Colbertia is an extinct genus of oldfieldthomasiid notoungulate. It lived from the Early to the Middle Eocene, and its fossilized remains were discovered in Argentina and Brazil.
Vassallia is an extinct genus of cingulate belonging to the family Pampatheriidae. It lived between the Middle Oligocene and the Early Pliocene in what is now South America.
Size comparison of Notostylops murinus with a human.
Life reconstruction.
