Toxodon is an extinct genus of South American mammals from the Late Miocene to early Holocene epochs. It is a member of Notoungulata, one of several now extinct orders of South American native ungulates distinct from living perissodactyls and artiodactyls. It was among the largest and last members of its order, and was probably the most common large hoofed mammal in South America of its time.
Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined from the late Neogene onwards, with only the large toxodontids persisting until the end of the Pleistocene, perishing as part of the Quaternary extinction event among with most other large mammals in the Americas. Collagen analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls, including rhinoceroses, tapirs and equines as part of the clade Panperissodactyla. However their relationships to other South American ungulates are uncertain. Several groups of notoungulates separately evolved ever-growing cheek teeth.
Toxodontia is a suborder of the meridiungulate order Notoungulata. Most of the members of the five included families, including the largest notoungulates, share several dental, auditory and tarsal specializations. The group is named after Toxodon, the first example of the group to be discovered by science.
Macrauchenia was a large, long-necked and long-limbed, three-toed native South American ungulate in the order Litopterna. The genus gives its name to its family, the Macraucheniidae or "robust litopterns". Like other litopterns, it is most closely related to the odd-toed ungulates (Perissodactyla), from which litopterns diverged approximately 66 million years ago. The oldest fossils in the genus date to the late Miocene, around seven million years ago, and M. patachonica disappears from the fossil record during the late Pleistocene, around 20,000-10,000 years ago. M. patachonica is one of the last and best known member of the family and is known primarily from the Luján Formation in Argentina, but is known from localities across southern South America. Another genus of macraucheniid Xenorhinotherium was present in northeast Brazil and Venezuela during the Late Pleistocene. The type specimen was discovered by Charles Darwin during the voyage of the Beagle. In life, Macrauchenia may have resembled a humpless camel, though the two taxa are not closely related. It fed on plants in a variety of environments across what is now South America. Among the species described, M. patachonica and M. ullomensis are considered valid; M. boliviensis is considered a nomen dubium; and M. antiqua has been moved to the genus Promacrauchenia.
Glyptotherium is a genus of glyptodont in the family Chlamyphoridae that lived from the Early Pliocene, about 3.6 million years ago, to the Late Pleistocene, around 15,000 years ago. It had a wide distribution, living in the United States, Mexico, Guatemala, Costa Rica, Honduras, El Salvador, Panama, Venezuela, and Brazil. The genus was first described in 1903 by American paleontologist Henry Fairfield Osborn with the type species being, G. texanum, based on fossils that had been found in the Pliocene Blancan Beds in Llano Estacado, Texas, USA. Glyptotherium fossils have since been unearthed from many more fossil sites, from Florida to Colombia. Another species, G. cylindricum, was named in 1912 by fossil hunter Barnum Brown on the basis of a partial skeleton that had been unearthed from the Pleistocene deposits in Jalisco, Mexico. The two species differ in several aspects, including age, with G. texanum being from the older Early Pliocene to Early Pleistocene strata, whereas G. cylindricum is exclusive to the Late Pleistocene.
Toxodontidae is an extinct family of notoungulate mammals, known from the Oligocene to the Holocene of South America, with one genus, Mixotoxodon, also known from the Pleistocene of Central America and southern North America. Member of the family were medium to large-sized, ranging from around 350–400 kilograms (770–880 lb) in Nesodon to 1,000–1,200 kilograms (2,200–2,600 lb) in Toxodon, and had medium to high-crowned dentition, which in derived members of the group evolved into ever-growing cheek teeth. Isotopic analyses have led to the conclusion that Pleistocene members of the family were flexible mixed feeders.
Eremotherium is an extinct genus of giant ground sloth in the family Megatheriidae. Eremotherium lived in the southern North America, Central America, and northern South America from the Pliocene, around 5.3 million years ago, to the end of the Late Pleistocene, around 10,000 years ago. Eremotherium was widespread in tropical and subtropical lowlands and lived there in partly open and closed landscapes, while its close relative Megatherium lived in more temperate climes of South America. Both genera reached the size of today's elephants and were among the largest mammals in the Americas. Characteristic of Eremotherium was its robust physique with comparatively long limbs and front and hind feet especially for later representatives- three fingers. However, the skull is relatively gracile, the teeth are uniform and high-crowned. Like today's sloths, Eremotherium was purely herbivorous and was probably a mixed feeder that dined on leaves and grasses. Eremotherium was a generalist that could adapt its diet to the respective local and climatic conditions of many regions. Finds of Eremotherium are common and widespread, with fossils being found as far north as South Carolina in the United States and as far south as Rio Grande Do Sul, and many complete skeletons have been unearthed.
Mixotoxodon is an extinct genus of notoungulate of the family Toxodontidae inhabiting South America, Central America and parts of southern North America during the Pleistocene epoch, from 1,800,000—12,000 years ago.
Nesodon is a genus of Miocene mammal belonging to the extinct order Notoungulata which inhabited southern South America during the Late Oligocene to Miocene living from 29.0 to 16.3 Ma and existed for approximately 12.7 million years. It had a relatively large size, weighing up to 554 kg (1221 lbs) and reaching 1.5 m in height.
Glyptodon is a genus of glyptodont, an extinct group of large, herbivorous armadillos, that lived from the Pliocene, around 3.2 million years ago, to the early Holocene, around 11,000 years ago, in Brazil, Uruguay, Paraguay, Bolivia, Peru, Argentina, and Colombia. It is one of, if not the, best known genus of glyptodont. Glyptodon has a long and storied past, being the first named extinct cingulate and the type genus of the subfamily Glyptodontinae. Fossils of Glyptodon have been recorded as early as 1814 from Pleistocene aged deposits from Uruguay, though many were incorrectly referred to the ground sloth Megatherium by early paleontologists.
Catonyx is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Pliocene and Pleistocene epochs. It lived from 2.5 Ma to about 10,000 years ago, existing for approximately 2.49 million years. The most recent date obtained is about 9600 B.P.
Notiomastodon is an extinct genus of gomphothere proboscidean, endemic to South America from the Pleistocene to the beginning of the Holocene. Notiomastodon specimens reached a size similar to that of the modern Asian elephant. Like other brevirostrine gomphotheres such as Cuvieronius and Stegomastodon, Notiomastodon had a shortened lower jaw and lacked lower tusks.
Arctotherium is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. Arctotherium migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama during the late Pliocene. The genus consists of one early giant form, A. angustidens, and several succeeding smaller species, which were within the size range of modern bears. Arctotherium was adapted to open and mixed habitat. They are genetically closer to the spectacled bear, than to Arctodus of North America, implying the two extinct forms evolved large size in a convergent manner.
Protopithecus is an extinct genus of large New World monkey that lived during the Pleistocene. Fossils have been found in the Toca da Boa Vista cave of Brazil, as well as other locales in the country. Fossils of another large, but less robust ateline monkey, Caipora, were also discovered in Toca da Boa Vista.
Hoffstetterius is an extinct genus of toxodontid notoungulate mammal, belonging to the subfamily Toxodontinae whose remains were discovered in the Middle to Late Miocene Mauri Formation in the La Paz Department in Bolivia. The only described species is the type Hoffstetterius imperator.
Valgipes is an extinct genus of scelidotheriid ground sloth, endemic to intertropical Brazil and Uruguay during the Late Pleistocene. Thought to have been a forest-dwelling browser, Valgipes is a monotypic genus with a complex and long taxonomic history, and is a close relative of Catonyx and Proscelidodon.
Cullinia is an extinct genus of litoptern, an order of South American native ungulates that included horse-like and camel-like animals such as Macrauchenia. It is only known from fragmentary remains. Cullinia levis is known from Chasicoan remains found in the Arroyo Chasicó Formation of Argentina, and remains from the Brazilian state of Acre and the Huayquerian Ituzaingó Formation have been assigned to Cullinia sp..
Falcontoxodon is an extinct genus of toxodontid notoungulate that lived from the late Pliocene to the Pleistocene in what is now Venezuela. Fossils of this genus have been found in the Chapadmalalan-Uquian Codore Formation, as well as in the more recent Ensenadan San Gregorio Formation.
Proadinotherium is an extinct genus of toxodontid. It lived between the Late Oligocene and the Early Miocene in what is now South America.
Gyrinodon is an extinct genus of toxodontid notoungulate that lived from the Late Miocene and Early Pliocene in what is now Brazil and Colombia.
