Ahytherium Temporal range: Late Pleistocene (Lujanian) | |
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Skull | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Pilosa |
Family: | † Megalonychidae |
Genus: | † Ahytherium Cartelle et al., 2008 |
Species: | †A. aureum |
Binomial name | |
†Ahytherium aureum Cartelle et al., 2008 | |
Ahytherium is an extinct genus of megalonychid sloth that lived during the Pleistocene of what is now Brazil. It contains a single species, A. aureum. [1] [2]
The almost-complete skeleton of Ahytherium alongside remains of another extinct sloth species, Australonyx , were discovered in Poço Azul, an underwater cave in Chapada Diamantina National Park in 2005. It was described by Castor Cartelle of Pontifícia Universidade Católica de Minas Gerais. The bones, which had a length of about 3 metres (9.8 ft) when put together, belong to an animal which presumably was still growing. [2]
This animal is known from well-preserved and nearly complete fossils, including a skeleton of an immature specimen, which in life must have been about three meters long, one meter tall and weighing perhaps half a ton. Like all ground sloths, Ahytherium was equipped with a robust body and legs equipped with powerful claws. Ahytherium was similar to other ground sloths such as Megalonyx but possessed some features that clearly distinguished it: the head, for example, was extremely short and tall, and possessed dorsally swollen frontal bones. The zygomatic arches were wide, particularly the frontal processes, while the lacrimal bones were narrow, blade-shaped and directed anterolaterally. The dentition included canine teeth that were thin, curved and oval in cross-section. The humerus was equipped with a poorly developed deltopectoral crest, while the femur possessed a large trochanter located distally. The shape of the caudal vertebrae indicates that the tail was dorsoventrally flattened.
Ahytherium is a genus of the extinct family Megalonychidae within the suborder Folivora and the superorder Xenarthra. The Megalonychidae represent a very diverse group. The closest relatives of the Megalonychidae are the Megatheriidae and the Nothrotheriidae. The former include the largest known representatives of the sloths, the latter consist of rather smaller members of the sloths. All three families together are relegated to the superfamily of Megatherioidea. Within the sloths, the Megalonychidae form a very old lineage, with the oldest known fossils dating to the Oligocene of Patagonia. [3] Characteristic features are found in the caniniform and incisiform design of the anterior tooth in each case, as well as in the molar-like (molariform) designed posterior teeth. The latter are characterized by two transverse ridges (bilophodont) on the occlusal surface, which points to a rather leaf-eating diet in the Megalonychidae. Unlike the Megatheriidae and the Nothrotheriidae, the hind foot is plantigrade in shape and not twisted, thus retaining its original shape. [4] The Megalonychidae were once widespread, with fossil remains found in both South America and Central America, as well as in North America as far north as the Arctic. [4] In a classic view based on skeletal anatomy comparisons-based view, the Megalonychidae originally included the sloths of the West Indies as well as the extant two-toed sloths of the genus Choloepus, [5] which are now considered to belong to the superfamily Mylodontoidea. [6] [7] Molecular genetic However, studies together with protein analyses failed to establish closer relationships between these individual groups. [6] [7]
Much of the fossil material of Megalonychidae is fragmentary and largely incomplete. The systematic Relationships of the individual representatives of the group could be worked out therefore so far only insufficiently. However, due to the richness of forms, different lines of development can be traced. One includes largely South American representatives such as Megistonyx or Ahytherium and Ortotherium , respectively, another consists of the North American forms Megalonyx and Pliometanastes (here, since based on skeletal features, also Caribbean sloths such as Megalocnus or Neocnus ). [5] [8] [9] Currently, however, it is not possible to determine direct ancestors for the North American representatives of Megalonychidae. Consequently, their relationship to the South American forms is rather unknown.
Below is a phylogenetic tree of the Megalonychidae, based on the work of Stinnesbeck and colleagues (2021). [10]
Ahytherium must have been a large, rather slow-moving terrestrial sloth that could defend itself from carnivores with its powerful claws. The flattened shape of the tail suggests that Ahytherium might have been a good swimmer. [2] It is suggested to have been a mixed feeder (both browsing and grazing). [11]
Megatherium is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Late Pleistocene. It is best known for the elephant-sized type species Megatherium americanum, primarily known from the Pampas, but ranging southwards to northernmost Patagonia and northwards to southern Bolivia during the late Middle Pleistocene and Late Pleistocene. Various other species belonging to the subgenus Pseudomegatherium ranging in size comparable to considerably smaller than M. americanum are known from the Andean region.
Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. They varied widely in size with the largest, belonging to genera Lestodon, Eremotherium and Megatherium, being around the size of elephants. Ground sloths represent a paraphyletic group, as living tree sloths are thought to have evolved from ground sloth ancestors.
Megatheriidae is a family of extinct ground sloths that lived from approximately 23 mya—11,000 years ago.
Scelidotheriidae is a family of extinct ground sloths within the order Pilosa, suborder Folivora and superfamily Mylodontoidea, related to the other extinct mylodontoid family, Mylodontidae, as well as to the living two-toed sloth family Choloepodidae. The only other extant family of the suborder Folivora is the distantly related Bradypodidae. Erected as the family Scelidotheriidae by Ameghino in 1889, the taxon was demoted to a subfamily of Mylodontidae by Gaudin in 1995. However, recent collagen sequence data indicates the group is less closely related to Mylodon and Lestodon than Choloepus is, and thus it has been elevated back to full family status by Presslee et al. (2019).
Megalonyx is an extinct genus of ground sloths of the family Megalonychidae, native to North America. It evolved during the Pliocene Epoch and became extinct at the end of the Late Pleistocene, living from ~5 million to ~13,000 years ago. The type species, M. jeffersonii, the youngest and largest known species, measured about 3 meters (9.8 ft) in length and weighed up to 1,000 kilograms (2,200 lb).
Megalonychidae is an extinct family of sloths including the extinct Megalonyx. Megalonychids first appeared in the early Oligocene, about 35 million years (Ma) ago, in southern Argentina (Patagonia). There is, however, one possible find dating to the Eocene, about 40 Ma ago, on Seymour Island in Antarctica. They first reached North America by island-hopping across the Central American Seaway, about 9 million years ago, prior to formation of the Isthmus of Panama about 2.7 million years ago. Some megalonychid lineages increased in size as time passed. The first species of these were small and may have been partly tree-dwelling, whereas the Pliocene species were already approximately half the size of the huge Late Pleistocene Megalonyx jeffersonii from the last ice age.
Eremotherium is an extinct genus of giant ground sloth in the family Megatheriidae. Eremotherium lived in southern North America, Central America, and northern South America from the Pliocene, around 5.3 million years ago, to the end of the Late Pleistocene, around 10,000 years ago. Eremotherium was one of the largest ground sloths, with a body size comparable to elephants, weighing around 4–6.5 tonnes and measuring about 6 metres (20 ft) long, slightly larger than its close relative Megatherium.
Catonyx is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Pliocene and Pleistocene epochs. It lived from 2.5 Ma to about 10,000 years ago, existing for approximately 2.49 million years. The most recent date obtained is about 9600 B.P.
Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately 17.49 million years. Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built Nothrotheriops, which reached a length of about 2.75 metres (9.0 ft). While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today.
Australonyx is an extinct genus of ground sloths, endemic to South America during the Late Pleistocene. It was found in Brazil.
Diabolotherium is an extinct genus of megatheriine ground sloth, known from the Late Pleistocene of Peru. Unlike most other extinct mainland sloths, it seems to have been a climber, similar to extinct sloths from the Caribbean. Fossils of the genus were found at the coastal Piedra Escrita site and the Andean Casa del Diablo cave.
Valgipes is an extinct genus of scelidotheriid ground sloth, endemic to intertropical Brazil and Uruguay during the Late Pleistocene. Thought to have been a forest-dwelling browser, Valgipes is a monotypic genus with a complex and long taxonomic history, and is a close relative of Catonyx and Proscelidodon.
Megatheriinae is a subfamily of the Megatheriidae, an extinct family of ground sloths that lived from the Middle Miocene to the Early Holocene.
Megathericulus is an extinct genus of ground sloths in the Megatheriidae family. It lived during the Middle Miocene, 11-16 Ma in what is now South America. Fossils have been found principally in Argentina, Bolivia, and Peru. It is a smaller representative of the megatheres. Despite being one of the earliest-known members of the family, its dentition structure is associated with homodont teeth belonging to the more modern line of evolution. The genus was scientifically named in 1904. Only one species is currently recognized, Megathericulus patagonicus.
Proeremotherium is an extinct genus of megatheriine ground sloths in the family Megatheriidae. It lived during the Late Miocene and Early Pliocene of what is now Venezuela. So far, two largely complete skulls have been recovered in the Falcón Basin in Venezuela. The finds identify the animals as medium-sized representatives of the Megatheriidae. In the cranial anatomy, Proeremotherium resembles the later and giant Eremotherium. It is therefore assumed that the two ground sloths are directly related to each other.
Lakukullus is an extinct genus of nothrotheriid ground sloths that lived during the Middle Miocene around 13.8 to 11.8 million years ago of what is now Bolivia.
Aymaratherium is an extinct genus of nothrotheriid ground sloths that lived during the Late Miocene and Early Pliocene of Bolivia. Fossils of Aymaratherium have been found in the Pomata Ayte locality of the Umala Formation.
Mesopotamocnus is an extinct genus of megalonychid ground sloth that lived during the Late Miocene in what is now Argentina. Fossils have been found in the Ituzaingó Formation of Argentina.
Bolivartherium is an extinct genus of mylodontine mylodontid sloth that lived during the Late Miocene and Late Pliocene in what is now Venezuela. Fossils have been found in the Codore and Urumaco Formations of Venezuela.
Ortotherium is a genus of megalonychid ground sloth from the Late Miocene Ituzaingó Formation of Entre Rios Province, Argentina. Although many species were described, the only valid species of the genus is Ortotherium laticurvatum, with many species being junior synonyms. Ortotherium is known from very fragmentary material, all of which is material from the mandible and teeth. The holotype of O. laticurvatum consists of an incomplete left dentary that had been unearthed from a series of sediments known as ‘Conglomerado osifero’ in Paraná, Argentina. Argentina paleontologist Florentino Ameghino named the species in 1885, though he would go on to name four more, invalid, species of the genus. One species however, O. brevirostrum, has been reclassified as Mesopotamocnus.