Aymaratherium | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Pilosa |
Family: | † Nothrotheriidae |
Subfamily: | † Nothrotheriinae |
Genus: | † Aymaratherium Pujos et al., 2016 |
Species: | †A. jeani |
Binomial name | |
†Aymaratherium jeani Pujos et al., 2016 | |
Aymaratherium is an extinct genus of nothrotheriid ground sloths that lived during the Late Miocene and Early Pliocene of Bolivia. Fossils of Aymaratherium have been found in the Pomata Ayte locality of the Umala Formation. [1]
The genus name, Aymaratherium, is derived from the Aymara, a native ethnic group and language from the Andes, and therion, meaning "beast". The specific name honors Jean Joinville Vacher, for his friendship and constant support for palaeontological investigations over the years. [1]
Aymaratherium was first described in 2016 on the basis of a nearly complete right dentary (specimen MNHN-Bol-V 008954), which was made of the holotype. In addition, three complete humeri (MNHN-Bol-V 003789, 012874 and 012875), a right astragalus (MNHN-Bol-V 012983), and a complete right calcaneus (MNHN-Bol-V 003307) have also been assigned to the genus. [1] The locality of the holotype was recovered from the Pomata-Ayte locality, in the Umala Formation, which is considered to be Early Pliocene in age, and is situated at the Oruro Department in Carangas Province near the city of the same name. Other vertebrate fauna found at the locality include the macraucheniid litoptern Macrauchenia sp., [2] the toxodontid notoungulate Posnanskytherium, several xenarthrans such as the sloths Simomylodon uccasamamensis [3] Megatherium altiplanicum , [4] and the pampatheriid cingulate Plaina .
Aymaratherium is a small to medium-sized nothrotheriid that was of similar size to Nothrotherium and Thalassocnus . According to Pujos et al. (2016), Aymaratherium was diagnosed under the autapomorphies; the presence of lower dentition consisting of a single small caniniform and three large molariform teeth, the absence of a diastema between c1 and m1, a very high hypsodonty index, which is also seen in Megatherium and Megalonyx, and a deep buccinator fossa and extremely thin dorsal margin of the spout. [1] The molariform teeth of the type are mesially convex and the most mesial tooth closely approaches the dorsolateral margin of the mandible meaning that the individual was an adult or subadult at the time of its death. [1]
The 2016 study that described Aymaratherium recovers it as a derived nothrotheriid, within the subfamily Nothrotheriinae, as the sister taxon to the tribe Nothrotheriini. Below is a parsimony tree establishing the relationships between the genera of the Nothrotheriidae, as proposed by Pujos et al, 2016. [1]
Megatherioidea |
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Megatherium is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Pleistocene. It is best known for the elephant-sized type species M. americanum, sometimes called the giant ground sloth, or the megathere, native to the Pampas through southern Bolivia during the Pleistocene. Various other smaller species belonging to the subgenus Pseudomegatherium are known from the Andes.
Hapalops is an extinct genus of ground sloth from the Early to Late Miocene of Brazil, Bolivia, Colombia, and Argentina in South America.
Glossotherium is an extinct genus of mylodontid ground sloths of the subfamily Mylodontinae, which includes large ground-dwelling sloths. It represents one of the best known members of the family, along with Mylodon and Paramylodon. Reconstructed animals were between 3 to 4 metres long and possibly weighed up to 1700 kg. The majority of finds of Glossotherium date from the Middle and Upper Pleistocene, around 300,000 to 10,000 years ago, with a few dating older, possibly as far back as the Pliocene, about 3 million years ago. The range included large parts of South America, east of the Andes roughly from latitude 20 to 40 degrees south, leaving out the Amazon Basin in the north. In western South America, finds are also documented north of the equator. The animals largely inhabited the open landscapes of the Pampas and northern savanna regions.
Eremotherium is an extinct genus of giant ground sloth, in the family Megatheriidae, the largest and most heavily built family of sloths. Eremotherium lived in the southern parts of North America and the northern parts of South America from the Pliocene, around 5.3 million years ago, to the end of the Late Pleistocene, around 10,000 years ago. Eremotherium was widespread in tropical and subtropical lowlands and lived there in partly open and closed landscapes, while its close relative Megatherium lived in more temperate climes. Both genera reached the size of today's elephants and were among the largest mammals in the Americas. Characteristic of Eremotherium was its robust physique with comparatively long limbs and front and hind feet especially for later representatives- three fingers. However, the skull is relatively gracile, the teeth are uniform and high-crowned. Like today's sloths, Eremotherium was a pure herbivore who dined on leaves and grasses, though it may have had semi-aquatic habits like modern hippos based on isotopic data. Eremotherium was a generalist that could adapt its diet to the respective local and climatic conditions of many regions. Finds of Eremotherium are common and widespread, with fossils being found as far north as South Carolina in the United States and as far south as Rio Grande Do Sul, and many complete skeletons have been unearthed.
Catonyx is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Pliocene and Pleistocene epochs. It lived from 2.5 Ma to about 10,000 years ago, existing for approximately 2.49 million years. The most recent date obtained is about 9600 B.P.
Scelidodon is an extinct genus of South American ground sloths. Its remains have been found in the Yupoí and Uspara Formations of Argentina, the Ulloma, Umala, Ñuapua and Tarija Formations of Bolivia, in Brazil, in Chile and in Peru. The youngest fossils have been dated to as recently as 9000 B.P.
Nothrotheriidae is a family of extinct ground sloths that lived from approximately 19.8 mya—10,000 years ago, existing for approximately 19.79 million years. Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built Nothrotheriops, which reached a length of about 2.75 metres (9.0 ft). While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today.
Macroeuphractus is a genus of extinct armadillos from the Late Miocene to Late Pliocene of South America. The genus is noted for its large size, with Macroeuphractus outesi being the largest non-pampathere or glyptodont armadillo discovered, as well as its specializations for carnivory, unique among all xenarthrans.
Hoffstetterius is an extinct genus of toxodontid notoungulate mammal, belonging to the subfamily Toxodontinae whose remains were discovered in the Middle to Late Miocene Mauri Formation in the La Paz Department in Bolivia. The only described species is the type Hoffstetterius imperator.
Australonyx is an extinct genus of ground sloths, endemic to South America during the Late Pleistocene. It was found in Brazil.
Megathericulus is an extinct genus of ground sloths in the Megatheriidae family. It lived during the Middle Miocene, 11-16 million years ago in what is now South America. Fossils have been found principally in Argentina Bolivia and Peru. It is a smaller representative of the megatheres. Despite being one of the earliest-known members of the family, its dentition structure is associated with homodont teeth belonging to the more modern line of development. The genus was scientifically named in 1904. Only one species is currently recognized, Megathericulus patagonicus.
Plesiotypotherium is an extinct genus of Notoungulate, belonging to the suborder Typotheria. It lived from the Middle to the Late Miocene, and its fossilized remains were discovered in South America.
Archaeomylodon is an extinct genus of mylodontine ground sloths that lived during the Middle Pleistocene of what is now Argentina. It is known so far only from a single skull, which in its dimensions corresponds to those of the giant Lestodon. However, the skull differs from this one by its narrower and higher snout. In addition, the anterior canine teeth, which are usually large in many mylodonts, are greatly reduced. The find comes from the Pampa region of South America and was deposited in about 700,000 years old sediments.
Mcdonaldocnus is an extinct genus of nothrotheriid ground sloths that lived during the Middle Miocene and Early Pliocene of what is now Bolivia and Argentina. It was originally placed in the genus Xyophorus but was subsequently recognized as a distinct genus by Gaudin and colleagues in 2022. The authors reassigned the material of "Xyophorus" bondesioi, Xyophorusvillarroeli and Xyophorus sp. to Mcdonaldocnus. Fossils of Mcdonaldocnus have been found in the Cerro Azul Formation of Argentina.
Baraguatherium is an extinct genus of ground sloths of the family Mylodontidae that lived during the Early Miocene of what is now Venezuela. It dates to the Early Miocene, around 20.44 to 15.97 million years ago and represents the oldest representative of its family in the northern part of South America to date. The structure of the teeth suggests that the genus represents a rather basal form within the Mylodontidae. Unlike other mylodonts, which tended to prefer open grasslands, Baraguatherium lived in a riverine, coastal tropical rainforest.
Simomylodon is an extinct genus of ground sloths from the family Mylodontidae. It lived from the Late Miocene to the Middle Pliocene of what is now Bolivia and Argentina, 5.3 to 2.8 million years ago. The most important find material comes from the central Altiplano in Bolivia and includes several skulls and dentition remains. Thus, the so far documented body skeleton is the best known and most significant of a Miocene representative of the Mylodontidae. On the basis of the remains, it can be concluded that it is a rather small member of the Mylodontidae. The construction of the limbs supports ground-dwelling locomotion, but this does not exclude occasional digging or climbing. The type and only known species is Simomylodon uccasamamensis.
Lakukullus is an extinct genus of nothrotheriid ground sloths that lived during the Middle Miocene around 13.8 to 11.8 million years ago of what is now Bolivia.
Mesopotamocnus is an extinct genus of megalonychid ground sloth that lived during the Late Miocene in what is now Argentina. Fossils have been found in the Ituzaingó Formation of Argentina.
Bolivartherium is an extinct genus of mylodontine mylodontid sloth that lived during the Late Miocene and Late Pliocene in what is now Venezuela. Fossils have been found in the Codore and Urumaco Formations of Venezuela.