Analcitherium

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Analcitherium
Temporal range: Early Miocene (Santacrucian)
~17.5  Ma
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Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Pilosa
Family: Scelidotheriidae
Genus: Analcitherium
Ameghino, 1891
Species:
A. antarticum
Binomial name
Analcitherium antarticum
Ameghino, 1891

Analcitherium is an extinct genus of scelidotheriid sloth that lived during the Early Miocene in what is now Argentina. Fossils have been found in the Santa Cruz Formation of Argentina. [1]

Taxonomy

Analcitherium was first named by Florentino Ameghino in 1891 based on fossils found in Argentina, dating to the Early Miocene. Originally thought to belong to a juvenile Nematherium , [2] it is now usually considered to be a distinct genus.

Analcitherium is a member of the Scelidotheriidae, a group of terrestrial sloths known from the Oligocene and Pleistocene that a characterized by an elongated snout. Although scelidotheriids are usually placed as a subfamily of the Mylodontidae, they are sometimes considered to be a separate family, Scelidotheriidae. [3]

Below is a phylogenetic tree of the Mylodontidae, based on the work of Varela et al. 2018. [4]

Mylodontidae  

Nematherium

Scelidotheriinae  

Analcitherium

Neonematherium

Sibyllotherium

Proscelidodon

Catonyx

Scelidotherium

Valgipes

Pseudoprepotherium

Orophodontinae? 

Brievabradys

Octodontotherium

Paroctodontotherium

Octomylodon

Urumacotherium

Baraguatherium

Octodontobradys

Mylodontinae  

Mylodon

Thinobadistes

Lestodon

Lestobradys

Bolivartherium

Simomylodon

Pleurolestodon

Glossotherium

Paramylodon

Related Research Articles

<span class="mw-page-title-main">Mylodontidae</span> Extinct family of ground sloths

Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa, living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another family of extinct ground sloths, Scelidotheriidae, as well as to the extant arboreal two-toed sloths, family Choloepodidae; together these make up the superfamily Mylodontoidea. Phylogenetic analyses based on morphology uncovered the relationship between Mylodontidae and Scelidotheriidae; in fact, the latter was for a time considered a subfamily of mylodontids. However, molecular sequence comparisons were needed for the correct placement of Choloepodidae. These studies have been carried out using mitochondrial DNA sequences as well as with collagen amino acid sequences. The latter results indicate that Choloepodidae is closer to Mylodontidae than Scelidotheriidae is. The only other living sloth family, Bradypodidae, belongs to a different sloth radiation, Megatherioidea.

<span class="mw-page-title-main">Scelidotheriidae</span> Extinct family of prehistoric ground sloths

Scelidotheriidae is a family of extinct ground sloths within the order Pilosa, suborder Folivora and superfamily Mylodontoidea, related to the other extinct mylodontoid family, Mylodontidae, as well as to the living two-toed sloth family Choloepodidae. The only other extant family of the suborder Folivora is the distantly related Bradypodidae. Erected as the family Scelidotheriidae by Ameghino in 1889, the taxon was demoted to a subfamily of Mylodontidae by Gaudin in 1995. However, recent collagen sequence data indicates the group is less closely related to Mylodon and Lestodon than Choloepus is, and thus it has been elevated back to full family status by Presslee et al. (2019).

<i>Scelidotherium</i> Extinct genus of ground sloths

Scelidotherium is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Late Pleistocene epoch. It lived from 780,000 to 11,000 years ago, existing for approximately 0.67 million years.

<span class="mw-page-title-main">Pilosa</span> Order of mammals

The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes anteaters and sloths. The name comes from the Latin word for "hairy".

<i>Protypotherium</i> Extinct genus of notoungulates

Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.

<i>Glossotherium</i> Extinct genus of ground sloths

Glossotherium is an extinct genus of large mylodontid ground sloths of the subfamily Mylodontinae. It represents one of the best-known members of the family, along with Mylodon and Paramylodon. Reconstructed animals were between 3 and 4 metres long and possibly weighed up to 1,002.6–1,500 kg. The majority of finds of Glossotherium date from the Middle and Upper Pleistocene, around 300,000 to 10,000 years ago, with a few dating older, as far back Pliocene, about 3.3-3 million years ago. The range included large parts of South America, east of the Andes roughly from latitude 20 to 40 degrees south, leaving out the Amazon Basin in the north. In western South America, finds are also documented north of the equator. The animals largely inhabited the open landscapes of the Pampas and northern savanna regions.

Chubutherium is an extinct genus of ground sloth from the Late Oligocene and Early Miocene of Chubut Province, Argentina.

<i>Catonyx</i> Extinct genus of ground sloths

Catonyx is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Pliocene and Pleistocene epochs. It lived from 2.5 Ma to about 10,000 years ago, existing for approximately 2.49 million years. The most recent date obtained is about 9600 B.P.

<i>Scelidodon</i> Extinct genus of ground sloths

Scelidodon is an extinct genus of South American ground sloths. Its remains have been found in the Yupoí and Uspara Formations of Argentina, the Ulloma, Umala, Ñuapua and Tarija Formations of Bolivia, in Brazil, in Chile and in Peru. The youngest fossils have been dated to as recently as 9000 B.P.

<span class="mw-page-title-main">Nothrotheriidae</span> Extinct family of ground sloths

Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately 17.49 million years. Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built Nothrotheriops, which reached a length of about 2.75 metres (9.0 ft). While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today.

<i>Nematherium</i> Extinct genus of ground sloths

Nematherium is an extinct genus of ground sloth belonging to Mylodontoidea, it is either considered to be a member of Mylodontidae or Scelidotheriidae. It lived during the Middle Miocene epoch (Santacrucian). Fossils have been found in the Cura-Mallín Formation of Chile and the Santa Cruz and Sarmiento Formations of Argentina.

<i>Pseudoprepotherium</i> Extinct genus of ground sloths

Pseudoprepotherium is an extinct genus of sloths of the family Mylodontidae. It was widespread across northern South America during the Early to Late Miocene epoch around 21 to 5.3 million years ago. Fossils of the animal have been found in Brazil, Venezuela, and Peru. Pseudoprepotherium lived in a tropical climate with a water-rich environment. Their known remains are limited to limb bones, except for a few skulls and teeth. Based on these remains, they were most likely medium to large-sized mylodontid. The genus was described in 1961 and currently contains three species, which were originally assigned to the genus Prepotherium.

<i>Proscelidodon</i> Extinct genus of ground sloths

Proscelidodon is an extinct genus of ground sloths in the family Scelidotheriidae. It lived during the Miocene and Pliocene of what is now Argentina and Bolivia. The genus was described in 1935.

Archaeomylodon is an extinct genus of mylodontine ground sloths that lived during the Middle Pleistocene of what is now Argentina. It is known so far only from a single skull, which in its dimensions corresponds to those of the giant Lestodon. However, the skull differs from this one by its narrower and higher snout. In addition, the anterior canine teeth, which are usually large in many mylodonts, are greatly reduced. The find comes from the Pampa region of South America and was deposited in about 700,000 years old sediments.

Urumacotherium is an extinct genus of ground sloths of the family Mylodontidae. It lived from the Middle Miocene to the Early Pliocene of what is now Brazil, Peru and Venezuela.

Baraguatherium is an extinct genus of ground sloths of the family Mylodontidae that lived during the Early Miocene of what is now Venezuela. It dates to the Early Miocene, around 20.44 to 15.97 million years ago and represents the oldest representative of its family in the northern part of South America to date. The structure of the teeth suggests that the genus represents a rather basal form within the Mylodontidae. Unlike other mylodonts, which tended to prefer open grasslands, Baraguatherium lived in a riverine, coastal tropical rainforest.

<i>Prepoplanops</i> Extinct genus of ground sloths

Prepoplanops is an extinct genus of ground sloth of the family Megatheriidae. It lived in the Miocene around 18 to 16 million years ago of what is now Argentina. The only known species is Prepoplanops boleadorensis.

Bolivartherium is an extinct genus of mylodontine mylodontid sloth that lived during the Late Miocene and Late Pliocene in what is now Venezuela. Fossils have been found in the Codore and Urumaco Formations of Venezuela.

Neonematherium is an extinct genus of scelidotheriid ground sloths that lived in Argentina, Chile, and Colombia during the Early to Late Miocene. Fossils have been found in the Honda Group of Colombia, and the Río Frías Formation of Chile.

<span class="mw-page-title-main">Santa Cruz Formation</span> Geological formation in Patagonia

The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and in adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with eponymous Santacrucian SALMA. The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which has a lithology predominantly consisting of tuffaceous deposits and fine grained sedimentary claystone and mudstone, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily claystone, mudstone, and sandstone. The environment of deposition is interpreted to have been mostly fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but likely became somewhat cooler and drier towards the end of the sequence. The Santa Cruz Formation is known for its abundance of South American native ungulates, as well as an abundance of rodents, xenarthrans, and metatherians.

References

  1. Toledo, Néstor; Cassini, Guillermo Hernán; Vizcaíno, Sergio Fabián; Bargo, M. Susana (June 2014). "Mass Estimation of Santacrucian Sloths from the Early Miocene Santa Cruz Formation of Patagonia, Argentina" (PDF). Acta Palaeontologica Polonica. 59 (2): 267–280. doi: 10.4202/app.2012.0009 . ISSN   0567-7920.
  2. Simpson, George Gaylord; Bird, Junius Bouton (1941). "A Miocene sloth from southern Chile" (PDF). American Museum Novitates (1156): 1–5.
  3. Presslee, S.; Slater, G. J.; Pujos, F.; Forasiepi, A. M.; Fischer, R.; Molloy, K.; Mackie, M.; Olsen, J. V.; Kramarz, A.; Taglioretti, M.; Scaglia, F.; Lezcano, M.; Lanata, J. L.; Southon, J.; Feranec, R.; Bloch, J.; Hajduk, A.; Martin, F. M.; Gismondi, R. S.; Reguero, M.; de Muizon, C.; Greenwood, A.; Chait, B. T.; Penkman, K.; Collins, M.; MacPhee, R.D.E. (2019). "Palaeoproteomics resolves sloth relationships" (PDF). Nature Ecology & Evolution. 3 (7): 1121–1130. Bibcode:2019NatEE...3.1121P. doi:10.1038/s41559-019-0909-z. PMID   31171860. S2CID   174813630.
  4. Varela, L.; Tambusso, P. S.; McDonald, H. G.; Fariña, R. A. (2018). "Phylogeny, Macroevolutionary Trends and Historical Biogeography of Sloths: Insights From a Bayesian Morphological Clock Analysis". Systematic Biology. 68 (2): 204–218. doi:10.1093/sysbio/syy058. PMID   30239971.