Santa Cruz Formation

Last updated
Santa Cruz Formation
Stratigraphic range: Burdigalian-Langhian
(Santacrucian-Friasian)
~18–15.2  Ma
O
S
D
C
P
T
J
K
Pg
N
Santa Cruz Formation strata.jpg
Type Geological formation
Sub-unitsEstancia La Costa Member, Estancia La Angelina Member (coastal section)
Underlies Cerro Boleadoras Formation
Overlies Monte Léon Formation
ThicknessOver 295 metres
Lithology
Primary
Location
Country Argentina, Chile
Extent Austral Basin
Type section
Named byFurque & Camacho
Locationnear Lago Argentino
Year defined1972
Santa Cruz in Argentina (+Falkland hatched).svg
Santa Cruz Province in Argentina, where the majority of the formation of exposed

The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with the eponymous Santacrucian age of the SALMA (South American land mammal age) timescale. [1] [2] The Santa Cruz Formation is known for its abundance of vertebrate fossils, including South American native ungulates (astrapotheres, litopterns, notoungulates), [3] as well as rodents, xenarthrans (armadillos, sloths, anteaters), and metatherians.

Contents

The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which consists predominately of tuffaceous deposits and fine grained mudrock, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily mudrock, and sandstone. The environment of deposition was mostly fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but likely became somewhat cooler and drier towards the end of the sequence. [1]

Stratigraphy

The Santa Cruz Formation is exposed in isolated outcrops across the Magallanes/Austral Basin extending from the Atlantic coast to the Andes, especially along the Santa Cruz River, as well as along the southern coastline of Santa Cruz Province. [4] While primarily located in Argentina, small outcrops are also found in Chile. [5] The base of the formation is defined by a marine regression event transitioning from the marine environment of the underlying Monte Léon Formation, which formed when large areas of Patagonia were submerged as a part of the Patagoniense Transgression.

The main source of sediment input to the basin was from the Andean orogeny to the west. The formation reaches a maximum thickness of over 295 meters, though the total thickness of the formation is strongly controlled by subsequent erosion and the 295 meters likely does not represent a complete sequence. The formation primarily consists of floodplain deposits. The lower parts of the formation have an abundance of tuffs and tuffaceous sediments. These likely originated from distant eruptions that were transported into the basin by aerial fallout, wind and/or river transport. The formation likely spans an approximately 3 million year interval in the late Early Miocene around 18 to 15.2 million years ago, during the Burdigalian and Langhian stages. [4]

Paleoenvironment

The environment of the Santa Cruz Formation is thought to have been relatively warm and humid, including a mix of open savanna, gallery forests and semi-deciduous forests. Permanent bodies of water such as lakes, ponds and streams are likely to have been present. [6]

Paleoflora

NameSpeciesLocalityMaterialNotesImage
Nothofagus [7] IndeterminateRincón del Buque, Punta SurLeaves, wood
Araucaria [7] IndeterminatePunta SurTwigMorphologically similar to A. marensii from the Eocene of Antarctica
Lauraceae [7] IndeterminatePunta SurWoodAssigned to form genus Laurinoxylon
Myrceugenia [7] M. chubutensePunta SurWoodA member of the family Myrtaceae
Eucryphiaceoxylon [7] E. eucryphioidesPunta SurWoodWood probably belonging to the genus Eucryphia
Faboideae [7] IndeterminatePunta SurWoodPossible affinities to Sophora (Fabaceae)
Doroteoxylon [7] D. vicenti-pereziiPunta SurWoodWood with affinities to the subfamily Caesalpinioideae
Akanioxylon [8] A. santacrucensisPunta SurWoodA member of the family Akaniaceae
Chloridoideae [7] Indeterminate Phytoliths Grass
Panicoideae [7]
Danthonioideae [7]
Pooideae [7]

Paleofauna

Invertebrates

NameSpeciesMaterialNotesImage
Crassostrea [9] C. orbignyiNumerous individuals in large beds at the base of the formationA marine true oyster
Diplodon [10] D. cf. colhuapiensisA freshwater bivalve belonging to Hyriidae
Stephadiscus [11] IndeterminateA terrestrial gastropod belonging to Charopidae
Gastrocopta [12] G.patagonicaA terrestrial gastropod belonging to Vertiginidae
? Scolodonta [12] IndeterminateA terrestrial gastropod belonging to Scolodontidae
Punctum [12] P. patagonicumA terrestrial gastropod belonging to Punctidae
Zilchogyra [12] Z. miocenicaA terrestrial gastropod belonging to Charopidae
Patagocharopa [12] P. enigmaticaA terrestrial gastropod of uncertain affinities, possibly a member of Charopidae
Porifera [7] Unspecified Spicules Freshwater sponge
Bacillariophyceae [7] Unspecified Frustules Freshwater diatoms
Chrysophyceae [7] UnspecifiedFreshwater algae

Amphibians

NameSpeciesLocalityMaterialNotesImage
Calyptocephalella [13] [14] C. cf. canqueliEstancia La Costa, Rinconada de los LópezSkull and jaw fragmentsA calyptocephalellid frog
Neobatrachia [13] IndeterminateEstancia La CostaSkull fragments and presacral vertebrateA frog originally assigned to "Leptodactylidae", but requires further investigation

Birds

NameSpeciesLocalityMaterialNotesImage
Anisolornis [15] A. excavatusA bird of uncertain affinities, authors have varyingly suggested affinities to trumpeters or limpkins
Ankonetta [15] A. larriestraiPartial tarsometatarsusA basal member of Anatidae
Badiostes [15] B. patagonicusA member of the family Falconidae
Brontornis [15] B. burmeisteriControversial taxonomic position, either considered a terror bird or an anserimorph
Brontornis burmeisteri femore sinistro.JPG
Dryornis [16] [17] D. pampeanusD. hatcheriTwo partial left humerusA New World vulture.
Eutelornis [15] E. patagonicus

E. australis

Limb fragmentsA member of Anseriformes, relationships of the species to each other or to other Ansiferiformes is uncertain
Liptornis [15] L. hesternusA member of Anhingidae
Macranhinga [15] IndeterminateA member of Anhingidae
Miocariama [15] [18] M. santacrucensisPartial cranium and tibiotarsi fragmentsA seriema
Nothurinae [15] 2 Indeterminate species Tinamou
Opisthodactylus [15] O. patagonicusLimb, vertebra and skull fragmentsA member of Rheidae
Patagornis [15] P. marshiA terror bird
Patagornis marshi.png
Phorusrhacos [19] P. longissimusPartial skullsA terror bird
Phorusrhacos longissimus skull 824.jpg
Protibis [15] P. cnemialisdistal end of tibiotarsusPotentially a spoonbill
Psilopterus [15] P. lemoinei

P. bachmanni

A terror bird
Psilopterus australis - AMNH - DSC06300.JPG
Thegornis [15] T. musculosus

T. debilis

A member of the family Falconidae
Archaeopsophia [20] A. aoniA member of the family Psophiidae (trumpeters)
Chehuenia [20] C. facongrandeiA roller
Kaikenia [20] K. mourerchauvireaAn anatid belonging to the subfamily Tadorninae
Peioa [20] P. australisA member of Anseriformes.

Reptiles

NameSpeciesLocalityMaterialNotesImage
Pristidactylus [21] IndeterminateLa CuevaFragmentary jaw bonesOriginally assigned to the dubious genus "Erichosaurus"
Pleurodonta [13] [21]
Tupinambis [13] Monte LeónFragmentary dentaries and maxillaA teiid lizard
Colubridae [13] [21] Cerro ObservatorioPartial trunk vertebraeA snake

Mammals

Meridiungulates

Astrapotheres
NameSpeciesLocalityMaterialNotesImage
Astrapotherium [22] [23] [24] [25] [26] [2] [27] A. magnum

A. nanum

A. sp.

An astrapotheriid
Astrapotherium magnum NT.jpg
Litopterns
NameSpeciesLocalityMaterialNotesImage
Adianthus [28] A. bucatusAn adianthid litoptern
Anisolophus [22] [3] [23] [29] [26] [2] A. australis

A. floweri

A proterotheriid litoptern
Diadiaphorus [30] [3] [23] [2] [29] [31] [27] D. majusculus

D. sanctaecrucis

D. sp.

A proterotheriid litoptern
Diadiaphorus majusculus NT.jpg
Tetramerorhinus [32] [30] [3] [23] [2] [26] [24] [25] [29] [31] [27] T. cingulatum

T. fleaglei

T. mixtum

T. lucarius

T. prosistens

T. sp.

Estancia La Costa MemberA proterotheriid litoptern
Tetramerorhinus jaw.png
Theosodon [33] [30] [3] [23] [34] [2] [26] [24] [25] [29] [31] [27] T. fontanae

T. garretorum

T. gracilis

T. karaikensis

T. lydekkeri

T. patagonicum

Estancia La Costa MemberA macraucheniid litoptern
Theosodon NT.jpg
Thoatherium [30] T. minisculum

T. sp.

Estancia La Costa MemberA proterotheriid litoptern
Thoatheriumknight.jpg
Notoungulates
NameSpeciesLocalityMaterialNotesImage
Adinotherium [3] [23] [35] [26] [24] [27] A. ovinum

A. robustum

A. sp.

A toxodontid notoungulate
Adinotherium ovinum.jpg
Cochilius [36] C. sp.An interatheriid notoungulate
Cochilius volvens.jpg
Hegetotherium [22] [3] [23] [2] [26] [24] [25] [37] H. mirabile

H. sp.

A hegetotheriid notoungulate
Hegetotherium.svg
Homalodotherium [22] [3] [23] [2] [26] [24] [25] [27] H. cunninghamiA homalodotheriid notoungulate
Homalodotherium FMNH.jpg
Interatherium [38] [22] [3] [23] [35] [2] [26] [24] [25] [27] I. anguliferum

I. brevifrons

I. dentatum

I. interruptum

I. robustum

I. rodens

I. supernum

An interatheriid notoungulate
Interatherium.jpg
Neoicochilus [36] N. undulatusAn interatheriid notoungulate
Nesodon [22] [3] [23] [35] [2] [26] [24] [25] [27] N. imbricatusA toxodontid notoungulate
Nesodon imbricatus skeleton reconstruction.jpg
Notohippus [39] N. toxodontoidesA notohippid notoungulate
Pachyrukhos [22] [3] [23] [2] [26] [24] [25] [37] P. moyaniA hegetotheriid notoungulate
Skeleton-of-Pachyrukhos-from-Sinclair-1909.png
Patriarchus [38] [40] P. palmidensAn interatheriid notoungulate
Protypotherium [33] [38] [22] [3] [23] [2] [26] [24] [25] [40] [27] P. attenuatum

P. australe

P. praerutilum

P. sp.

An interatheriid notoungulate
ProtypotheriumSinclair 1.jpg

Xenarthrans

Pilosa
NameSpeciesLocalityMaterialNotesImage
Analcimorphus [41] [42] A. giganteus

A. inversus

Estancia La Costa MemberA basal megatherioid sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE XLVII) (6859444178).jpg
Analcitherium [43] [41] [42] A. antarcticumEstancia La Costa MemberA scelidotheriid sloth
Eucholoeops [44] E. fronto

E. ingens

E. litoralis

E. titans

Estancia La Costa MemberA megalonychid ground sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE LV) (7005566005).jpg
Hapalops [26] H. longiceps

H. elongatus

H. indifferens

H. angustipalatus

H. platycephalus

H. ponderosus

H. rectangularis

A ground sloth belonging to Megatherioidea
Hapalops skeleton.jpg
Hyperleptus [45] IndeterminateA megatherioid ground sloth of uncertain affinities
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE LIV) (7005565749).jpg
Megalonychotherium M. atavusA megalonychid ground sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE XLVI) (7005563487).jpg
Mylodontidae [45] Indeterminate
Nematherium [45] N. angulatumA ground sloth belonging to Mylodontoidea
Nematherium angulatum.JPG
Pelecyodon [45] P. cristatusA ground sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE LII) (7005565177).jpg
Planops [45] P. magnusA megatheriid ground sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE LVIII) (7005566787).jpg
Protamandua [45] P. rothiAn anteater
Prepotherium [45] P. potens

P. filholi

A ground sloth
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE LIX) (6859447460).jpg
Megalonychidae IndeterminateA ground sloth
Schismotherium [46] [47] S. fractumA ground sloth
Xyophorus [47] X. atlanticus

X. latirostris

A nothrotheriid ground sloth
Cingulata
NameSpeciesLocalityMaterialNotesImage
Anantiosodon [48] A. rarusAn armadillo
cf. Asterostemma [48] cf. A. depressaA glyptodont
Cochlops [48] C. muricatusA glyptodont
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE XIX) (6859436306).jpg
Eucinepeltus [48] E. petesatusA glyptodont
Metopotoxus [48] M. laevatusA glyptodont
Parutaetus [48] P. sp.An armadillo
Peltephilus [48] P. ferox

P. giganteus

P. nanus

P. pumilus

P. strepens

A horned armadillo
Peltephilus ferox skull.jpg
Proeutatus [48] [26] P. carinatus

P. deleo

P. lagena

P. oenophorus

P. robustus

An armadillo
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE XI) (7005553277) (cropped).jpg
Propalaehoplophorus [48] P. australis

P. incisivusP. minus

A glyptodont
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE XXII) (7005556677).jpg
Prozaedyus [48] [26] P. exilis

P. proximus

An armadillo
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE V) (6859432278).jpg
Stenotatus [26] S. patagonicusAn armadillo
Reports of the Princeton University Expeditions to Patagonia, 1896-1899 (PLATE VII) (6859432828).jpg
Stegotherium [48] S. tauberi

S. tessellatum

An armadillo
Stegotherium tesselatum.jpg
Vetelia [48] [49] V. punctaAn armadillo
Vetelia.png

Metatherians

NameSpeciesLocalityMaterialNotesImage
Abderites [50] A. meridionalisA member of Abderitidae (Paucituberculata)
Acdestis [50] A. oweniiA. lemaireiA member of Palaeothentidae (Paucituberculata)
Acrocyon [51] A. sectoriusA borhyaenid sparassodont
Acyon [51] A. tricuspidatusA hathliacynid sparassodont
Arctodictis [51] A. muniziA borhyaenid sparassodont
Borhyaena [51] B. tuberataA borhyaenid sparassodont
Borhyaena tuberata.JPG
Cladosictis [51] C. patagoniaA sparassodont
Lycopsis [51] L. torresiA sparassodont
Microbiotherium [50] M. acicula

M. patagonicum

M. gallegosense

M. tehuelchum

A member of Microbiotheriidae (Microbiotheria)
Palaeothentes [50] P. aratae

P. minutus

P. intermedius

P. lemoinei

P. pascuali

A member of Palaeothentidae (Paucituberculata)
Palaeothentes lepidus mandible.jpg
Perathereutes [51] P. pungensA hathliacynid sparassodont
Phonocdromus [50] P. gracilisA member of Pichipilidae (Paucituberculata)
Prothylacinus [51] P. patagonicusA sparassodont
Prothylacinus patagonicus.png
Pseudonotictis [51] P. pusillusA hathliacynid sparassodont
Sipalocyon [51] S. gracilis

S. obusta

A hathliacynid sparassodont
Stilotherium [50] S. dissimileA member of Caenolestidae (Paucituberculata)

Rodents

NameSpeciesLocalityMaterialNotesImage
Acarechimys [47] A. minutus

A. minutissmus

A. constans

A. gracilis

A member of Octodontoidea
Acaremys [52] [26] A. murinus

A. messor

A member of Acaremyidae (Octodontoidea)
Adelphomys [52] A. candidusA member of Octodontoidea
Dudumus [52] Indeterminate, potentially new speciesA member of Octodontoidea
Eocardia [52] E. montana

"E". excavata

"E". fissa

A member of Cavioidea
Eocardia.jpg
Neoreomys [52] N. australisA member of Cavioidea
Perimys [52] P. erutus

P. onustus

P. incavatus

A member of Chinchilloidea
Perimys tooth.png
Phanomys [52] P. mixtus

P. vetulus

A member of Cavioidea
Pliolagostomus [52] P. notatusA member of Chinchilloidea
Prolagostomus [52] P. pusilllusA member of Chinchilloidea
Prospaniomys [52] Indeterminate, potentially new speciesA member of Octodontoidea
Pseudoacaremys [52] P. kramarziA member of the family Acaremyidae (Octodontoidea)
Schistomys [52] S. erroA member of Cavioidea
Sciamys [52] S. principalis

S. varians

S. latidens

A member of the family Acaremyidae (Octodontoidea)
Scleromys [52] S. angustusA member of Chinchilloidea
Spaniomys [52] S. riparius

S. regularis

A member of Octodontoidea
Steiromys [52] S. dentatus

S. duplicatus

A member of Erethizontoidea

Primates

NameSpeciesLocalityMaterialNotesImage
Homunculus [53] [54] [55] H. patagonicus, H. vizcainoiA New World monkey
Killikaike [56] K. blakeiA New World monkey, some authors regard the taxon as a synonym of Homunculuspatagonicus, while others regard them as distinct.
Killikaike blakei.png

Meridiolestida

NameSpeciesLocalityMaterialNotesImage
Necrolestes [57] N. patagonicusSkull and postcranial remainsA mole-like meridiolestidan, youngest known member of the group
Necrolestes.svg

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Ortotherium is a genus of megalonychid ground sloth from the Late Miocene Ituzaingó Formation of Entre Rios Province, Argentina. Although many species were described, the only valid species of the genus is Ortotherium laticurvatum, with many species being junior synonyms. Ortotherium is known from very fragmentary material, all of which is material from the mandible and teeth. The holotype of O. laticurvatum consists of an incomplete left dentary that had been unearthed from a series of sediments known as ‘Conglomerado osifero’ in Paraná, Argentina. Argentina paleontologist Florentino Ameghino named the species in 1885, though he would go on to name four more, invalid, species of the genus. One species however, O. brevirostrum, has been reclassified as Mesopotamocnus.

<span class="mw-page-title-main">Peltephilidae</span> Family of South American cingulates (armadillos)

Peltephilidae is a family of South American cingulates (armadillos) that lived for over 40 million years, but peaked in diversity towards the end of the Oligocene and beginning of the Miocene in what is now Argentina. They were exclusive to South America due to its geographic isolation at the time, one of many of the continent's strange endemic families. Peltephilids are one of the earliest known cingulates, diverging from the rest of Cingulata in the Early Eocene.

References

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