Santa Cruz Formation | |
---|---|
Stratigraphic range: Burdigalian-Langhian (Santacrucian-Friasian) ~ | |
Type | Geological formation |
Sub-units | Estancia La Costa Member, Estancia La Angelina Member (coastal section) |
Underlies | Cerro Boleadoras Formation |
Overlies | Monte Léon Formation |
Thickness | Over 295 metres |
Lithology | |
Primary | |
Location | |
Country | Argentina, Chile |
Extent | Austral Basin |
Type section | |
Named by | Furque & Camacho |
Location | near Lago Argentino |
Year defined | 1972 |
Santa Cruz Province in Argentina, where the majority of the formation of exposed |
The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with the eponymous Santacrucian age of the SALMA (South American land mammal age) timescale. [1] [2] The Santa Cruz Formation is known for its abundance of vertebrate fossils, including South American native ungulates (astrapotheres, litopterns, notoungulates), [3] as well as rodents, xenarthrans (armadillos, sloths, anteaters), and metatherians.
The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which consists predominately of tuffaceous deposits and fine grained mudrock, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily mudrock, and sandstone. The environment of deposition was mostly fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but likely became somewhat cooler and drier towards the end of the sequence. [1]
The Santa Cruz Formation is exposed in isolated outcrops across the Magallanes/Austral Basin extending from the Atlantic coast to the Andes, especially along the Santa Cruz River, as well as along the southern coastline of Santa Cruz Province. [4] While primarily located in Argentina, small outcrops are also found in Chile. [5] The base of the formation is defined by a marine regression event transitioning from the marine environment of the underlying Monte Léon Formation, which formed when large areas of Patagonia were submerged as a part of the Patagoniense Transgression.
The main source of sediment input to the basin was from the Andean orogeny to the west. The formation reaches a maximum thickness of over 295 meters, though the total thickness of the formation is strongly controlled by subsequent erosion and the 295 meters likely does not represent a complete sequence. The formation primarily consists of floodplain deposits. The lower parts of the formation have an abundance of tuffs and tuffaceous sediments. These likely originated from distant eruptions that were transported into the basin by aerial fallout, wind and/or river transport. The formation likely spans an approximately 3 million year interval in the late Early Miocene around 18 to 15.2 million years ago, during the Burdigalian and Langhian stages. [4]
The environment of the Santa Cruz Formation is thought to have been relatively warm and humid, including a mix of open savanna, gallery forests and semi-deciduous forests. Permanent bodies of water such as lakes, ponds and streams are likely to have been present. [6]
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Nothofagus [7] | Indeterminate | Rincón del Buque, Punta Sur | Leaves, wood | ||
Araucaria [7] | Indeterminate | Punta Sur | Twig | Morphologically similar to A. marensii from the Eocene of Antarctica | |
Lauraceae [7] | Indeterminate | Punta Sur | Wood | Assigned to form genus Laurinoxylon | |
Myrceugenia [7] | M. chubutense | Punta Sur | Wood | A member of the family Myrtaceae | |
Eucryphiaceoxylon [7] | E. eucryphioides | Punta Sur | Wood | Wood probably belonging to the genus Eucryphia | |
Faboideae [7] | Indeterminate | Punta Sur | Wood | Possible affinities to Sophora (Fabaceae) | |
Doroteoxylon [7] | D. vicenti-perezii | Punta Sur | Wood | Wood with affinities to the subfamily Caesalpinioideae | |
Akanioxylon [8] | A. santacrucensis | Punta Sur | Wood | A member of the family Akaniaceae | |
Chloridoideae [7] | Indeterminate | Phytoliths | Grass | ||
Panicoideae [7] | |||||
Danthonioideae [7] | |||||
Pooideae [7] |
Name | Species | Material | Notes | Image |
---|---|---|---|---|
Crassostrea [9] | C. orbignyi | Numerous individuals in large beds at the base of the formation | A marine true oyster | |
Diplodon [10] | D. cf. colhuapiensis | A freshwater bivalve belonging to Hyriidae | ||
Stephadiscus [11] | Indeterminate | A terrestrial gastropod belonging to Charopidae | ||
Gastrocopta [12] | G.patagonica | A terrestrial gastropod belonging to Vertiginidae | ||
? Scolodonta [12] | Indeterminate | A terrestrial gastropod belonging to Scolodontidae | ||
Punctum [12] | P. patagonicum | A terrestrial gastropod belonging to Punctidae | ||
Zilchogyra [12] | Z. miocenica | A terrestrial gastropod belonging to Charopidae | ||
Patagocharopa [12] | P. enigmatica | A terrestrial gastropod of uncertain affinities, possibly a member of Charopidae | ||
Porifera [7] | Unspecified | Spicules | Freshwater sponge | |
Bacillariophyceae [7] | Unspecified | Frustules | Freshwater diatoms | |
Chrysophyceae [7] | Unspecified | Freshwater algae |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Calyptocephalella [13] [14] | C. cf. canqueli | Estancia La Costa, Rinconada de los López | Skull and jaw fragments | A calyptocephalellid frog | |
Neobatrachia [13] | Indeterminate | Estancia La Costa | Skull fragments and presacral vertebrate | A frog originally assigned to "Leptodactylidae", but requires further investigation |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Anisolornis [15] | A. excavatus | A bird of uncertain affinities, authors have varyingly suggested affinities to trumpeters or limpkins | |||
Ankonetta [15] | A. larriestrai | Partial tarsometatarsus | A basal member of Anatidae | ||
Badiostes [15] | B. patagonicus | A member of the family Falconidae | |||
Brontornis [15] | B. burmeisteri | Controversial taxonomic position, either considered a terror bird or an anserimorph | |||
Dryornis [16] [17] | D. pampeanusD. hatcheri | Two partial left humerus | A New World vulture. | ||
Eutelornis [15] | E. patagonicus E. australis | Limb fragments | A member of Anseriformes, relationships of the species to each other or to other Ansiferiformes is uncertain | ||
Liptornis [15] | L. hesternus | A member of Anhingidae | |||
Macranhinga [15] | Indeterminate | A member of Anhingidae | |||
Miocariama [15] [18] | M. santacrucensis | Partial cranium and tibiotarsi fragments | A seriema | ||
Nothurinae [15] | 2 Indeterminate species | Tinamou | |||
Opisthodactylus [15] | O. patagonicus | Limb, vertebra and skull fragments | A member of Rheidae | ||
Patagornis [15] | P. marshi | A terror bird | |||
Phorusrhacos [19] | P. longissimus | Partial skulls | A terror bird | ||
Protibis [15] | P. cnemialis | distal end of tibiotarsus | Potentially a spoonbill | ||
Psilopterus [15] | P. lemoinei P. bachmanni | A terror bird | |||
Thegornis [15] | T. musculosus T. debilis | A member of the family Falconidae | |||
Archaeopsophia [20] | A. aoni | A member of the family Psophiidae (trumpeters) | |||
Chehuenia [20] | C. facongrandei | A roller | |||
Kaikenia [20] | K. mourerchauvirea | An anatid belonging to the subfamily Tadorninae | |||
Peioa [20] | P. australis | A member of Anseriformes. |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Pristidactylus [21] | Indeterminate | La Cueva | Fragmentary jaw bones | Originally assigned to the dubious genus "Erichosaurus" | |
Pleurodonta [13] [21] | |||||
Tupinambis [13] | Monte León | Fragmentary dentaries and maxilla | A teiid lizard | ||
Colubridae [13] [21] | Cerro Observatorio | Partial trunk vertebrae | A snake |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Astrapotherium [22] [23] [24] [25] [26] [2] [27] | A. magnum A. nanum A. sp. | An astrapotheriid |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Adianthus [28] | A. bucatus | An adianthid litoptern | |||
Anisolophus [22] [3] [23] [29] [26] [2] | A. australis A. floweri | A proterotheriid litoptern | |||
Diadiaphorus [30] [3] [23] [2] [29] [31] [27] | D. majusculus D. sanctaecrucis D. sp. | A proterotheriid litoptern | |||
Tetramerorhinus [32] [30] [3] [23] [2] [26] [24] [25] [29] [31] [27] | T. cingulatum T. fleaglei T. mixtum T. lucarius T. prosistens T. sp. | Estancia La Costa Member | A proterotheriid litoptern | ||
Theosodon [33] [30] [3] [23] [34] [2] [26] [24] [25] [29] [31] [27] | T. fontanae T. garretorum T. gracilis T. karaikensis T. lydekkeri T. patagonicum | Estancia La Costa Member | A macraucheniid litoptern | ||
Thoatherium [30] | T. minisculum T. sp. | Estancia La Costa Member | A proterotheriid litoptern |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Adinotherium [3] [23] [35] [26] [24] [27] | A. ovinum A. robustum A. sp. | A toxodontid notoungulate | |||
Cochilius [36] | C. sp. | An interatheriid notoungulate | |||
Hegetotherium [22] [3] [23] [2] [26] [24] [25] [37] | H. mirabile H. sp. | A hegetotheriid notoungulate | |||
Homalodotherium [22] [3] [23] [2] [26] [24] [25] [27] | H. cunninghami | A homalodotheriid notoungulate | |||
Interatherium [38] [22] [3] [23] [35] [2] [26] [24] [25] [27] | I. anguliferum I. brevifrons I. dentatum I. interruptum I. robustum I. rodens I. supernum | An interatheriid notoungulate | |||
Neoicochilus [36] | N. undulatus | An interatheriid notoungulate | |||
Nesodon [22] [3] [23] [35] [2] [26] [24] [25] [27] | N. imbricatus | A toxodontid notoungulate | |||
Notohippus [39] | N. toxodontoides | A notohippid notoungulate | |||
Pachyrukhos [22] [3] [23] [2] [26] [24] [25] [37] | P. moyani | A hegetotheriid notoungulate | |||
Patriarchus [38] [40] | P. palmidens | An interatheriid notoungulate | |||
Protypotherium [33] [38] [22] [3] [23] [2] [26] [24] [25] [40] [27] | P. attenuatum P. australe P. praerutilum P. sp. | An interatheriid notoungulate |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Analcimorphus [41] [42] | A. giganteus A. inversus | Estancia La Costa Member | A basal megatherioid sloth | ||
Analcitherium [43] [41] [42] | A. antarcticum | Estancia La Costa Member | A scelidotheriid sloth | ||
Eucholoeops [44] | E. fronto E. ingens E. litoralis E. titans | Estancia La Costa Member | A megalonychid ground sloth | ||
Hapalops [26] | H. longiceps H. elongatus H. indifferens H. angustipalatus H. platycephalus H. ponderosus H. rectangularis | A ground sloth belonging to Megatherioidea | |||
Hyperleptus [45] | Indeterminate | A megatherioid ground sloth of uncertain affinities | |||
Megalonychotherium | M. atavus | A megalonychid ground sloth | |||
Mylodontidae [45] | Indeterminate | ||||
Nematherium [45] | N. angulatum | A ground sloth belonging to Mylodontoidea | |||
Pelecyodon [45] | P. cristatus | A ground sloth | |||
Planops [45] | P. magnus | A megatheriid ground sloth | |||
Protamandua [45] | P. rothi | An anteater | |||
Prepotherium [45] | P. potens P. filholi | A ground sloth | |||
Megalonychidae | Indeterminate | A ground sloth | |||
Schismotherium [46] [47] | S. fractum | A ground sloth | |||
Xyophorus [47] | X. atlanticus X. latirostris | A nothrotheriid ground sloth |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Anantiosodon [48] | A. rarus | An armadillo | |||
cf. Asterostemma [48] | cf. A. depressa | A glyptodont | |||
Cochlops [48] | C. muricatus | A glyptodont | |||
Eucinepeltus [48] | E. petesatus | A glyptodont | |||
Metopotoxus [48] | M. laevatus | A glyptodont | |||
Parutaetus [48] | P. sp. | An armadillo | |||
Peltephilus [48] | P. ferox P. giganteus P. nanus P. pumilus P. strepens | A horned armadillo | |||
Proeutatus [48] [26] | P. carinatus P. deleo P. lagena P. oenophorus P. robustus | An armadillo | |||
Propalaehoplophorus [48] | P. australis P. incisivusP. minus | A glyptodont | |||
Prozaedyus [48] [26] | P. exilis P. proximus | An armadillo | |||
Stenotatus [26] | S. patagonicus | An armadillo | |||
Stegotherium [48] | S. tauberi S. tessellatum | An armadillo | |||
Vetelia [48] [49] | V. puncta | An armadillo | |||
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Abderites [50] | A. meridionalis | A member of Abderitidae (Paucituberculata) | |||
Acdestis [50] | A. oweniiA. lemairei | A member of Palaeothentidae (Paucituberculata) | |||
Acrocyon [51] | A. sectorius | A borhyaenid sparassodont | |||
Acyon [51] | A. tricuspidatus | A hathliacynid sparassodont | |||
Arctodictis [51] | A. munizi | A borhyaenid sparassodont | |||
Borhyaena [51] | B. tuberata | A borhyaenid sparassodont | |||
Cladosictis [51] | C. patagonia | A sparassodont | |||
Lycopsis [51] | L. torresi | A sparassodont | |||
Microbiotherium [50] | M. acicula M. patagonicum M. gallegosense M. tehuelchum | A member of Microbiotheriidae (Microbiotheria) | |||
Palaeothentes [50] | P. aratae P. minutus P. intermedius P. lemoinei P. pascuali | A member of Palaeothentidae (Paucituberculata) | |||
Perathereutes [51] | P. pungens | A hathliacynid sparassodont | |||
Phonocdromus [50] | P. gracilis | A member of Pichipilidae (Paucituberculata) | |||
Prothylacinus [51] | P. patagonicus | A sparassodont | |||
Pseudonotictis [51] | P. pusillus | A hathliacynid sparassodont | |||
Sipalocyon [51] | S. gracilis S. obusta | A hathliacynid sparassodont | |||
Stilotherium [50] | S. dissimile | A member of Caenolestidae (Paucituberculata) |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Acarechimys [47] | A. minutus A. minutissmus A. constans A. gracilis | A member of Octodontoidea | |||
Acaremys [52] [26] | A. murinus A. messor | A member of Acaremyidae (Octodontoidea) | |||
Adelphomys [52] | A. candidus | A member of Octodontoidea | |||
Dudumus [52] | Indeterminate, potentially new species | A member of Octodontoidea | |||
Eocardia [52] | E. montana "E". excavata "E". fissa | A member of Cavioidea | |||
Neoreomys [52] | N. australis | A member of Cavioidea | |||
Perimys [52] | P. erutus P. onustus P. incavatus | A member of Chinchilloidea | |||
Phanomys [52] | P. mixtus P. vetulus | A member of Cavioidea | |||
Pliolagostomus [52] | P. notatus | A member of Chinchilloidea | |||
Prolagostomus [52] | P. pusilllus | A member of Chinchilloidea | |||
Prospaniomys [52] | Indeterminate, potentially new species | A member of Octodontoidea | |||
Pseudoacaremys [52] | P. kramarzi | A member of the family Acaremyidae (Octodontoidea) | |||
Schistomys [52] | S. erro | A member of Cavioidea | |||
Sciamys [52] | S. principalis S. varians S. latidens | A member of the family Acaremyidae (Octodontoidea) | |||
Scleromys [52] | S. angustus | A member of Chinchilloidea | |||
Spaniomys [52] | S. riparius S. regularis | A member of Octodontoidea | |||
Steiromys [52] | S. dentatus S. duplicatus | A member of Erethizontoidea |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Homunculus [53] [54] [55] | H. patagonicus, H. vizcainoi | A New World monkey | |||
Killikaike [56] | K. blakei | A New World monkey, some authors regard the taxon as a synonym of Homunculuspatagonicus, while others regard them as distinct. |
Name | Species | Locality | Material | Notes | Image |
---|---|---|---|---|---|
Necrolestes [57] | N. patagonicus | Skull and postcranial remains | A mole-like meridiolestidan, youngest known member of the group |
Homunculus is an extinct genus of New World monkey that lived in Patagonia during the Miocene. Two species are known: Homunculus patagonicus and Homunculus vizcainoi, which are known from material found in the Santa Cruz Formation in the far south of Argentina.
Killikaike is an extinct genus of New World monkey. The genus includes one species, Killikaike blakei, that lived in Argentina during the Early Miocene.
Peltephilus, the horned armadillo, is an extinct genus of armadillo xenarthran mammals that first inhabited Argentina during the Oligocene epoch, and became extinct in the Miocene epoch. Notably, the scutes on its head were so developed that they formed horns. Aside from the horned gophers of North America, it is the only known fossorial horned mammal. P. ferox had skull about 11.7 centimetres (4.6 in), and estimated body mass is around 11.07 kilograms (24.4 lb).
Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.
Patagornis is a genus of extinct flightless predatory birds of the family Phorusrhacidae. Known as "terror birds", these lived in what is now Argentina during the Early and Middle Miocene; the Santa Cruz Formation in Patagonia contains numerous specimens. Patagornis was an agile, medium sized Patagornithine and was likely a pursuit predator.
Psilopterus is an extinct genus of phorusrhacid from the Middle Oligocene to possibly the Late Pleistocene of Argentina and Uruguay. Compared to other phorusrhacids, members of the genus are both relatively gracile and diminutive, and include the smallest known species of terror bird: with the head raised P. bachmanni was 70–80 centimeters (2.3–2.6 ft) in height and weighed about 5 kilograms (11 lb), while the largest members of the genus were only about 8 kilograms (18 lb). The birds resemble the modern cariama, except with a heavier build and considerably smaller wings. Fossil finds in Uruguay indicate the genus may have survived until 96,040 ± 6,300 years ago, millions of years after the larger phorusrhacids became extinct.
The Magallanes Basin or Austral Basin is a major sedimentary basin in southern Patagonia. The basin covers a surface of about 170,000 to 200,000 square kilometres and has a NNW-SSE oriented shape. The basin is bounded to the west by the Andes mountains and is separated from the Malvinas Basin to the east by the Río Chico-Dungeness High. The basin evolved from being an extensional back-arc basin in the Mesozoic to being a compressional foreland basin in the Cenozoic. Rocks within the basin are Jurassic in age and include the Cerro Toro Formation. Three ages of the SALMA classification are defined in the basin; the Early Miocene Santacrucian from the Santa Cruz Formation and Friasian from the Río Frías Formation and the Pleistocene Ensenadan from the La Ensenada Formation.
The Santacrucian age is a period of geologic time within the Early Miocene epoch of the Neogene, used more specifically with SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age.
Stegotherium is an extinct genus of long-nosed armadillo, belonging to the Dasypodidae family alongside the nine-banded armadillo. It is currently the only genus recognized as a member of the tribe Stegotheriini. It lived during the Early Miocene of Patagonia and was found in Colhuehuapian rocks from the Sarmiento Formation, Santacrucian rocks from the Santa Cruz Formation, and potentially also in Colloncuran rocks from the Middle Miocene Collón Curá Formation. Its strange, almost toothless and elongated skull indicates a specialization for myrmecophagy, the eating of ants, unique among the order Cingulata, which includes pampatheres, glyptodonts and all the extant species of armadillos.
Hegetotherium is an extinct genus of mammals from the Early to Middle Miocene of South America. Fossils of this genus have been found in the Cerro Bandera, Cerro Boleadoras, Chichinales, Collón Curá, Santa Cruz and Sarmiento Formations of Argentina, the Nazareno Formation of Bolivia, and the Galera and Río Frías Formations of Chile.
Thegornis is an extinct genus of herpetotherine falconid that lived during the Miocene of South America. The genus was erected by Florentino Ameghino in 1895 based on two species, T. musculosus and T. debilis. However, T. debilis was later suggested to be an invalid species, with the differences between it and T. musculosus being due to sexual dimorphism. Two additional species, T. spivacowi and T. sosae, were subsequently named in later years by Federico Agnolín. Its skull and postcranial morphology are similar to the laughing falcon and forest falcon, which together form the clade Herpetotherinae. The seriema Noriegavis's holotype was transferred to this genus and the well-preserved specimen described in 2015 attributed to Noriegavis has been classified into Miocariama.
Proterotherium is an extinct genus of litoptern mammal of the family Proterotheriidae that lived during the Late Miocene of Argentina and Chile. Fossils of this genus have been found in the Ituzaingó Formation of Argentina, and the Galera Formation of Chile.
Anisolophus is an extinct genus of proterotheriid from the Early to Middle Miocene of Argentina. The genus was named by Burmeister in 1885 to accommodate the species Anchitherium australe, which they had named earlier in 1879. Soria then referred the species Licaphrium floweri and Anisolophus minisculus to the genus, making Licaphrium, named in 1887 by Florentino Ameghino, a junior synonym of the genus. Both A. australis and A. floweri are known from the Santacrucian age Santa Cruz Formation, while A. minisculus is known from the Collón Curá Formation.
Prepoplanops is an extinct genus of ground sloth of the family Megatheriidae. It lived in the Miocene around 18 to 16 million years ago of what is now Argentina. The only known species is Prepoplanops boleadorensis.
Eucinepeltus is an extinct genus of glyptodont. It lived during the Early Miocene, and its fossilized remains were discovered in South America.
Cochlops is an extinct genus of glyptodont. It lived from the Early to Middle Miocene, and its fossilized remains have been found in South America.
Analcitherium is an extinct genus of scelidotheriid sloth that lived during the Early Miocene in what is now Argentina. Fossils have been found in the Santa Cruz Formation of Argentina.
Perimys is an extinct genus of neoepiblemid rodent that lived from the Early to Late Miocene in what is now South America. Fossils have been found in the Cerro Bandera, Cerro Boleadoras, Ituzaingó, Santa Cruz, and Sarmiento Formations of Argentina, and the Galera, Santa Cruz and Río Frías Formations of Chile.
Ortotherium is a genus of megalonychid ground sloth from the Late Miocene Ituzaingó Formation of Entre Rios Province, Argentina. Although many species were described, the only valid species of the genus is Ortotherium laticurvatum, with many species being junior synonyms. Ortotherium is known from very fragmentary material, all of which is material from the mandible and teeth. The holotype of O. laticurvatum consists of an incomplete left dentary that had been unearthed from a series of sediments known as ‘Conglomerado osifero’ in Paraná, Argentina. Argentina paleontologist Florentino Ameghino named the species in 1885, though he would go on to name four more, invalid, species of the genus. One species however, O. brevirostrum, has been reclassified as Mesopotamocnus.
Peltephilidae is a family of South American cingulates (armadillos) that lived for over 40 million years, but peaked in diversity towards the end of the Oligocene and beginning of the Miocene in what is now Argentina. They were exclusive to South America due to its geographic isolation at the time, one of many of the continent's strange endemic families. Peltephilids are one of the earliest known cingulates, diverging from the rest of Cingulata in the Early Eocene.
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