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Proeutatus Temporal range: Early Miocene ~ | |
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Reconstructed skeleton of Proeutatus | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Cingulata |
Family: | Chlamyphoridae |
Genus: | † Proeutatus Ameghino, 1891 |
Type species | |
†Proeutatus oenophorum Ameghino, 1887 | |
Species | |
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Synonyms | |
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Proeutatus is an extinct genus of xenarthran, belonging to the order Cingulata. It lived during the Early Miocene, and its fossilized remains were discovered in South America.
This animal was vaguely similar to the extant six-banded armadillo, and was similarly sized. The skull was approximately 12 centimeters long, the whole animal not exceeding more than half a meter in length for a weight between 5 and 6 kilograms. Proeutatus had a quite sturdy body, a high reared skull with an elongated snout and a sturdy set of teeth compared to those of many other armadillos.
The skull of Proeutatus had some characteristics distinguishing it from the more similar forms, such as Stenotatus and Eutatus ; the premaxilla had an upper tooth, the zygomatic arch was expanded and had a suborbital process ; the mandible was more robust, with an almost vertically ascending branch and a higher coronoid process. The general shape of the skull was, in lateral view, concave, while it was straight and downwards tending for Eutatus. A sagittal crest was present, suggesting the existence of powerful masticatory muscles. The dentition was very robust, and the teeth possessed a posterolabial lobe, indicating a particular chewing movement.
The first remains of Proeutatus were found in Argentina and described in 1887 by Florentino Ameghino, who attributed them to the genus Eutatus. In 1891, Ameghino created the genus Proeutatus for those remains ; the two known species are Proeutatus oenophorus, the type species, and P. postpuntum.
Proeutatus was historically considered to be a member of the group Eutatinae, composed of medium to large armadillos specialized in an herbivorous diet. More recent researches indicates that it was a basal member of the family Chlamyphoridae, close to the origin of the glyptodonts and the Pampatheres.
The structure of its jaws and teeth indicates that it had an omnivorous diet, with notable specializations for the consumption of vegetal matters. Proeutatus was among the armadillos the most specialized towards an herbivorous diet.
Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. Dasypodids and chlamyphorids, the armadillos, are the only surviving families in the order. Two groups of cingulates much larger than extant armadillos existed until recently: pampatheriids, which reached weights of up to 200 kg (440 lb) and chlamyphorid glyptodonts, which attained masses of 2,000 kg (4,400 lb) or more.
Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.
Interatherium is an extinct genus of interatheriid mammal from the Early to Middle Miocene (Colhuehuapian-Mayoan). Fossils have been found in the Santa Cruz, Collón Curá and Sarmiento Formations in Argentina.
Glyptodon is a genus of glyptodont that lived from the Pleistocene, around 2.5 million years ago, to the Early Holocene, around 11,000 years ago, in Argentina, Uruguay, Paraguay, Bolivia, Peru, Brazil, and Colombia. It was the first named extinct cingulate and is the type genus of Glyptodontinae, and, or, Glyptodontinae. Many species have been named for the genus, though few are considered valid, and it is one of, if not the, best known genus of glyptodont. Hundreds of specimens have been referred to the genus, but the holotype, or name specimen, of the type species, G. clavipes, was described in 1839 by notable British paleontologist Sir Richard Owen. The holotype used by Owen is a chimera of fossils from 3 different localities, including a molariform used for the name of the genus that actually belongs to Panochthus, making it a species inquirenda.
Holmesina is a genus of pampathere, an extinct group of armadillo-like creatures that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell was made up of flexible plates which allowed the animal to move more easily. Holmesina species were herbivores that grazed on coarse vegetation; armadillos are mostly insectivorous or omnivorous.
Stegotherium is an extinct genus of long-nosed armadillo, belonging to the Dasypodidae family alongside the nine-banded armadillo. It is currently the only genus recognized as a member of the tribe Stegotheriini. It lived during the Early Miocene of Patagonia and was found in Colhuehuapian rocks from the Sarmiento Formation, Santacrucian rocks from the Santa Cruz Formation, and potentially also in Colloncuran rocks from the Middle Miocene Collón Curá Formation. Its strange, almost toothless and elongated skull indicates a specialization for myrmecophagy, the eating of ants, unique among the order Cingulata, which includes pampatheres, glyptodonts and all the extant species of armadillos.
Necrolestes is an extinct genus of mammals, which lived during the Early Miocene in what is now Argentine Patagonia. It is the most recent known genus of Meridiolestida, an extinct group of mammals more closely related to therians than to monotremes that dominated South America during the Late Cretaceous. It contains two species, N. patagonensis and N. mirabilis, The type species N. patagonensis was named by Florentino Ameghino in 1891 based on remains found by his brother, Carlos Ameghino in Patagonia. Fossils of Necrolestes have been found in the Sarmiento and Santa Cruz Formations. Its morphology suggests that is was a digging, subterranean dwelling mole-like mammal that fed on invertebrates.
Utaetus is an extinct genus of mammal in the order Cingulata, related to the modern armadillos. The genus contains a single species, Utaetus buccatus. It lived in the Late Paleocene to Late Eocene and its fossil remains were found in Argentina and Brazil in South America.
Epipeltephilus is an extinct genus of armadillo, belonging to the family Peltephilidae, the "horned armadillos", whose most famous relative was Peltephilus. Epipeltephilus is the last known member of its family, becoming extinct during the Chasicoan period. It was found in the Rio Mayo Formation and the Arroyo Chasicó Formation of Argentina, and in northern Chile.
Prosotherium is an extinct genus of hegetotheriid notoungulate. It lived during the Late Oligocene, and its fossilized remains were found in South America.
Astrapothericulus is an extinct genus of mammals, belonging to the order Astrapotheria. It lived during the Lower Miocene in what is now South America.
Eomorphippus is an extinct genus of notohippid notoungulate that lived from the Late Eocene to the Early Oligocene in what is today South America.
Morphippus is an extinct genus of notohippid notoungulate that lived during the Middle to Late Oligocene in what is now South America.
Stenotatus is an extinct genus of cingulate, belonging to the family Dasypodidae. It lived from the Early to the Late Miocene in South America.
Proeuphractus is an extinct genus of xenarthran, related to the modern armadillos. It lived from the Early to the Late Miocene, and its fossilized remains were discovered in South America.
Eucinepeltus is an extinct genus of Glyptodont. It lived during the Early Miocene, and its fossilized remains were discovered in South America.
Glyptatelus is an extinct genus of glyptodont. It lived from the Late Eocene to the Middle Oligocene in what is now Argentina and Bolivia.
Cochlops is an extinct genus of glyptodont. It lived from the Early to Middle Miocene, and its fossilized remains have been found in South America.
Australohyaena is an extinct genus of carnivorous mammal, belonging to the order Sparassodonta. It lived during the Late Oligocene, and its fossilized remains were discovered in Argentina.
Arminiheringia is an extinct genus of sparassodont. It lived during the Early Eocene in South America.