| Pampatherium Temporal range: | |
|---|---|
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Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Cingulata |
| Family: | † Pampatheriidae |
| Genus: | † Pampatherium Ameghino, 1875 [1] |
| Species | |
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Pampatherium is an extinct genus of xenarthran that lived in the Americas during the Pleistocene. Some species went extinct right at the Pleistocene-Holocene border.
Pampatherium humboldtii and P. typum lived in South America (mostly Brazil) during the Pleistocene, with P. humboldtii surviving into the very Late Pleistocene. [2] [3] [4]
Pampatherium mexicanum was the only North American species, reaching as far north as Sonora, Mexico. It lived during the Rancholabrean. [5]
Pampatherium resembled a very large armadillo. One species, P. humboldtii, weighed up to 209.5 kg (462 lb). [6] Pampatheres generally resembled armadillos, particularly in the shape of it skull, long snout, and the presence of three areas on the carapace (movable bands, scapular and pelvic shields). Among the features that distinguish them from armadillos are their posterior teeth, which are bilobate rather than peg-like. Their endocranial morphology is also similar to glyptodonts. [6]
The osteoderms of Pampatherium have little ornamentation, lack a depressed marginal band, and those from the posterior buckler are mostly rectangular. [7]
Pampatherium is thought to have excavated paleoburrows and would have both fought with and have been preyed upon by Arctotherium angustidens in these dens. [8]
Xenarthra is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 Mya, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene.
Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. Dasypodids and chlamyphorids, the armadillos, are the only surviving families in the order. Two groups of cingulates much larger than extant armadillos existed until recently: pampatheriids, which reached weights of up to 200 kg (440 lb) and chlamyphorid glyptodonts, which attained masses of 2,000 kg (4,400 lb) or more.
Doedicurus, or Dædicurus, is an extinct genus of glyptodont from South America containing one species, D. clavicaudatus. Glyptodonts are a member of the family Chlamyphoridae, which also includes some modern armadillo species, and they are classified in the superorder Xenarthra alongside sloths and anteaters. Being a glyptodont, it was a rotund animal with heavy armor and a carapace. Averaging at an approximate 1,400 kg (3,100 lb), it was one of the largest glyptodonts to have ever lived. Though glyptodonts were quadrupeds, large ones like Doedicurus may have been able to stand on two legs like other xenarthrans. It notably sported a spiked tail club, which may have weighed 40 or 65 kg in life, and it may have swung this in defense against predators or in fights with other Doedicurus at speeds of perhaps 11 m/s.
Glyptotherium is a genus of glyptodont that lived from the Early Pliocene, about 4.9 million years ago, to the Early Holocene, around 7,000 years ago, in the United States, Mexico, Guatemala, Costa Rica, Honduras, El Salvador, Panama, Venezuela, and Brazil. The genus was first described in 1903 by American paleontologist Henry Fairfield Osborn with the type species being, G. texanum, based on fossils that had been found in the Pliocene Blancan Beds in Llano Estacado, Texas, USA. The genus has since been discovered in many more fossil sites. Another species, G. cylindricum, was named in 1912 by fossil hunter Barnum Brown on the basis of a partial carapace, teeth, and several additional fossils that had been unearthed from the Pleistocene deposits in Jalisco, Mexico.
Pampatheriidae is an extinct family of large plantigrade armored armadillos related to extant armadillos in the order Cingulata. However, pampatheriids have existed as a separate lineage since at least the middle Eocene Mustersan age, 45 to 48 million years ago. Pampatheres evolved in South America during its long period of Cenozoic isolation. Although widespread, they were less diverse and abundant than the armadillos. Holmesina spread to North America after the formation of the Isthmus of Panama as part of the Great American Interchange. They finally disappeared on both continents in the end-Pleistocene extinctions, about 12,000 years ago.
Panochthus is an extinct genus of glyptodont, which lived in the Gran Chaco-Pampean region of Argentina, Brazil, Bolivia, Paraguay and Uruguay during the Pleistocene epoch.
Glyptodon is a genus of glyptodont that lived from the Pleistocene, around 2.5 million years ago, to the Early Holocene, around 11,000 years ago, in Argentina, Uruguay, Paraguay, Bolivia, Peru, Brazil, and Colombia. It was the first named extinct cingulate and is the type genus of Glyptodontinae, and, or, Glyptodontinae. Many species have been named for the genus, though few are considered valid, and it is one of, if not the, best known genus of glyptodont. Hundreds of specimens have been referred to the genus, but the holotype, or name specimen, of the type species, G. clavipes, was described in 1839 by notable British paleontologist Sir Richard Owen. It was roughly the same size and weight as a Volkswagen Beetle, 800–840 kg (1,760–1,850 lb). With its rounded, bony shell and squat limbs, it superficially resembled a turtle, and the much earlier dinosaurian ankylosaur – providing an example of the convergent evolution of unrelated lineages into similar forms. In 2016 an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae. For this reason, glyptodonts and all armadillos but Dasypus were relocated to a new family, Chlamyphoridae, and glyptodonts were demoted from the former family Glyptodontidae to a subfamily.
Holmesina is a genus of pampathere, an extinct group of armadillo-like creatures that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell was made up of flexible plates which allowed the animal to move more easily. Holmesina species were herbivores that grazed on coarse vegetation; armadillos are mostly insectivorous or omnivorous.
Catonyx is an extinct genus of ground sloth of the family Scelidotheriidae, endemic to South America during the Pliocene and Pleistocene epochs. It lived from 2.5 Ma to about 10,000 years ago, existing for approximately 2.49 million years. The most recent date obtained is about 9600 B.P.
Arctotherium is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. Arctotherium migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama during the late Pliocene. The genus consists of one early giant form, A. angustidens, and several succeeding smaller species, which were within the size range of modern bears. Arctotherium was adapted to open and mixed habitat. They are genetically closer to the spectacled bear, than to Arctodus of North America, implying the two extinct forms evolved large size in a convergent manner.
Chlamyphoridae is a family of cingulate mammals. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, there had been speculation that the extant family Dasypodidae could be paraphyletic based on morphological evidence. In 2016, an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae. For this reason, all extant armadillos but Dasypus were relocated to a new family.
Macroeuphractus is a genus of extinct armadillos from the Late Miocene to Late Pliocene of South America. The genus is noted for its large size, with Macroeuphractus outesi being the largest non-pampathere or glyptodont armadillo discovered, as well as its specializations for carnivory, unique among all xenarthrans.
Valgipes is an extinct genus of scelidotheriid ground sloth, endemic to intertropical Brazil and Uruguay during the Late Pleistocene. Thought to have been a forest-dwelling browser, Valgipes is a monotypic genus with a complex and long taxonomic history, and is a close relative of Catonyx and Proscelidodon.
Cullinia is an extinct genus of litoptern, an order of South American native ungulates that included horse-like and camel-like animals such as Macrauchenia. It is only known from fragmentary remains. Cullinia levis is known from Chasicoan remains found in the Arroyo Chasicó Formation of Argentina, and remains from the Brazilian state of Acre and the Huayquerian Ituzaingó Formation have been assigned to Cullinia sp..
Kraglievichia is an extinct genus of cingulate belonging to the family Pampatheriidae. It lived from the Late Miocene to the Early Pliocene, and its fossilized remains were discovered in South America.
Scirrotherium is an extinct genus of pampatheres, a family of herbivorous cingulates, related to the similar but smaller modern armadillos, and with the now extinct glyptodonts, well-known from their shell-like armor. Its scientific name is derived from the Greek prefix "skiros-", "cover", and the suffix "-therion, "beast", while the name of the type species, hondaensis, honors the town of Honda, in the Tolima Department of Colombia. Scirrotherium is one of several genera of xenarthrans found in the La Venta fauna, dated from the Middle Miocene.
Neoglyptatelus is an extinct genus of xenarthran, belonging to the order Cingulata. It lived from the Middle to the Late Miocene, and its fossilized remains are found in South America.
Proeutatus is an extinct genus of xenarthran, belonging to the order Cingulata. It lived during the Early Miocene, and its fossilized remains were discovered in South America.
Vassallia is an extinct genus of cingulate belonging to the family Pampatheriidae. It lived between the Middle Oligocene and the Early Pliocene in what is now South America.
Dasypus neogaeus is an extinct species of armadillo, belonging to the genus Dasypus, alongside the modern nine-banded armadillo. The only known fossil is a single osteoderm, though it has been lost, that was found in the Late Miocene strata of Argentina.
