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Pampatheriidae Temporal range: | |
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Holmesina occidentalis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Cingulata |
Family: | † Pampatheriidae Paula Couto 1954 |
Genera | |
Pampatheriidae ("Pampas beasts") is an extinct family of large cingulates related to armadillos. They first appeared in South America during the mid-Miocene, and Holmesina and Pampatherium spread to North America during the Pleistocene after the formation of the Isthmus of Panama as part of the Great American Interchange. They became extinct as part of the end-Pleistocene extinction event, about 12,000 years ago. [2]
The placement of the Eocene genus Machlydotherium in the family is considered doubtful. The oldest undoubted member of the group is Scirrotherium from La Venta, Colombia, dating to the mid-Miocene. [2] Analysis of ear morphology suggests that they are most closely related to the much larger glyptodonts, which genetic evidence indicates is nested with modern armadillos as part of the family Chlamyphoridae, which by extension also places pampatheres within this group. [3]
Phylogeny after Tambusso et al. (2021): [4]
Cingulata |
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Pampatheres are believed to have attained a weight of up to 200 kg (440 lb). Like three-banded armadillos, and unlike glyptodonts, their armored shell was given some flexibility by three movable lateral bands of scutes. [5] The osteoderms (bony plates in the skin comprising the armor) of pampatheres were each covered by a single keratinized scute, unlike osteoderms of armadillos, which have more than one scute. [5]
A study of pampathere jaw biomechanics showed that their masticatory musculature was more powerful and more adapted for transverse movements than that of armadillos, leading to the conclusion that much of their diet was coarse vegetation. They are thought to have been primarily grazers, unlike armadillos, which are omnivorous or insectivorous. [5] The variation between species in the expression of adaptations for grinding coarse vegetation correlates with the aridity of their habitat; such adaptations are most pronounced in Pampatherium typum , which lived in the arid Pampas, and least pronounced in Holmesina occidentalis , which lived in humid lowlands. [6]
Pampatheres were widely distributed across South America during the Pleistocene ranging from northern Argentina, eastwards to northeastern Brazil, and westwards to Colombia and Peru. [7] As part of the Great American Interchange, pampatheres dispersed northwards into Central and North America, with members of the genus Holmesina reaching the United States, as far north as Kansas, and as far east as Florida and South Carolina. [8]
Glyptodonts are an extinct clade of large, heavily armoured armadillos, reaching up to 1.5 metres (4.9 ft) in height, and maximum body masses of around 2 tonnes. They had short, deep skulls, a fused vertebral column, and a large bony carapace made up of hundreds of individual scutes. Some glyptodonts had clubbed tails, similar to ankylosaurid dinosaurs.
Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. Dasypodids and chlamyphorids, the armadillos, are the only surviving families in the order. Two groups of cingulates much larger than extant armadillos existed until recently: pampatheriids, which reached weights of up to 200 kg (440 lb) and chlamyphorid glyptodonts, which attained masses of 2,000 kg (4,400 lb) or more.
Peltephilus, the horned armadillo, is an extinct genus of armadillo xenarthran mammals that first inhabited Argentina during the Oligocene epoch, and became extinct in the Miocene epoch. Notably, the scutes on its head were so developed that they formed horns. Aside from the horned gophers of North America, it is the only known fossorial horned mammal. P. ferox had skull about 11.7 centimetres (4.6 in), and estimated body mass is around 11.07 kilograms (24.4 lb).
Glyptotherium is a genus of glyptodont in the family Chlamyphoridae that lived from the Early Pliocene, about 3.6 million years ago, to the Late Pleistocene, around 15,000 years ago. It had a wide distribution, living in the United States, Mexico, Guatemala, Costa Rica, Honduras, El Salvador, Panama, Venezuela, and Brazil. The genus was first described in 1903 by American paleontologist Henry Fairfield Osborn with the type species being, G. texanum, based on fossils that had been found in the Pliocene Blancan Beds in Llano Estacado, Texas, USA. Glyptotherium fossils have since been unearthed from many more fossil sites, from Florida to Colombia. Another species, G. cylindricum, was named in 1912 by fossil hunter Barnum Brown on the basis of a partial skeleton that had been unearthed from the Pleistocene deposits in Jalisco, Mexico. The two species differ in several aspects, including age, with G. texanum being from the older Early Pliocene to Early Pleistocene strata, whereas G. cylindricum is exclusive to the Late Pleistocene.
Scalabrinitherium is an extinct genus of mammals of the family Macraucheniidae. Fossils of this animal were found among the fossils of prehistoric xenarthrans in the Ituzaingó Formation of Argentina.
Glyptodon is a genus of glyptodont, an extinct group of large, herbivorous armadillos, that lived from the Pliocene, around 3.2 million years ago, to the early Holocene, around 11,000 years ago, in South America. It is one of, if not the, best known genus of glyptodont. Glyptodon has a long and storied past, being the first named extinct cingulate and the type genus of the subfamily Glyptodontinae. Fossils of Glyptodon have been recorded as early as 1814 from Pleistocene aged deposits from Uruguay, though many were incorrectly referred to the ground sloth Megatherium by early paleontologists.
Holmesina is an extinct genus of pampathere, an extinct group of armadillo-like xenarthrans that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell was made up of flexible plates which allowed the animal to move more easily.
Stegotherium is an extinct genus of long-nosed armadillo, belonging to the Dasypodidae family alongside the nine-banded armadillo. It is currently the only genus recognized as a member of the tribe Stegotheriini. It lived during the Early Miocene of Patagonia and was found in Colhuehuapian rocks from the Sarmiento Formation, Santacrucian rocks from the Santa Cruz Formation, and potentially also in Colloncuran rocks from the Middle Miocene Collón Curá Formation. Its strange, almost toothless and elongated skull indicates a specialization for myrmecophagy, the eating of ants, unique among the order Cingulata, which includes pampatheres, glyptodonts and all the extant species of armadillos.
Neosclerocalyptus was an extinct genus of glyptodont that lived during the Pliocene, Pleistocene, and Holocene of Southern South America, mostly Argentina. It was small compared to many glyptodonts at only around 2 meters long and 360 kilograms.
Pampatherium is an extinct genus of xenarthran that lived in the Americas during the Pleistocene. Some species went extinct right at the Pleistocene-Holocene border.
Tonnicinctus is an extinct species of pampatheriid that lived in Argentina during the Pleistocene and Holocene.
Proeremotherium is an extinct genus of megatheriine ground sloths in the family Megatheriidae. It lived during the Late Miocene and Early Pliocene of what is now Venezuela. So far, two largely complete skulls have been recovered in the Falcón Basin in Venezuela. The finds identify the animals as medium-sized representatives of the Megatheriidae. In the cranial anatomy, Proeremotherium resembles the later and giant Eremotherium. It is therefore assumed that the two ground sloths are directly related to each other.
Kraglievichia is an extinct genus of cingulate belonging to the family Pampatheriidae. It lived from the Late Miocene to the Early Pliocene, and its fossilized remains were discovered in South America.
Scirrotherium is an extinct genus of pampatheres, a family of herbivorous cingulates, related to the similar but smaller modern armadillos, and with the now extinct glyptodonts, well-known from their shell-like armor. Its scientific name is derived from the Greek prefix "skiros-", "cover", and the suffix "-therion, "beast", while the name of the type species, hondaensis, honors the town of Honda, in the Tolima Department of Colombia. Scirrotherium is one of several genera of xenarthrans found in the La Venta fauna, dated from the Middle Miocene.
Neoglyptatelus is an extinct genus of xenarthran, belonging to the order Cingulata. It lived from the Middle to the Late Miocene, and its fossilized remains are found in South America.
Proeuphractus is an extinct genus of xenarthran, related to the modern armadillos. It lived from the Early to the Late Miocene, and its fossilized remains were discovered in South America.
Vassallia is an extinct genus of cingulate belonging to the family Pampatheriidae. It lived between the Middle Oligocene and the Early Pliocene in what is now South America.
Dasypus neogaeus is an extinct species of armadillo, belonging to the genus Dasypus, alongside the modern nine-banded armadillo. The only known fossil is a single osteoderm, though it has been lost, that was found in the Late Miocene strata of Argentina.
Caiman australis is an extinct species of caiman described in 1858 on the basis of a left maxilla that was collected from the Upper Miocene age Ituzaingó Formation of Entre Rios, Argentina.
Peltephilidae is a family of South American cingulates (armadillos) that lived for over 40 million years, but peaked in diversity towards the end of the Oligocene and beginning of the Miocene in what is now Argentina. They were exclusive to South America due to its geographic isolation at the time, one of many of the continent's strange endemic families. Peltephilids are one of the earliest known cingulates, diverging from the rest of Cingulata in the Early Eocene.