Boreostemma | |
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Tail of Boreostemma from the Honda Group | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Cingulata |
Family: | Chlamyphoridae |
Subfamily: | † Glyptodontinae |
Genus: | † Boreostemma Carlini et al. 2008 |
Type species | |
†Boreostemma pliocena Carlini et al. 2008 | |
Species | |
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Boreostemma is an extinct genus of glyptodonts from northern South America. Fossils assigned to the genus were first described as belonging to Asterostemma from southern South America, but have been placed in the new genus Boreostemma by Carlini et al. in 2008. The type species is B. pliocena. Fossils of Boreostemma have been found in the Honda Group of Colombia, in Peru and Venezuela.
The genus name Boreostemma is a combination of stemma, taken from Asterostemma , and boreo is derived from borealis, meaning "northern", to distinguish the northern South American genus from the southern Asterostemma. [1]
Boreostemma has been placed as closely related to the later Glyptodon and Glyptotherium . Cladogram after Barasoain et al. 2022: [2]
Glyptodonts |
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Four species have been described in the genus Boreostemma. [3] Unspecified Boreostemma fossils were found in the Ipururo Formation of the department of Ucayali, Amazonian Peru. [4] [5] [6] Asterostemma contains the type species A. depressa and several nomina dubia. The genus has been found in the Early Miocene (Santacrucian in the SALMA classification) Santa Cruz Formation of the province of Santa Cruz, Argentina and in the Solimões Formation in the state of Acre, Amazonian Brazil. [7] [8]
The type species B. pliocena was described in 2008 by Carlini et al. based on 30 osteoderms palate fragment with M6-7 alveoli. [9] Fossils have been described from the type locality along the road to Tío Gregorio in the Codore Formation at the Lagerstätte of Urumaco. [10]
B. gigantea was described as Asterostemma gigantea in 1997 by Carlini et al. based on a partial skeleton. [11] Fossils have been recovered at the Duke Localities 32, 108, 113, 114 and 121W in the La Victoria Formation of the Honda Group at the Konzentrat-Lagerstätte La Venta. [12] [13] [14] [15] [16]
B. acostae was described as Asterostemma acostae in 1983 by Carlos Villarroel based on a carapace fragment with 44 plates. [17] Fossils have been found in the Cerbatana Member, and the Chunchullo Sandstone of the La Victoria Formation, of the Honda Group at the Konzentrat-Lagerstätte La Venta. [18] [19]
B. venezolensis was described as Asterostemma venezolensis in 1947 by George Gaylord Simpson, [20] based on most of the carapace and the anterior end of the lower jaw, including teeth. [21] Fossils have been uncovered from the Santa Inés Formation along the banks of the Güere River in the state of Anzoátegui in Venezuela. [22]
Glyptodonts are an extinct clade of large, heavily armoured armadillos, reaching up to 1.5 metres (4.9 ft) in height, and maximum body masses of around 2 tonnes. They had short, deep skulls, a fused vertebral column, and a large bony carapace made up of hundreds of individual scutes. Some glyptodonts had clubbed tails, similar to ankylosaurid dinosaurs.
Doedicurus is an extinct genus of glyptodont from South America containing one species, D. clavicaudatus. Glyptodonts are a member of the family Chlamyphoridae, which also includes some modern armadillo species, and they are classified in the superorder Xenarthra alongside sloths and anteaters. Being a glyptodont, it was a rotund animal with heavy armor and a carapace. Averaging at an approximate 1,400 kg (3,100 lb), it was one of the largest glyptodonts to have ever lived. Though glyptodonts were quadrupeds, large ones like Doedicurus may have been able to stand on two legs like other xenarthrans. It notably sported a spiked tail club, which may have weighed 40 or 65 kg in life, and it may have swung this in defense against predators or in fights with other Doedicurus at speeds of perhaps 11 m/s.
Glyptotherium is a genus of glyptodont in the family Chlamyphoridae that lived from the Early Pliocene, about 3.6 million years ago, to the Late Pleistocene, around 15,000 years ago. It had a wide distribution, living in the United States, Mexico, Guatemala, Costa Rica, Honduras, El Salvador, Panama, Venezuela, and Brazil. The genus was first described in 1903 by American paleontologist Henry Fairfield Osborn with the type species being, G. texanum, based on fossils that had been found in the Pliocene Blancan Beds in Llano Estacado, Texas, USA. Glyptotherium fossils have since been unearthed from many more fossil sites, from Florida to Colombia. Another species, G. cylindricum, was named in 1912 by fossil hunter Barnum Brown on the basis of a partial skeleton that had been unearthed from the Pleistocene deposits in Jalisco, Mexico. The two species differ in several aspects, including age, with G. texanum being from the older Early Pliocene to Early Pleistocene strata, whereas G. cylindricum is exclusive to the Late Pleistocene.
Pampatheriidae is an extinct family of large cingulates related to armadillos. They first appeared in South America during the mid-Miocene, and Holmesina and Pampatherium spread to North America during the Pleistocene after the formation of the Isthmus of Panama as part of the Great American Interchange. They became extinct as part of the end-Pleistocene extinction event, about 12,000 years ago.
Hoplophorus is an extinct genus of glyptodont, a subfamily of armadillos. The only confidently known species was H. euphractus, found in Pleistocene deposits in Brazil, though fossils possibly from another species are known from Bolivia.
Panochthus is an extinct genus of glyptodont, which lived in the Gran Chaco-Pampean region of Argentina, Brazil, Bolivia, Paraguay and Uruguay during the Pleistocene epoch.
Glyptodon is a genus of glyptodont, an extinct group of large, herbivorous armadillos, that lived from the Pliocene, around 3.2 million years ago, to the early Holocene, around 11,000 years ago, in South America. It is one of, if not the, best known genus of glyptodont. Glyptodon has a long and storied past, being the first named extinct cingulate and the type genus of the subfamily Glyptodontinae. Fossils of Glyptodon have been recorded as early as 1814 from Pleistocene aged deposits from Uruguay, though many were incorrectly referred to the ground sloth Megatherium by early paleontologists.
Propalaehoplophorus, also written as Propalaeohoplophorus, is an extinct genus of glyptodont, which lived in South America during the Early Miocene epoch.
Stegotherium is an extinct genus of long-nosed armadillo, belonging to the Dasypodidae family alongside the nine-banded armadillo. It is currently the only genus recognized as a member of the tribe Stegotheriini. It lived during the Early Miocene of Patagonia and was found in Colhuehuapian rocks from the Sarmiento Formation, Santacrucian rocks from the Santa Cruz Formation, and potentially also in Colloncuran rocks from the Middle Miocene Collón Curá Formation. Its strange, almost toothless and elongated skull indicates a specialization for myrmecophagy, the eating of ants, unique among the order Cingulata, which includes pampatheres, glyptodonts and all the extant species of armadillos.
Eleutherocercus was a genus of glyptodonts that lived during the Late Miocene and Early Pliocene in South America. Fossils of the genus have been found in the Huayquerian Ituzaingó Formation and the Montehermosan Monte Hermoso Formation in Argentina.
Neosclerocalyptus was an extinct genus of glyptodont that lived during the Pliocene, Pleistocene, and Holocene of Southern South America, mostly Argentina. It was small compared to many glyptodonts at only around 2 meters long and 360 kilograms.
Lomaphorus is a possibly dubious extinct genus of glyptodont that lived during the Pleistocene in eastern Argentina. Although many species have been referred, the genus itself is possibly dubious or synonymous with other glyptodonts like Neoslerocalyptus from the same region.
Kelenkura is an extinct genus of heavily armored mammals belonging to the subfamily Glyptodontinae, from the family Chlamyphoridae that contain most of the modern armadillos. It was a medium-sized South American animal, distantly related to Doedicurus. Fossils of this genus were recovered in the Arroyo Chasicó Formation and in the Loma de Las Tapias Formation of Argentina in rocks dating back to the Late Miocene epoch.
Proeuphractus is an extinct genus of xenarthran, related to the modern armadillos. It lived from the Early to the Late Miocene, and its fossilized remains were discovered in South America.
Palaehoplophorus is an extinct genus of glyptodont. It lived from the Middle to the Late Miocene, and its fossilized remains were discovered in South America.
Neuryurus is an extinct genus of glyptodont. It lived from the Late Pliocene to the Early Holocene, and its fossilized remains were discovered in South America.
Comaphorus is a dubious extinct genus of glyptodont. It lived during the Late Miocene in Argentina, but only one fossil has ever been referred to the animal.
Asterostemma is an extinct genus of glyptodont. It lived during the Middle Miocene, and its fossilized remains were discovered in South America.
Cochlops is an extinct genus of glyptodont. It lived from the Early to Middle Miocene, and its fossilized remains have been found in South America.
Peltephilidae is a family of South American cingulates (armadillos) that lived for over 40 million years, but peaked in diversity towards the end of the Oligocene and beginning of the Miocene in what is now Argentina. They were exclusive to South America due to its geographic isolation at the time, one of many of the continent's strange endemic families. Peltephilids are one of the earliest known cingulates, diverging from the rest of Cingulata in the Early Eocene.