The Knife Tooth Cat (Machairodus) is a genus of large machairodont or ''saber-toothed cat'' that lived in Africa, Eurasia and North America during the late Miocene. It is the animal from which the subfamily Machairodontinae gets its name and has since become a wastebasket taxon over the years as many genera of sabertooth cat have been and are still occasionally lumped into it.
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. It is one of two groups placed in the clade Theria alongside Eutheria, which contains the placentals.
A saber-tooth is any member of various extinct groups of predatory therapsids, predominantly carnivoran mammals, that are characterized by long, curved saber-shaped canine teeth which protruded from the mouth when closed. Saber-toothed mammals have been found almost worldwide from the Eocene epoch to the end of the Pleistocene epoch.
Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae. They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million until about 10,000 years ago.
Creodonta is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the Oxyaenids and the Hyenodonts, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa.
Sparassodonta is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has showed that sparassodonts died out long before eutherian carnivores arrived in South America. Sparassodonts have been referred to as borhyaenoids by some authors, but currently the term Borhyaenoidea refers to a restricted subgroup of sparassodonts comprising borhyaenids and their close relatives.
Thylacosmilus is an extinct genus of saber-toothed metatherian mammals that inhabited South America from the Late Miocene to Pliocene epochs. Though Thylacosmilus looks similar to the "saber-toothed cats", it was not a felid, like the well-known North American Smilodon, but a sparassodont, a group closely related to marsupials, and only superficially resembled other saber-toothed mammals due to convergent evolution. A 2005 study found that the bite forces of Thylacosmilus and Smilodon were low, which indicates the killing-techniques of saber-toothed animals differed from those of extant species. Remains of Thylacosmilus have been found primarily in Catamarca, Entre Ríos, and La Pampa Provinces in northern Argentina.
Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified fourth upper premolar and the first lower molar. These teeth are also referred to as sectorial teeth.
Borhyaenidae is an extinct metatherian family of low-slung, heavily built predatory mammals in the order Sparassodonta. Borhyaenids are not true marsupials, but members of a sister taxon, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful jaws, like those of the unrelated placentalians Hyaenodon and Andrewsarchus, for crushing bones. Borhyaenids grew up to an average of 5 to 6 feet long.
Durophagy is the eating behavior of animals that consume hard-shelled or exoskeleton-bearing organisms, such as corals, shelled mollusks, or crabs. It is mostly used to describe fish, but is also used when describing reptiles, including fossil turtles, placodonts and invertebrates, as well as "bone-crushing" mammalian carnivores such as hyenas. Durophagy requires special adaptions, such as blunt, strong teeth and a heavy jaw. Bite force is necessary to overcome the physical constraints of consuming more durable prey and gain a competitive advantage over other organisms by gaining access to more diverse or exclusive food resources earlier in life. Those with greater bite forces require less time to consume certain prey items as a greater bite force can increase the net rate of energy intake when foraging and enhance fitness in durophagous species.
Dukecynus is an extinct genus of meat-eating metatherian belonging to the order Sparassodonta, which lived in South America during the Middle Miocene (Laventan), between about 13.8 and 11.8 million years ago. The name of the genus meaning "Duke dog", for Duke University and the Greek word cynos, dog, for the pretended similarity of this animal with dogs. A single species known so far, Dukecynus magnus. The species name "magnus" derives from Latin for big, to reflect their great size.
Proborhyaenidae is an extinct family of metatherian mammals of the order Sparassodonta, which lived in South America from the Eocene (Mustersan) until the Oligocene (Deseadan). Sometimes it has been included as a subfamily of their relatives, the borhyaenids. The largest species, Proborhyaena gigantea, is estimated to be about the size of a spectacled bear, with its skull reaching 60 cm (2.0 ft) in length, and body mass estimates up to approximately 90–200 kg (200–440 lb), making the proborhyaenids some of the largest known metatherians. Proborhyaenid remains have been found in western Bolivia, Uruguay, southern Brazil, and the provinces of Mendoza, Salta, and Chubut, in Argentina.
Thylacosmilidae is an extinct family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like Barbourofelis and Smilodon.
Patagosmilus is an extinct genus of meat-eating metatherian mammal of the family Thylacosmilidae, that lived in the Middle Miocene in South America. Like other representatives of this family, such as Thylacosmilus atrox and Anachlysictis gracilis, it was characterized by its elongated fangs of the upper jaw, similar to the well known "sabertooth cats" (Machairodontinae), of which they were ecological equivalents. Despite being geologically younger than Anachlysictis, the morphology of Patagosmilus suggests that this species was more closely related to Thylacosmilus than Anachlysictis, though in other respects this species is less specialized than Thylacosmilus.
Hondadelphys is an extinct genus of carnivorous sparassodonts, known from the Middle Miocene of Colombia. The type species, H. fieldsi, was described in 1976 from the fossil locality of La Venta, which hosts fossils from the Villavieja Formation. Hondadelphys was originally interpreted as belonging to the opossum family Didelphidae, but subsequently assigned to its own family, Hondadelphidae and interpreted as a basal sparassodont. The genus name refers to the Honda Group, the stratigraphic group in which the fossils of this animal were first found, combined with delphys (Greek for "womb", a common suffix used for opossum-like metatherians).
Magericyon is an extinct genus of Amphicyonid ("bear-dog") that lived during the Miocene 10-9 Ma in what is now Spain.
Anatoliadelphys maasae is an extinct genus of predatory metatherian mammal from the Eocene of Anatolia. It was an arboreal, cat-sized animal, with powerful crushing jaws similar to those of the modern Tasmanian devil. Although most mammalian predators of the northern hemisphere in this time period were placentals, Europe was an archipelago, and the island landmass now forming Turkey might have been devoid of competing mammalian predators, though this may not matter since other carnivorous metatherians are also known from the Cenozoic in the Northern Hemisphere. Nonetheless, it stands as a reminder that mammalian faunas in the Paleogene of the Northern Hemisphere were more complex than previously thought, and metatherians did not immediately lose their hold as major predators after their success in the Cretaceous.
Kerberos ("Cerberus") is an extinct genus of hyainailourid hyaenodonts in the subfamily Hyainailourinae, that lived in Europe. It contains the single species Kerberos langebadreae.
Arminiheringia is an extinct genus of sparassodont. It lived during the Early Eocene in South America.
Proborhyaena is an extinct genus of proborhyaenid sparassodont that lived during the Oligocene of what is now South America. It is considered to be the largest of the sparassodonts.
