Catopsbaatar is a genus of multituberculate, an extinct order of rodent-like mammals. It lived in what is now Mongolia during the late Campanian age of the Late Cretaceous epoch, about 72 million years ago. The first fossils were collected in the early 1970s, and the animal was named as a new species of the genus Djadochtatherium in 1974, D. catopsaloides. The specific name refers to the animal's similarity to the genus Catopsalis. The species was moved to the genus Catopsalis in 1979, and received its own genus in 1994. Five skulls, one molar, and one skeleton with a skull are known; the last is the genus' most complete specimen. Catopsbaatar was a member of the family Djadochtatheriidae.
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives.
Didelphodon is a genus of stagodont metatherians from the Late Cretaceous of North America.
Deltatheridium is an extinct species of metatherian. It lived in what is now Mongolia during the Upper Cretaceous, circa 80 million years ago. A study in 2022 strongly suggested that Deltatherium was a marsupial, making it the earliest known member of this group.
Pucadelphys is an extinct genus of non-marsupial metatherian. The genus contains a single species, P. andinus. Fossils of Pucadelphys have been found in the Santa Lucía Formation in Tiupampa in Bolivia.
Sparassodonta is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has showed that sparassodonts died out long before eutherian carnivores arrived in South America. Sparassodonts have been referred to as borhyaenoids by some authors, but currently the term Borhyaenoidea refers to a restricted subgroup of sparassodonts comprising borhyaenids and their close relatives.
Thylacosmilus is an extinct genus of saber-toothed metatherian mammals that inhabited South America from the Late Miocene to Pliocene epochs. Though Thylacosmilus looks similar to the "saber-toothed cats", it was not a felid, like the well-known North American Smilodon, but a sparassodont, a group closely related to marsupials, and only superficially resembled other saber-toothed mammals due to convergent evolution. A 2005 study found that the bite forces of Thylacosmilus and Smilodon were low, which indicates the killing-techniques of saber-toothed animals differed from those of extant species. Remains of Thylacosmilus have been found primarily in Catamarca, Entre Ríos, and La Pampa Provinces in northern Argentina.
Alphadon is an extinct genus of small, primitive mammal that was a member of the metatherians, a group of mammals that includes modern-day marsupials. Its fossils were first discovered and named by George Gaylord Simpson in 1929.
Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, Gurbanodelta kara, is known from the late Paleocene (Gashatan) of China. Their fossils are restricted to Central Asia and North America. This order can be defined as all metatherians closer to Deltatheridium than to Marsupialia.
Asiatheriidae is a family of extinct metatherians in the order Asiadelphia known from Cretaceous Asia. Different from the Ameridelphia, they lacked a prominent distolateral process on the scaphoid bone and possessed a more slender fibula. Their dentition is different from most other marsupials.
Several mammals are known from the Mesozoic of Madagascar. The Bathonian Ambondro, known from a piece of jaw with three teeth, is the earliest known mammal with molars showing the modern, tribosphenic pattern that is characteristic of marsupial and placental mammals. Interpretations of its affinities have differed; one proposal places it in a group known as Australosphenida with other Mesozoic tribosphenic mammals from the southern continents (Gondwana) as well as the monotremes, while others favor closer affinities with northern (Laurasian) tribosphenic mammals or specifically with placentals. At least five species are known from the Maastrichtian, including a yet undescribed species known from a nearly complete skeleton that may represent a completely new group of mammals. The gondwanathere Lavanify, known from two teeth, is most closely related to other gondwanatheres found in India and Argentina. Two other teeth may represent another gondwanathere or a different kind of mammal. One molar fragment is one of the few known remains of a multituberculate mammal from Gondwana and another has been interpreted as either a marsupial or a placental.
Deltatheridiidae is an extinct family of basal carnivorous metatherians that lived during the Cretaceous and Paleogene. They were closely related to marsupials. Their fossils are restricted to Central Asia and North America. They mostly disappeared in the KT event, but a ghost lineage, currently represented by Gurbanodelta, survived until the late Paleocene by decreasing in size and becoming insectivorous.
Stagodontidae is an extinct family of carnivorous metatherian mammals that inhabited North America and Europe during the late Cretaceous, and possibly to the Eocene in South America.
Pappotherium is an extinct genus of mammals from the Albian of Texas, US, known from a fossilized maxilla fragment bearing two tribosphenic molars, discovered within the Glen Rose Formation near Decatur, Wise County, Texas.
Groeberiidae is a family of strange non-placental mammals from the Eocene and Oligocene epochs of Patagonia, Argentina and Chile, South America. Originally classified as paucituberculate marsupials, they were suggested to be late representatives of the allothere clade Gondwanatheria. However, the relationship of the type genus, Groeberia, to Gondwanatheria has been firmly rejected by other scholars.
The "Gurlin Tsav" skull is a currently unnamed carnivorous metatherian fossil from the Nemegt Formation of Mongolia. Composed of a single semi-complete skull, this specimen is notable in regards to the evolution and systematics of Metatheria as a whole, and thus nigh-omnipresent in phylogenetic analyses of this group.
Tsagandelta is a genus of deltatheroidean therian mammal that lived in Asia during the Late Cretaceous. Distantly related to modern marsupials, it is part of Deltatheroida, a lineage of carnivorous metatherians common in the Cretaceous of Asia and among the most successful non-theropod carnivores of the region. It represents the first known mammal from the Bayan Shireh Formation.
Peradectes is an extinct genus of small metatherian mammals known from the latest Cretaceous to Eocene of North and South America and Europe. The first discovered fossil of P. elegans, was one of 15 Peradectes specimens described in 1921 from the Mason pocket fossil beds in Colorado. The monophyly of the genus has been questioned.
Holoclemensia is an extinct genus of mammal of uncertain phylogenetic placement. It lived during the Early Cretaceous and its fossil remains were discovered in Texas.
Ambolestes is an extinct genus of eutherian mammal from the Early Cretaceous of China. It includes a single species, Ambolestes zhoui, known from a single complete skeleton recovered from the Yixian Formation, part of the fossiliferous Jehol biota. Ambolestes is one of the most basal eutherians, presenting a combination of features from both early eutherians (stem-placentals) and early metatherians (stem-marsupials). This is responsible for the generic name of Ambolestes: "ambo" is Latin for "both", while "-lestes" is a popular suffix for fossil mammals. The species name honors influential Jehol paleontologist Zhou Zhonghe.
