| Stagodontidae | |
|---|---|
 | |
| Didelphodon skull diagram | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Clade: | † Archimetatheria |
| Family: | † Stagodontidae Marsh, 1889 |
| Genera | |
† Didelphodon Contents | |
Stagodontidae is an extinct family of carnivorous metatherian mammals that inhabited North America and Europe [3] during the late Cretaceous, [1] and possibly to the Eocene in South America.
Currently, the family includes four genera, Eodelphis , Didelphodon , Fumodelphodon and Hoodootherium , which together include some seven different species. [1] [2] In addition, the Cenomanian species Pariadens kirklandi might be a member of the family. [2] Carneiro and Oliveira (2017) considered the species Eobrasilia coutoi from the early Eocene (Itaboraian) of Brazil to be a stagodontid; [4] if confirmed it would make it the only known Cenozoic and the only known South American member of the family. Stagodontids were some of the largest known Cretaceous mammals, ranging from 0.4 to 2.0 kilograms (0.88–4.41 lb) in mass. [5] One of the most unusual features of stagodontids are their robust, bulbous premolars, which are thought to have been used to crush freshwater mollusks, [6] a diet that apparently evolved independently at least twice within this clade. [7] Postcranial remains suggest that stagodontids may have been semi-aquatic. [8] [9] The most well described forms are found in Laramidia, but they are also present on Appalachian and South American sites, further leading credence to their aquatic habits. [10] Cretaceous fossils were also found in France, suggesting a pan-Laurasian distribution for Cretaceous metatherians. [3]
The evolution of Didelphodon and other large stagodontids (as well as large deltatheroideans like Nanocuris ) occurs after the local extinction of eutriconodont mammals, suggesting passive or direct ecological replacement. [11] They are considered rare in any given fauna they appear in, probably due to their specialised habits. [12]
Stagodontids were once thought to be closely related to the Sparassodonta, but later studies suggest they belong to a more ancient branch of the metatherian family tree, possibly closely related to pediomyids, [13] [14] being in particular closest to Pariadens , which forms the immediate outgroup to Stagodontidae. [15] [16] With the possible exception of Eobrasilia (see above), stagodontids are last known from the Maastrichtian, and are thought to have gone extinct in the K-T Extinction.
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives.
Didelphodon is a genus of stagodont metatherians from the Late Cretaceous of North America.
Eodelphis, from eo- plus [Di]delphis, thus meaning "very early opossum", is a genus of stagodont metatherians from the Late Cretaceous of North America, with distinctive crushing dentition. Named species include E. browni and the more advanced E. cutleri. Both come from the Late Campanian of Dinosaur Provincial Park, Alberta. Specimens are also known from the Judith River Formation of Montana. E. cutleri is related to the Maastrichtian genus Didelphodon as indicated by its enlarged premolars and more robust jaw. Eodelphis was probably an aquatic predator like its relative Didelphodon, and may have weighed about 0.6 kg (1.3 lb), making it one of the largest mammals of its time.
Pucadelphys is an extinct genus of non-marsupial metatherian. The genus contains a single species, P. andinus. Fossils of Pucadelphys have been found in the Santa Lucía Formation in Tiupampa in Bolivia.
Sparassodonta is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has showed that sparassodonts died out long before eutherian carnivores arrived in South America. Sparassodonts have been referred to as borhyaenoids by some authors, but currently the term Borhyaenoidea refers to a restricted subgroup of sparassodonts comprising borhyaenids and their close relatives.
Alphadon is an extinct genus of small, primitive mammal that was a member of the metatherians, a group of mammals that includes modern-day marsupials. Its fossils were first discovered and named by George Gaylord Simpson in 1929.
Borhyaenidae is an extinct metatherian family of low-slung, heavily built predatory mammals in the order Sparassodonta. Borhyaenids are not true marsupials, but members of a sister taxon, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful jaws, like those of the unrelated placentalians Hyaenodon and Andrewsarchus, for crushing bones. Borhyaenids grew up to an average of 5 to 6 feet long. Borhyaena is the type genus of this group.
Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, Gurbanodelta kara, is known from the late Paleocene (Gashatan) of China. Their fossils are restricted to Central Asia and North America. This order can be defined as all metatherians closer to Deltatheridium than to Marsupialia.
The Itaboraian age is a period within the Early Eocene geologic time epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age.
The Alcântara Formation is a geological formation in northeastern Brazil whose strata date back to the Cenomanian of the Late Cretaceous.
Proborhyaenidae is an extinct family of metatherian mammals of the order Sparassodonta, which lived in South America from the Eocene (Mustersan) until the Oligocene (Deseadan). Sometimes it has been included as a subfamily of their relatives, the borhyaenids. The largest species, Proborhyaena gigantea, is estimated to be about the size of a spectacled bear, with its skull reaching 60 cm (2.0 ft) in length, and body mass estimates up to approximately 90–200 kg (200–440 lb), making the proborhyaenids some of the largest known metatherians. Proborhyaenid remains have been found in western Bolivia, Uruguay, southern Brazil, and the provinces of Mendoza, Salta, and Chubut, in Argentina.
Deltatheridiidae is an extinct family of basal carnivorous metatherians that lived during the Cretaceous and Paleogene. They were closely related to marsupials. Their fossils are restricted to Central Asia and North America. They mostly disappeared in the KT event, but a ghost lineage, currently represented by Gurbanodelta, survived until the late Paleocene by decreasing in size and becoming insectivorous.
Oxlestes is an extinct mammal from the Late Cretaceous of Asia, more specifically from the Cenomanian of Uzbekistan. A carnivorous species of uncertain affinities, it is notable for its relatively large size, being among the largest of all Mesozoic mammals. Due to the limited amount of material, it has been considered a nomen dubium.
The "Gurlin Tsav" skull is a currently unnamed carnivorous metatherian fossil from the Nemegt Formation of Mongolia. Composed of a single semi-complete skull, this specimen is notable in regards to the evolution and systematics of Metatheria as a whole, and thus nigh-omnipresent in phylogenetic analyses of this group.
Anatoliadelphys maasae is an extinct genus of predatory metatherian mammal from the Eocene of Anatolia. It was an arboreal, cat-sized animal, with powerful crushing jaws similar to those of the modern Tasmanian devil. Although most mammalian predators of the northern hemisphere in this time period were placentals, Europe was an archipelago, and the island landmass now forming Turkey might have been devoid of competing mammalian predators, though this may not matter since other carnivorous metatherians are also known from the Cenozoic in the Northern Hemisphere. Nonetheless, it stands as a reminder that mammalian faunas in the Paleogene of the Northern Hemisphere were more complex than previously thought, and metatherians did not immediately lose their hold as major predators after their success in the Cretaceous.
Utaetus is an extinct genus of mammal in the order Cingulata, related to the modern armadillos. The genus contains two species, Utaetus buccatus and U. magnum. It lived in the Late Paleocene to Late Eocene and its fossil remains were found in Argentina and Brazil in South America.
The Itaboraí Formation is a highly fossiliferous geologic formation and Lagerstätte of the Itaboraí Basin in Rio de Janeiro, southeastern Brazil. The formation reaching a thickness of 100 metres (330 ft) is the defining unit for the Itaboraian South American land mammal age (SALMA), dating to the Early Eocene, approximately 53 to 50 Ma.
Ernestokokenia is an extinct genus of mammal, belonging to the Didolodontidae. It lived during the Early Eocene and the Middle Eocene, and its fossils were discovered in South America.
Brachytherium is an extinct genus of proterotheriid mammal from the Late Miocene to Late Pliocene of Argentina. It is represented by the type and only species B. cuspidatum, a taxon named in 1883 by Ameghino for a partial mandible with teeth. Though it was considered a dubious taxon at times, Brachytherium was revised as valid by Schmidt in 2015, who also synonymized the species Proterotherium gradatum and Lophogonodon paranensis, expanding the material known, all of which is from the Ituzaingó Formation. Some material previously referred to Brachytherium has been given the new name Neobrachytherium.
Albertatherium is an extinct genus of alphadontid metatherians that lived during the Late Cretaceous of North America. The genus contains two species, Albertatherium primus, and Albertatherium secundus. Fossils have been found in the Eagle Formation of Montana and the Milk River Formation of Alberta.
