Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a pouch. Living marsupials include kangaroos, koalas, opossums, Tasmanian devils, wombats, wallabies, and bandicoots among others, while many extinct species, such as the thylacine, Thylacoleo, and Diprotodon, are also known.
Opossums are members of the marsupial order Didelphimorphia endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North America in the Great American Interchange following the connection of North and South America.
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives.
Sparassodonta is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has showed that sparassodonts died out long before eutherian carnivores arrived in South America. Sparassodonts have been referred to as borhyaenoids by some authors, but currently the term Borhyaenoidea refers to a restricted subgroup of sparassodonts comprising borhyaenids and their close relatives.
The bushy-tailed opossum is an opossum from South America. It was first described by English zoologist Oldfield Thomas in 1912. It is a medium-sized opossum characterized by a large, oval, dark ears, fawn to cinnamon coat with a buff to gray underside, grayish limbs, and a furry tail. Little is known of the behavior of the bushy-tailed opossum; less than 25 specimens are known. It appears to be arboreal (tree-living), nocturnal and solitary. The diet probably comprises insects, eggs and plant material. This opossum has been captured from heavy, humid, tropical forests; it has been reported from Bolivia, Brazil, Colombia, Ecuador and Peru. The IUCN classifies it as least concern.
Herpetotheriidae is an extinct family of metatherians, closely related to marsupials. Species of this family are generally reconstructed as terrestrial, and are considered morphologically similar to modern opossums. Fossils of herpetotheriids come from North America, Asia, Europe, Africa, and perhaps South America. The oldest representative is Maastrichtidelphys from the latest Cretaceous (Maastrichtian) of the Netherlands and the youngest member is Amphiperatherium from the Middle Miocene of Europe. The group has been suggested to be paraphyletic, with an analysis of petrosal anatomy finding that North American Herpetotherium was more closely related to marsupials than the European Peratherium and Amphiperatherium.
Peratherium is a genus of metatherian mammals in the family Herpetotheriidae that lived in Europe and Africa from the Early Eocene to the Early Miocene. Species include the following:
Cryptonanus is a genus of opossums from South America. It includes five species found from Bolivia to Uruguay and eastern Brazil, one of which is now extinct. Although the first species were discovered in 1931, the genus was not recognized as distinct from Gracilinanus until 2005. It includes small opossums with generally grayish, sometimes reddish, fur that are mainly distinguished from other opossums by characters of the skull.
Stagodontidae is an extinct family of carnivorous metatherian mammals that inhabited North America and Europe during the late Cretaceous, and possibly to the Eocene in South America.
Siamoperadectes is a genus of non-marsupial metatherian from the Miocene of Thailand. A member of Peradectidae, it is the first member of its clade known from South Asia, and among the last non-marsupial metatherians.
Groeberiidae is a family of strange non-placental mammals from the Eocene and Oligocene epochs of Patagonia, Argentina and Chile, South America. Originally classified as paucituberculate marsupials, they were suggested to be late representatives of the allothere clade Gondwanatheria. However, the relationship of the type genus, Groeberia, to Gondwanatheria has been firmly rejected by other scholars.
Anatoliadelphys maasae is an extinct genus of predatory metatherian mammal from the Eocene of Anatolia. It was an arboreal, cat-sized animal, with powerful crushing jaws similar to those of the modern Tasmanian devil. Although most mammalian predators of the northern hemisphere in this time period were placentals, Europe was an archipelago, and the island landmass now forming Turkey might have been devoid of competing mammalian predators, though this may not matter since other carnivorous metatherians are also known from the Cenozoic in the Northern Hemisphere. Nonetheless, it stands as a reminder that mammalian faunas in the Paleogene of the Northern Hemisphere were more complex than previously thought, and metatherians did not immediately lose their hold as major predators after their success in the Cretaceous.
Peradectes is an extinct genus of small metatherian mammals known from the latest Cretaceous to Eocene of North and South America and Europe. The first discovered fossil of P. elegans, was one of 15 Peradectes specimens described in 1921 from the Mason pocket fossil beds in Colorado. The monophyly of the genus has been questioned.
Gypsonictops is an extinct genus of leptictidan mammals of the family Gypsonictopidae, which was described in 1927 by George Gaylord Simpson. Species in this genus were small mammals and the first representatives of the order Leptictida, that appeared during the Upper Cretaceous.
Ernestokokenia is an extinct genus of mammal, belonging to the Didolodontidae. It lived during the Early Eocene and the Middle Eocene, and its fossils were discovered in South America.
Holoclemensia is an extinct genus of mammal of uncertain phylogenetic placement. It lived during the Early Cretaceous and its fossil remains were discovered in Texas.
Asiatherium is an extinct genus of mammal, probably belonging to Metatheria. It lived during the Late Cretaceous, and its fossilized remains were discovered in Mongolia.
Amphiperatherium is an extinct genus of metatherian mammal, closely related to marsupials. It ranged from the Early Eocene to the Middle Miocene in Europe. It is the most recent metatherian known from the continent.
Arminiheringia is an extinct genus of sparassodont. It lived during the Early Eocene in South America.
Ambolestes is an extinct genus of eutherian mammal from the Early Cretaceous of China. It includes a single species, Ambolestes zhoui, known from a single complete skeleton recovered from the Yixian Formation, part of the fossiliferous Jehol biota. Ambolestes is one of the most basal eutherians, presenting a combination of features from both early eutherians (stem-placentals) and early metatherians (stem-marsupials). This is responsible for the generic name of Ambolestes: "ambo" is Latin for "both", while "-lestes" is a popular suffix for fossil mammals. The species name honors influential Jehol paleontologist Zhou Zhonghe.
