Tsagandelta Temporal range: Late Cretaceous, | |
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Preserved left dentary in lateral view, holotype PSS-MAE 629 | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Deltatheroida |
Family: | † Deltatheridiidae |
Genus: | † Tsagandelta Rougier et al. 2015 |
Species: | †T. dashzevegi |
Binomial name | |
†Tsagandelta dashzevegi Rougier et al. 2015 | |
Tsagandelta (meaning "white crest") is a genus of deltatheroidean therian mammal that lived in Asia during the Late Cretaceous. Distantly related to modern marsupials, it is part of Deltatheroida, a lineage of carnivorous metatherians common in the Cretaceous of Asia and among the most successful non-theropod carnivores of the region. It represents the first known mammal from the Bayan Shireh Formation.
The holotype was first discovered and collected in 2002 during the field expedition conducted by the Mongolian Academy of Sciences and the American Museum of Natural History, the specimen only preserved a partial left dentary. It was discovered on the outcrops at the Tsagan Tsonj locality in the Bayan Shireh Formation, [1] the age is estimated to be around 102 million to 86 million years ago, between the Cenomanian and Santonian stages of the Late Cretaceous epoch. [2] The generic name, Tsagandelta, is derived from the Mongolian цагаан (tsagaan, meaning white) and the Greek δέλτᾰ (delta, meaning crest or point-shaped) in reference to the withish sediments at the Tsagan Tsonj locality and the prominent dentition of the Deltatheroida. The specific name, dashzevegi, is in honor to the Mongolian paleontologist Demberlyin Dashzeveg who did notable research on early mammals. [1]
Tsagandelta is currently known from one specimen, the holotype PSS-MAE 629. This specimen is composed of a left dentary fragment containing an almost intact second molar, the base of the third molar and the roots of the first premolar; various other tooth sockets are empty, and the dental formula is probably similar to that of Deltatheridium . [1] Based on comparisons with the related Lotheridium , the preserved dentary indicates that Tsagandelta was slightly smaller than the former; Lotheridium is about the size of a modern marten. [3]
Tsagandelta was classified within the Deltatheridiidae, being more derived than Sulestes and Oklatheridium but less so than the rest of the family. This phylogenetic position has been argued to suggest an Asian origin for Deltatheroida. [1]
Like most deltatheroideans, Tsagandelta was a carnivore, its molar anatomy similar to the carnassials of other carnivorous therians. Tsagandelta and its relatives (as well as the unrelated eutriconodonts) are among the Mesozoic mammals most clearly specialised to meat-eating. [1] [3]
Tsagandelta hails from the Bayan Shireh Formation, dating from the Cenomanian to the Santonian. [1] [2] It is adjacent to the Iren Dabasu Formation, these formations bear a very similar fauna. [4] A great diversity of reptiles are known from the Bayan Shireh Formation, mainly compromising dinosaur and non-dinosaur taxa that likely preyed on Tsagandelta, mammals however, are pooly known from this formation. [5] [6] [7] [8]
Enigmosaurus is a genus of therizinosauroid that lived in Asia during the Late Cretaceous period. It was a medium-sized, ground-dwelling, bipedal herbivore that represents the third therizinosaur taxon from the Bayan Shireh Formation, although it is known from the lower part. The genus is monotypic, including only the type species E. mongoliensis, known from a well preserved pelvis and other tentative body remains.
Garudimimus is a genus of ornithomimosaur that lived in Asia during the Late Cretaceous. The genus is known from a single specimen found in 1981 by a Soviet-Mongolian paleontological expedition in the Bayan Shireh Formation and formally described in the same year by Rinchen Barsbold; the only species is Garudimimus brevipes. Several interpretations about the anatomical traits of Garudimimus were made in posterior examinations of the specimen, but most of them were criticized during its comprehensive redescription in 2005. Extensive undescribed ornithomimosaur remains at the type locality of Garudimimus may represent additional specimens of the genus.
Alectrosaurus is a genus of tyrannosauroid theropod dinosaur that lived in Asia during the Late Cretaceous period, about some 96 million years ago in what is now the Iren Dabasu Formation.
Achillobator is a genus of large dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous period about 96 million to 89 million years ago in what is now the Bayan Shireh Formation. The genus is currently monotypic, only including the type species A. giganticus. The first remains were found in 1989 during a Mongolian-Russian field expedition in Mongolia and later described in 1999. Remains at the type locality of Achillobator may represent additional specimens. It represents the first and largest dromaeosaurid known from the Bayan Shireh Formation.
Nomingia is a genus of oviraptorid theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia.
Bagaceratops is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 72 to 71 million years ago. Bagaceratops remains have been reported from the Barun Goyot Formation and Bayan Mandahu Formation. One specimen may argue the possible presence of Bagaceratops in the Djadochta Formation.
The Bayan Shireh Formation is a geological formation in Mongolia, that dates to the Cretaceous period. It was first described and established by Vasiliev et al. 1959.
Arstanosaurus is a genus of hadrosauroid dinosaur from the Santonian-Campanian-age Upper Cretaceous Bostobe Formation, Kazakhstan. It has had a confusing history, being considered both a hadrosaurid and a ceratopsid, or both at the same time (chimeric).
Tsagantegia is a genus of medium-sized ankylosaurid thyreophoran dinosaur that lived in Asia during the Late Cretaceous period. The genus is monotypic, including only the type species, T. longicranialis. The specimen consists of a very partial individual, comprising the skull and lacking postcranial remains. Since it only preserves the skull, Tsagantegia is mainly characterized by its elongated snout and the flattened facial osteoderms, greatly differing from other ankylosaurs.
Talarurus is a genus of ankylosaurid dinosaur that lived in Asia during the Late Cretaceous period, about 96 million to 89 million years ago. The first remains of Talarurus were discovered in 1948 and later described by the Russian paleontologist Evgeny Maleev with the type species T. plicatospineus. It is known from multiple yet sparse specimens, making it one of the most well known ankylosaurines, along with Pinacosaurus. Elements from the specimens consists of various bones from the body; five skulls have been discovered and assigned to the genus, although the first two were very fragmented.
Graciliceratops is a genus of neoceratopsian dinosaurs that lived in Asia during the Late Cretaceous period.
Erketu is a genus of somphospondylan sauropod dinosaur that lived in Asia during the Late Cretaceous roughly between 96 million and 89 million years ago. Its fossils were found in Mongolia between 2002 and 2003 during a field expedition and first described in 2006; later on in 2010 due to some cervicals that were left behind in the expedition. Erketu represents one of the first sauropods described from the Bayan Shireh Formation. The elongated cervical vertebrae of Erketu indicates that it was the sauropod with the longest neck relative to its body size.
Erlikosaurus is a genus of therizinosaurid that lived in Asia during the Late Cretaceous period. The fossils, a skull and some post-cranial fragments, were found in the Bayan Shireh Formation of Mongolia in 1972, dating to around 96 million and 89 million years ago. These remains were later described by Altangerel Perle and Rinchen Barsbold in 1980, naming the new genus and species Erlikosaurus andrewsi. It represents the second therizinosaur taxon from this formation with the most complete skull among members of this peculiar family of dinosaurs.
The Barun Goyot Formation is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia.
The Djadochta Formation is a highly fossiliferous geological formation situated in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million to 71 million years ago. The type locality is the Bayn Dzak locality, famously known as the Flaming Cliffs. Reptile and mammal remains are among the fossils that have been recovered from the formation.
The Iren Dabasu Formation is a Late Cretaceous geologic formation in the Iren Nor region of Inner Mongolia. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation. The formation was first described and defined by Henry Fairfield Osborn in 1922 and it is located in the Iren Nor region of China.
Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, Gurbanodelta kara, is known from the late Paleocene (Gashatan) of China. Their fossils are restricted to Central Asia and North America. This order can be defined as all metatherians closer to Deltatheridium than to Marsupialia.
Oxlestes is an extinct mammal from the Late Cretaceous of Asia, more specifically from the Cenomanian of Uzbekistan. A carnivorous species of uncertain affinities, it is notable for its relatively large size, being among the largest of all Mesozoic mammals. Due to the limited amount of material, it has been considered a nomen dubium.
Gobihadros is a genus of basal hadrosauroid dinosaur that lived in Asia during the Late Cretaceous period in what is now the Bayan Shireh Formation (Cenomanian-Santonian). It contains only the type species Gobihadros mongoliensis. It has an estimated length of 7.5 m (25 ft).
Natovenator is a genus of halszkaraptorine dromaeosaurid dinosaur from the Late Cretaceous Barun Goyot Formation of Mongolia. The genus is known from a single species, N. polydontus. Natovenator is crucial to the understanding of halszkaraptorines due to it providing more support for the semi-aquatic lifestyle that has been proposed for this clade. This discovery is important as the semi-aquatic lifestyles of halszkaraptorines was contested in early 2022.