This is a list of gomphothere fossils found in South America. Gomphotheres were elephant-like mammals that lived from the Middle Miocene (approximately 12 million years ago) to the Holocene (6000 years BP).
The following species have been described in twentieth and twenty-first century paleontological literature about South America. [2] An alternative proposal is considered within the scientific community, [3] [4] [5] [6] listed below.
Modern classification
Original classification
Formations are ordered according to their youngest age
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Pleistocene | Río Rujero, Tarija | 1,718 m (5,636 ft) | Cuvieronius hyodon | ||
Ulloma Fm. | Río Desaguadero, Ulloma, Oruro | 3,880 m (12,730 ft) | Cuvieronius hyodon | ||
Anzaldo, Cochabamba | 3,038 m (9,967 ft) | Cuvieronius sp. | |||
Betanzos Basin, between Potosí and Sucre | 3,390 m (11,120 ft) | Cuvieronius hyodon | |||
Valle de Carohuaycho, between Camiri and Charagua | 1,080 m (3,540 ft) | Cuvieronius "andinum" | |||
Chiquitos | 167 m (548 ft) | Cuvieronius "andinum" | |||
Villa Montes, Tarija | 413 m (1,355 ft) | Notiomastodon platensis | |||
Mid Pleistocene | Ñuapua Fm. | Ñuapua 1, Chuquisaca | 595 m (1,952 ft) | Stegomastodon sp. | |
Early Pleistocene | Tarija Fm. | Tarija, Tarija | 973 m (3,192 ft) | Cuvieronius hyodon, C. tropicus , Notiomastodon sp., Stegomastodon sp. | |
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Holocene | Lagoa do Rumo/Chapada Diamantina, Baixa Grande, Bahia | 400 m (1,300 ft) | Stegomastodon waringi | ||
Late Pleistocene | Garrincho, São Raimundo Nonato, Piauí | 390 m (1,280 ft) | Haplomastodon chimborazi | ||
Toca da Janela da Barra do Antoniao, São Raimundo Nonato, Piauí | 300 m (980 ft) | Haplomastodon sp. | |||
Gruta dos Brejões, Morro do Chapéu, Bahia | 500 m (1,600 ft) | Stegomastodon waringi | |||
Toca dos Ossos, Ourolândia, Bahia | 770 m (2,530 ft) | Stegomastodon waringi | |||
Fazenda Suse II, Vitória da Conquista, Bahia | 900 m (3,000 ft) | Stegomastodon waringi | |||
Janaúba, Minas Gerais | 580 m (1,900 ft) | Stegomastodon waringi | |||
Gruta do Ingá, Pains, Minas Gerais | 700 m (2,300 ft) | Stegomastodon waringi | |||
Poté, Minas Gerais | 560 m (1,840 ft) | Stegomastodon waringi | |||
Lapa do Caetano, Pedro Leopoldo, Minas Gerais | 685 m (2,247 ft) | Stegomastodon waringi | |||
Lajedo de Escada, Apodi, Rio Grande do Norte | 122 m (400 ft) | Stegomastodon waringi ("= Notiomastodon platensis") | |||
Caverna do Japonês & Nascente do Formoso, Serra da Bodoquena, Bonito, Mato Grosso do Sul | 373 m (1,224 ft) | Stegomastodon waringi | |||
Picos II, Piranhas, Alagoas | 255 m (837 ft) | Notiomastodon platensis | |||
Touro Passo Fm. | Touro Passo Creek, Uruguaiana, Rio Grande do Sul | 52 m (171 ft) | Gomphotheriidae indet. | ||
Pleistocene | Aguas do Araxá, Minas Gerais | 973 m (3,192 ft) | Haplomastodon waringi, Stegomastodon waringi | ||
Inajá Fm. | Inajá, Pernambuco | 355 m (1,165 ft) | Haplomastodon chimborazi | ||
Lage Grande, Pesqueira, Pernambuco | 740 m (2,430 ft) | Haplomastodon chimborazi, Notiomastodon vidali | |||
Roçado & Quixabinha, Petrolândia, Pernambuco | 320 m (1,050 ft) | Haplomastodon sp. | |||
Santa Cruz do Capibaribe & Panelas, Pernambuco | 440 m (1,440 ft) | Haplomastodon sp. (="Stegomastodon") | |||
Águas Belas & Santa Maria da Boa Vista, Pernambuco | 360 m (1,180 ft) | Gomphotheriidae indet. | |||
Lagoa das Pedras, Salgueiro, Pernambuco | 430 m (1,410 ft) | Haplomastodon waringi | |||
Lagoa Lajinha, Petrolina, Pernambuco | 390 m (1,280 ft) | Stegomastodon waringi "(=H. waringi)" | |||
Fazenda Nova, Brejo da Madre de Deus, Pernambuco | 470 m (1,540 ft) | Stegomastodon sp. "(=Haplomastodon sp.)", S. waringi , Notiomastodon platensis | |||
Jirau & João Cativo, Itapipoca, Ceará | 65 m (213 ft) | Notiomastodon platensis | |||
Taguara/Araras, Nova Mamoré & Porto Velho, Rondônia | 150 m (490 ft) | Notiomastodon platensis | |||
Sitio Novo, Matina, Bahia | 490 m (1,610 ft) | Notiomastodon platensis | |||
Vitória da Conquista, Bahia | 950 m (3,120 ft) | Notiomastodon platensis | |||
Fazenda Caraíba, Coronel João Sá, Bahia | 266 m (873 ft) | Stegomastodon waringi, Notiomastodon platensis | |||
Serro do Gronda, Espírito Santo | 25 m (82 ft) | Haplomastodon chimborazi | |||
Taperoá, Paraíba | 530 m (1,740 ft) | Haplomastodon sp. | |||
Upper Ribeira, São Paulo | 360 m (1,180 ft) | Stegomastodon waringi | |||
Tortonian | Solimões Fm. | Río Acre, Acre | 274 m (899 ft) | Amahuacatherium peruvium | |
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Pleistocene | Taguatagua, O'Higgins | 206 m (676 ft) | C. hyodon (rev. Stegomastodon sp.) | ||
Tierras Blancas & Los Vilos, La Ligua, Valparaíso | 943 m (3,094 ft) | C. hyodon (rev. Stegomastodon sp.) | |||
Parral, Cauquenes, Maule | 150 m (490 ft) | Cuvieronius hyodon | |||
San Pablo de Tramalhué, Pilauco Bajo, Osorno | 14 m (46 ft) | Cuvieronius hyodon | |||
Río Bueno, Valdivia | 10 m (33 ft) | Cuvieronius hyodon | |||
Paredones & Colchagua | 55 m (180 ft) | Cuvieronius hyodon | |||
Alto de Boroa, Temuco | 131 m (430 ft) | Cuvieronius hyodon | |||
Limahuida, Río Choapa | 282 m (925 ft) | Cuvieronius hyodon | |||
Casablanca | 313 m (1,027 ft) | Cuvieronius hyodon | |||
Chillán | 138 m (453 ft) | Cuvieronius hyodon | |||
Monte Verde | 59 m (194 ft) | C. hyodon (rev. Stegomastodon sp.) | |||
Lagunillas, Valparaíso | 212 m (696 ft) | cf. hyodon | |||
Catapilco, La Ligua | 103 m (338 ft) | cf. hyodon | |||
Marchigüe, Colchagua | 170 m (560 ft) | Stegomastodon sp. | |||
El Trebal 1, Padre Hurtado, Metropolitan Region | 424 m (1,391 ft) | Stegomastodon platensis | |||
Río Salado 1, Chiu Chiu | 3,012 m (9,882 ft) | Gomphotheriidae indet. | |||
Quebrada Quereo Fm. | Quereo | 20 m (66 ft) | Gomphotheriidae indet. | ||
Malloco, Santiago | 482 m (1,581 ft) | Gomphotheriidae indet. | |||
Algarrobo | 5 m (16 ft) | Gomphotheriidae indet. | |||
Melipilla | 269 m (883 ft) | Gomphotheriidae indet. | |||
Cerrillos, Santiago | 516 m (1,693 ft) | Gomphotheriidae indet. | |||
Chorro de la Vieja, Colchagua | 1,813 m (5,948 ft) | Gomphotheriidae indet. | |||
Cachagua | 5 m (16 ft) | Gomphotheriidae indet. | |||
Estero de Coyanco, Laja | 121 m (397 ft) | Gomphotheriidae indet. | |||
Pliocene | Las Pozas/Chacabuco | 695 m (2,280 ft) | C. tropicus (rev. Stegomastodon sp.) | ||
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Mid to Late Pleistocene | Tablazo Fm. | Río Cañas, Montecristi, Manabí | 100 m (330 ft) | Haplomastodon chimborazi | |
Pleistocene | Imbabura, Imbabura | 3,200 m (10,500 ft) | Cuvieronius hyodon (TL) | ||
Region de La Paz, Carchi | 2,690 m (8,830 ft) | Haplomastodon chimborazi | |||
Hacienca Pucará, Carchi | 3,000 m (9,800 ft) | Haplomastodon chimborazi | |||
Calderón/Carapungo, Quito, Pichincha | 2,630 m (8,630 ft) | Haplomastodon chimborazi | |||
Alangasí, Quito, Pichincha | 2,539 m (8,330 ft) | Haplomastodon chimborazi | |||
Punín, Chalán, Chimborazo | 2,994 m (9,823 ft) | Haplomastodon chimborazi | |||
Tumbaco, Pichincha | 2,360 m (7,740 ft) | Haplomastodon chimborazi | |||
Llano Chico, Quito, Pichincha | 2,610 m (8,560 ft) | Haplomastodon chimborazi | |||
Quebrada La Tola, Quito, Pichincha | 2,343 m (7,687 ft) | Haplomastodon chimborazi | |||
Checa, Pichincha | 2,520 m (8,270 ft) | Haplomastodon chimborazi | |||
La Carolina, Santa Elena | 5 m (16 ft) | Haplomastodon guayasensis | |||
Tanque Loma, Santa Elena | 21 m (69 ft) | Gomphotheriidae indet. (cf. N. platensis) | |||
Quebrada Pistud, Bolívar, Carchi | 2,760 m (9,060 ft) | Haplomastodon chimborazi | |||
Baños, Cuenca | 2,760 m (9,060 ft) | Cuvieronius hyodon | |||
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Pleistocene | General Bruguer/Riacho Negro, Presidente Hayes | 55 m (180 ft) | Stegomastodon sp. | ||
Puerto Santa Rosa, San Pedro | 72 m (236 ft) | cf. Notiomastodon | |||
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Late Pleistocene | La Brea, Talara, Piura | 3 m (9.8 ft) | Stegomastodon sp. | ||
Cupinisque Desert, La Libertad | 2 m (6.6 ft) | Stegomastodon sp. | |||
Ocucaje, Ica | 500 m (1,600 ft) | Stegomastodon sp. | |||
Quebrada El Jahuay, Arequipa | 1 m (3.3 ft) | Stegomastodon sp. | |||
La Huaca, Piura | 17 m (56 ft) | Stegomastodon sp. | |||
Ayusbamba, Cuzco | 3,500 m (11,500 ft) | Cuvieronius hyodon | |||
Quipan, Canta, Lima | 2,200 m (7,200 ft) | Stegomastodon waringi | |||
Pasamayo Quarry, Lima | 10 m (33 ft) | Stegomastodon waringi | |||
Pleistocene | Sacaco 2, Arequipa | 750 m (2,460 ft) | Stegomastodon sp. | ||
Celendín, Cajamarca | 2,500 m (8,200 ft) | Stegomastodon sp. | |||
Tortonian | Madre de Dios Fm. | Cerro Colorado, Madre de Dios | 300 m (980 ft) | Amahuacatherium peruvium | |
Age | Formation | Location | Present elevation | Fossils | Notes |
---|---|---|---|---|---|
Late Pleistocene | Dolores Fm. | Campo Viñoles, Cerro Largo | 105 m (344 ft) | Stegomastodon waringi | |
Mid to Late Pleistocene | Casil Quarry, Montevideo | 17 m (56 ft) | Stegomastodon waringi | ||
Some authors, Lucas, Mothé, Avilla et al., propose a reclassification of the South American gomphotheres as follows: [3] [4] [5] [6]
Other researchers as Labarca, Prado and Alberdi agree with the placement of species into Notiomastodon, but reject the idea of a single species. They continue to use Stegomastodon waringi for South American gomphotheres from Chile. [90]
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Proboscideans include some of the largest known land mammals. The largest land mammal of all time may have been a proboscidean; the elephant Palaeoloxodon namadicus has been estimated to be up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, surpassing the paraceratheres, the otherwise largest known land mammals, though this estimate was made based on a single fragmentary femur and is speculative. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.
Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.
Gomphotherium is an extinct genus of gomphothere proboscidean from the Neogene of Eurasia, Africa and North America. The genus is probably paraphyletic.
Cuvieronius is an extinct New World genus of gomphothere which ranged from southern North America to western South America during the Pleistocene epoch. Among the last gomphotheres, it became extinct at the end of the Pleistocene, approximately 12,000 years ago, following the arrival of humans to the Americas.
Stegomastodon is an extinct genus of gomphotheres, a family of proboscideans. It ranged throughout North America from the early Blancan ~4 Ma, to the early Irvingtonian. The South American species have been synonymized with Notiomastodon platensis.
Mixotoxodon is an extinct genus of notoungulate of the family Toxodontidae inhabiting South America, Central America and parts of southern North America during the Pleistocene epoch, from 1,800,000—12,000 years ago.
Rhynchotherium is an extinct genus of proboscidea endemic to North America and Central America during the Miocene through Pliocene from 13.650 to 3.6 Ma, living for approximately 10 million years.
The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Ma during the Paleocene and continuing through to the Late Pleistocene. These periods are referred to as ages, stages, or intervals and were established using geographic place names where fossil materials where obtained.
Notiomastodon is an extinct genus of gomphothere proboscidean, endemic to South America from the Pleistocene to the beginning of the Holocene. Notiomastodon specimens reached a size similar to that of the modern Asian elephant. Like other brevirostrine gomphotheres such as Cuvieronius and Stegomastodon, Notiomastodon had a shortened lower jaw and lacked lower tusks.
Choerolophodon is an extinct genus of proboscidean that lived during the Miocene of Eurasia and Africa. Fossils of Choerolophodon have been found in Africa, Southeast Europe, Turkey, Iraq, Iran, the Indian subcontinent, and China.
Surameryx is an extinct genus of herbivorous even-toed ungulates originally described as belonging to the extinct family Palaeomerycidae. A single species, S. acrensis, was described from the Late Miocene of the Madre de Dios Formation, South America. It was originally interpreted as one of the few northern mammals that entered South America before the Pliocene. However, both its identification as a member of the family Palaeomerycidae and claims about its Miocene age were subsequently challenged.
Sinomammut is a mammutid proboscidean from the Miocene of China. Only one species, S. tobieni, is known, named in 2016.
Tibitó is the second-oldest dated archaeological site on the Altiplano Cundiboyacense, Colombia. The rock shelter is located in the municipality Tocancipá, Cundinamarca, Colombia, in the northern part of the Bogotá savanna. At Tibitó, bone and stone tools and carbon have been found. Bones from Haplomastodon, Cuvieronius, Cerdocyon and white tailed deer from the deepest human trace containing layer of the site is carbon dated to be 11,740 ± 110 years old. The oldest dated sediments are lacustrine clays from an ancient Pleistocene lake.
The Sabana Formation is a geological formation of the Bogotá savanna, Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The formation consists mainly of shales with at the edges of the Bogotá savanna lignites and sandstones. The Sabana Formation dates to the Quaternary period; Middle to Late Pleistocene epoch, and has a maximum thickness of 320 metres (1,050 ft), varying greatly across the savanna. It is the uppermost formation of the lacustrine and fluvio-glacial sediments of paleolake Humboldt, that existed at the edge of the Eastern Hills until the latest Pleistocene.
The Sopas Formation is a Lujanian geologic formation in Uruguay.
The Libertad Formation is a Lujanian geologic formation in Uruguay.
The Dolores Formation is a Lujanian geologic formation in Uruguay.
Publications are in chronological order, most recent first, because of the advances in taxonomy and research