Ungulates are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving.
Mesonychia is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely when the last genus, Mongolestes, became extinct in the early Oligocene. In Asia, the record of their history suggests they grew gradually larger and more predatory over time, then shifted to scavenging and bone-crushing lifestyles before the group became extinct.
Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined during the Late Neogene, with only the large toxodontids persisting until the end of the Pleistocene. Collagen analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls, including rhinoceroses, tapirs and equines. but their relationships to other South American ungulates are uncertain. Several groups of notoungulates separately evolved ever growing teeth like rodents and lagomorphs, a distinction among ungulates only shared with Elasmotherium.
Phenacodus is an extinct genus of mammals from the late Paleocene through middle Eocene, about 55 million years ago. It is one of the earliest and most primitive of the ungulates, typifying the family Phenacodontidae and the order Perissodactyla.
Arctocyon is an extinct genus of ungulate mammals. Arctocyon was a "ground dwelling omnivore", that lived from 61.3-56.8 Ma. Synonyms of Arctocyon include Claenodon, and Neoclaenodon. Arctocyon was likely plantigrade, meaning that it walked with its feet flat on the ground, rather than on its toes.
Pantodonta is an extinct suborder of eutherian mammals. These herbivorous mammals were one of the first groups of large mammals to evolve after the end of the Cretaceous. The last pantodonts died out at the end of the Eocene.
Mesonychidae is an extinct family of small to large-sized omnivorous-carnivorous mammals. They were endemic to North America and Eurasia during the Early Paleocene to the Early Oligocene, and were the earliest group of large carnivorous mammals in Asia. They are not closely related to any living mammals. Mesonychid taxonomy has long been disputed and they have captured popular imagination as "wolves on hooves," animals that combine features of both ungulates and carnivores. Skulls and teeth have similar features to early whales, and the family was long thought to be the ancestors of cetaceans. Recent fossil discoveries have overturned this idea; the consensus is that whales are highly derived artiodactyls. Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define Mesonychia as basal to all ungulates, occupying a position between Perissodactyla and Ferae. In this case, the resemblances to early whales would be due to convergent evolution among ungulate-like herbivores that developed adaptations related to hunting or eating meat.
Arctocyonidae has been defined as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene . Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction.
Ankalagon saurognathus is an extinct carnivorous mammal of the family Mesonychidae, endemic to North America during the Paleocene epoch, existing for approximately 3.1 million years.
Carodnia is an extinct genus of South American ungulate known from the Early Eocene of Brazil, Argentina, and Peru. Carodnia is placed in the order Xenungulata together with Etayoa and Notoetayoa.
Tillodontia is an extinct suborder of eutherian mammals known from the Early Paleocene to Late Eocene of China, the Late Paleocene to Middle Eocene of North America where they display their maximum species diversity, the Middle Eocene of Pakistan, and the Early Eocene of Europe. Leaving no descendants, they are most closely related to the pantodonts, another extinct group. The tillodonts were medium- to large-sized animals that probably feed on roots and tubers in temperate to subtropical habitats.
Phenacodontidae is an extinct family of large herbivorous mammals traditionally placed in the “wastebasket taxon” Condylarthra, which may instead represent early-stage perissodactyls. They lived in the Paleocene and Eocene epochs and their fossil remains have been found in North America and Europe.
Triisodon is a genus of extinct mesonychian mammal that existed during the Early Paleocene of New Mexico, North America. The genus was named by Edward Drinker Cope in 1881 as a member of the Acreodi, a now invalid taxon that encompassed both creodonts and mesonychians. The premolar teeth have three points, hence the generic name. Cope described the type specimen of T. quivirensis as "about the size of a wolf." A smaller species has also been identified from the same region. Since material from this genus is incomplete, the exact size of adults and whether they showed sexual dimorphism or regional variations in size is unknown.
Ocepeia is an extinct genus of afrotherian mammal that lived in present-day Morocco during the middle Paleocene epoch, approximately 60 million years ago. First named and described in 2001, the type species is O. daouiensis from the Selandian stage of Morocco's Ouled Abdoun Basin. A second, larger species, O. grandis, is known from the Thanetian, a slightly younger stage in the same area. In life, the two species are estimated to have weighed about 3.5 kg (7.7 lb) and 10 kg (22 lb), respectively, and are believed to have been specialized leaf-eaters. The fossil skulls of Ocepeia are the oldest known afrotherian skulls, and the best-known of any Paleocene mammal in Africa.
Protictis is an extinct genus of carnivorous mammals known from the late Paleocene and early Eocene of North America. It belongs to the family Viverravidae of the Carnivoramorpha.
Gypsonictops is an extinct genus of leptictidan mammals of the monotypic family Gypsonictopidae, which was described in 1927 by George Gaylord Simpson. Species in this genus were small mammals and the first representatives of the order Leptictida, that appeared during the Upper Cretaceous.
Notiolofos is an extinct genus of sparnotheriodontid ungulate from the order Litopterna. The animal lived during the Eocene, in modern-day Antarctica. The genus contains two species, N. arquinotiensis, the type species, and N. regueroi.
Ernestokokenia is an extinct genus of mammal, belonging to the Didolodontidae. It lived during the Early Eocene and the Middle Eocene, and its fossils were discovered in South America.
Asmithwoodwardia is an extinct genus of mammals, from the order Litopterna. It lived during the Late Paleocene and the Early Eocene, and its fossilized remains were found in South America.
Protolipterna is an extinct genus of mammal, belonging to the order Litopterna. It lived during the Late Paleocene and the Early Eocene, in what is now South America.
