Anayatherium | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Notoungulata |
Family: | † Leontiniidae |
Genus: | † Anayatherium Shockey 2005 |
Type species | |
†Anayatherium ekecoa Shockey, 2005 | |
Species | |
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Anayatherium is an extinct genus of notoungulate belonging to the family Leontiniidae. It lived during the Late Oligocene, and its fossilized remains were discovered in South America.
It was a large-sized and heavy-shaped animal, that could reach the size of a cow. It may have reached a length of 2.5 meters ; its skull was 45 centimeters long, and its weight may have exceeded 350 kilograms.
Anayatherium was characterized by a very short muzzle, compared to other leontiniids. The shortening of the muzzle was reflected in the dentition by the loss of one of the upper teeth, probably a canine. The first upper incisor was larger than the second, as in Scarrittia , while the third and fourth premolars had vertical lingual grooves, as in Leontinia . The second upper incisor was of a typical shape, unlike Leontinia in which it had instead the shape of a canine ; in Anayatherium, the first incisor took the place of the canine. Unlike Scarrittia, the premolars had grooved protocones. The molars were low-crowned (brachydont), and only slightly high-crowned (hypsodont) on the lingual side.
The genus Anayatherium was first described in 2015 by Shockey, based on fossils found in the Salla Formation of Bolivia, in terrains dated from the Late Oligocene. Two species were attributed to the genus, A. ekecoa, the type species, and A. fortis, mainly distinguished by their respective sizes.
Anayatherium was a leontiniid, a family of notoungulates typical of the Oligocene and Miocene, with heavy and large bodies. Anayatherium seems to have belonged to a clade including Colpodon , Ancylocoelus , and the two derived genera Leontinia and Scarrittia.
Like all leontiniids, Anayatherium was a heavy quadrupedal folivore. Fossils of Anayatherium are unusually rare for a leontiniid ; leontiniids were usually common animals in their habitats. It is possible that the Salla Formation may not have been particularly suited environment for leontiniids during the Late Oligocene.
Toxodontia is a suborder of the meridiungulate order Notoungulata. Most of the members of the five included families, including the largest notoungulates, share several dental, auditory and tarsal specializations. The group is named after Toxodon, the first example of the group to be discovered by science.
Rhynchippus is an extinct genus of notoungulate mammals from the Late Oligocene of South America. The genus was first described by Florentino Ameghino in 1897 and the type species is R. equinus, with lectotype MACN A 52–31. Fossils of Rhynchippus have been found in the Agua de la Piedra and Sarmiento Formations of Argentina, the Salla and Petaca Formations of Bolivia, the Tremembé Formation of Brazil, and the Moquegua Formation of Peru.
Leontiniidae is an extinct family comprising eighteen genera of notoungulate mammals known from the Middle Eocene (Mustersan) to Late Miocene (Huayquerian) of South America.
Mesotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pleistocene of South America. Mesotheriids were small to medium-sized herbivorous mammals adapted for digging.
Ethegotherium is an extinct genus of Notoungulates, belonging to the suborder Typotheria. It lived from the Lower to the Middle Miocene, and its fossilized remains were discovered in South America. It might be a synonym of the genus Prohegetotherium.
Gualta is an extinct genus of leontiniid notoungulates. It lived during the Late Oligocene of what is now Argentina.
Eomorphippus is an extinct genus of notohippid notoungulate that lived from the Late Eocene to the Early Oligocene in what is today South America.
Eurygenium is an extinct genus of notoungulate belonging to the family Notohippidae. It lived during the Late Oligocene in what is today South America.
Ancylocoelus is an extinct genus of mammal, belonging to the order Notoungulata. It lived during the Late Oligocene, in what is today Argentina, in South America.
Colpodon is an extinct genus of herbivorous mammal, belonging to the order Notoungulata. It lived during the Early Miocene, in what is today Argentina and Chile, in South America.
Coquenia is an extinct genus of Notoungulate, belonging to the family Leontiniidae. It lived during the Middle Eocene, in what is today Argentina.
Elmerriggsia is an extinct genus of Notoungulate, belonging to the family Leontiniidae. It lived during the Late Oligocene in South America.
Martinmiguelia is an extinct genus of Notoungulate, belonging to the family Leontiniidae. It lived during the Middle Eocene, and its fossil remains were found in South America.
Taubatherium is an extinct genus of mammal, belonging to the order Notoungulata. It lived during the Late Oligocene, in what is today Brazil, in South America.
Termastherium is an extinct genus of leontiniid notoungulates that lived during the Early Oligocene of what is now Chile. Fossils of this genus have been found in the Oligocene-aged Abanico Formation of Chile.
Pascualihippus is an extinct genus of notoungulate belonging to the family Notohippidae. It lived during the Late Oligocene, in what is now Bolivia.
Hegetotheriopsis is an extinct genus of hegetotheriid notoungulate. It lived from the Late Oligocene to the Early Miocene, and its fossilized remains are found in Argentina.
Sallatherium is an extinct genus of Notoungulate, belonging to the suborder Typotheria. It lived during the Late Oligocene, and its fossilized remains were discovered in South America.
Rosendo is an extinct genus of notohippid notoungulates that lived during the Early Oligocene in what is now Argentina and Chile. Fossils of this genus have been found in the Sarmiento Formation and the Abanico Formations of Argentina and Chile.
Moqueguahippus is an extinct genus of notohippid notoungulates that lived during the Late Oligocene of what is now Peru. Fossils of this genus have been found in the Moquegua Formation of Peru, which it was named after.