Trionyx

Trionyx Temporal range: Miocene to recent, 23–0  Ma [1] PreꞒ Ꞓ O S D C P T J K Pg N
T. triunguis, the only extant species
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Trionychidae
Subfamily:	Trionychinae
Genus:	Trionyx Geoffroy Saint-Hilaire, 1809
Synonyms
Aspidonectes Wagler, 1830

Trionyx is a genus of softshell turtles belonging to the family Trionychidae. In the past many species in the family were classified in this genus, but today T. triunguis , the African or Nile softshell turtle, is the only extant softshell still classified as Trionyx. The other species still assigned to this genus are only known from fossils. T. triunguis is a relatively large, aquatic piscivore.

Species

The list of species follows a 2017 review of the Trionychidae by Georgios L. Georgalis and Walter G. Joyce. ^[2]

• Trionyx triunguis (Forsskål, 1775) – African or Nile softshell turtle
• † Trionyx pliocenicus Fucini, 1912 – Pliocene of Tuscany (Italy) ^[2]
• † Trionyx vindobonensis (Peters, 1855) – Miocene of Vienna (Austria) ^[2]

The following species are considered valid trionychid taxa, but with uncertain phylogenetic relationships. They are only referred to Trionyx provisionally due to its historic status as a wastebasket taxon. ^[2] The species from North America follow a 2015 review by Natasha S. Vitek and Walter G. Joyce ^[3] and the species from Europe and Asia follow the 2017 review by Georgalis and Joyce. ^[2]

• † "Trionyx" admirabilis(Hay, 1905) – Paleocene of New Mexico (United States) ^[3]
• † "Trionyx" aequa(Hay, 1908) – Eocene of Wyoming (United States) ^[3]
• † "Trionyx" allani(Gilmore, 1923) – Late Cretaceous of Alberta (Canada) ^[3]
• † "Trionyx" austerus(Hay, 1908) – Late Cretaceous of New Mexico (United States) ^[3]
• † "Trionyx" baynshirensisDanilov et al., 2014 – Middle Cretaceous of Dornogovi Aimag (Mongolia) ^[2]
• † "Trionyx" beecheri(Hay, 1904) – Late Cretaceous of Wyoming (United States) ^[3]
• † "Trionyx" boulengeriReinach, 1900 – Oligocene of Rhineland-Palatinate (Germany) ^[2]
• † "Trionyx" capelliniiNegri, 1892 – Eocene of Veneto (Italy) ^[2]
• † "Trionyx" dissolutusVitek & Danilov, 2014 – Late Cretaceous of Navoiy Region (Uzbekistan) ^[2]
• † "Trionyx" ellipticus(Hay, 1908) – Eocene of Utah (United States) ^[3]
• † "Trionyx" eloisae(Gilmore, 1919) – Paleocene of New Mexico (United States) ^[3]
• † "Trionyx" foveatusLeidy, 1857 – Late Cretaceous of Montana (United States) ^[3]
• † "Trionyx" gilbentuensisDanilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia) ^[2]
• † "Trionyx" gobiensisDanilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia) ^[2]
• † "Trionyx" gregarius(Gilmore, 1934) – Eocene of Inner Mongolia (China) ^[2]
• † "Trionyx" henriciOwen, 1849 – Eocene of Hampshire (United Kingdom) ^[2]
• † "Trionyx" ikoviensis Danilov et al., 2011 – Eocene of Luhansk (Ukraine) ^[2]
• † "Trionyx" impressus(Yeh, 1963) – Eocene of Yuangdong Province (China) ^[2]
• † "Trionyx" latus(Gilmore, 1919) – Late Cretaceous of Alberta (Canada) ^[3]
• † "Trionyx" leucopotamicusCope, 1891 – Eocene of Saskatchewan (Canada) ^[3]
• † "Trionyx" johnsoniGilmore, 1931 – Eocene of Inner Mongolia (China) ^[2]
• † "Trionyx" kansaiensis Vitek & Danilov, 2010 – Late Cretaceous of Khodzhent Province (Tajikistan) ^[2]
• † "Trionyx" linchuensis(Yeh, 1962) – Eocene of Shandong Province (China) ^[2]
• † "Trionyx" messelianusReinach, 1900 – Eocene of Hesse (Germany) ^[2]
• † "Trionyx" miensisOkazaki and Yoshida, 1977 – Pliocene of Mie Prefecture (Japan) ^[2]
• † "Trionyx" miocaenus(Matthew, 1924) – Eocene of Wyoming (United States) ^[3]
• † "Trionyx" minusculus(Chkhikvadze, 1973) – Eocene of East Kazakhstan Region (Kazakhstan) ^[2]
• † "Trionyx" mira(Hay, 1908) – Eocene of Wyoming (United States) ^[3]
• † "Trionyx" ninaeChkhikvadze, 1971 – Oligocene of Karagandy Region (Kazakhstan) ^[2]
• † "Trionyx" onomatoplokosGeorgalis & Joyce, 2017 – Late Cretaceous of Kyzylorda Region (Kazakhstan) ^[2]
• † "Trionyx" reesidei(Gilmore, 1919) – Paleocene of New Mexico (United States) ^[3]
• † "Trionyx" riabininiKuznetsov & Chkhikvadze, 1987 – Late Cretaceous of Kyzylorda Region (Kazakhstan) ^[2]
• † "Trionyx" robustus(Gilmore, 1919) – Late Cretaceous of New Mexico (United States) ^[3]
• † "Trionyx" shiluutulensisDanilov et al., 2014 – Late Cretaceous of Ömnögovi Aimag (Mongolia) ^[2]
• † "Trionyx" silvestrisWalker & Moody, 1974 – Eocene of Kent (United Kingdom) ^[2]
• † "Trionyx" singularis (Hay, 1907) – Paleocene of New Mexico (United States) ^[3]
• † "Trionyx" uintaensisLeidy, 1873 – Eocene of Wyoming (United States) ^[3]
• † "Trionyx" vegetus(Gilmore, 1919) – Paleocene of New Mexico (United States) ^[3]
• † "Trionyx" yixiensisLi, Tong et al., 2015 – Early Cretaceous of Heilongjiang Province (China) ^[2]

† = extinct

References

1. "Trionyx". FossilWorks . Retrieved 17 December 2021.
2. Georgalis, Georgios L.; Joyce, Walter G. (2017). "A Review of the Fossil Record of Old World Turtles of the Clade Pan-Trionychidae" (PDF). Bulletin of the Peabody Museum of Natural History. 58 (1): 115–208. doi:10.3374/014.058.0106. S2CID   89732695.
3. Vitek, Natasha S.; Joyce, Walter G. (2015). "A Review of the Fossil Record of New World Turtles of the Clade Pan-Trionychidae". Bulletin of the Peabody Museum of Natural History. 56 (2): 185–244. doi:10.3374/014.056.0204. ISSN   0079-032X. S2CID   86006102.

External links

• Biolib
• The Paleobiology Database