Podocnemididae is a family of pleurodire (side-necked) turtles, once widely distributed. Most of its 20 genera and 30 species are now extinct. Seven of its eight surviving species are native to South America: the genus Peltocephalus, with only one species ; and the genus Podocnemis, with six living species of South American side-necked river turtles. There is also one genus native to Madagascar: Erymnochelys, the Madagascan big-headed turtle, whose single species E. madagascariensis.
The Pleurodira are one of the two living suborders of turtles, the other being the Cryptodira. The division between these two suborders represents a very deep evolutionary divide between two very different types of turtles. The physical differences between them, although anatomical and largely internal, are nonetheless significant, and the zoogeographic implications of them are substantial. The Pleurodira are known more commonly as the side-necked turtles and the name Pleurodira quite literally translates to side neck, whereas the Cryptodira are known as hidden-necked turtles. The Pleurodira turtles are currently restricted to freshwater habitats in the Southern Hemisphere, largely to Australia, South America, and Africa. Within the Pleurodira, three living families are represented: Chelidae, also known as the Austro-South American side-necked turtles, the Pelomedusidae, also known as the African mud terrapins, and the Podocnemididae, also known as the American side-neck river turtles. However, they were cosmopolitan clade during the Cretaceous and most of the Cenozoic, and even occurred in marine environments around the world.
Paracryptodira is an extinct group of reptiles in the clade Testudinata, known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder sister to Cryptodira, they were then thought to be a very primitive lineage inside the Cryptodira according to the most common use of the latter taxon. They are now often regarded as late-diverging stem-turtles, lying outside the clade formed by Cryptodira and Pleurodira. The paracryptodires are said to have phylogenic relationships, noted as primary subclades, within the Baenidae and Pleurosternidae. Within each subclade, lies many biodiverse turtles that are continuously being investigated and added to the fossil record. Paracryptodires are divided into three main groups, Compsemydidae, known from the Late Jurassic to Paleocene of North America and Europe, Pleurosternidae, known from the Late Jurassic to Early Cretaceous of North America and Europe, and Baenidae, known from the Early Cretaceous to Eocene of North America. The latter two groups are more closely related to each other than to Compsemys, forming the clade Baenoidea.
Gilmoremys is an extinct genus of softshell turtle which lived during the late Cretaceous of North Dakota, Montana and Wyoming, United States.
Baenidae is an extinct family of paracryptodiran turtles known from the Early Cretaceous to Eocene of North America. While during the Early Cretaceous they are found across North America, during the Late Cretaceous they are only found in Laramidia, having disappeared from Appalachia. The majority of lineages survived the K-Pg Extinction, but the family was extinct by the latest Eocene. The name of the type genus, Baena, appears to be of Native American origin. They are primarily found in freshwater deposits, and are considered to be aquatic, with a largely generalist habit.
Sokatra is an extinct genus of pleurodiran turtle, known from the Cretaceous (Maastrichtian) Maevarano Formation of Madagascar. It belongs to the family Sahonachelyidae.
Araripemydidae is a family of freshwater aquatic turtles belonging to the order Pleurodira, known from the Early Cretaceous of South America and Africa. The family contains two recognised monotypic genera, Araripemys and Taquetochelys, from the Santana Group of Brazil and the Elrhaz Formation of Niger, respectively, which date to the Aptian-Albian stages of the Early Cretaceous. They are consider to be the most basal lineage within the Pelomedusoides. They are thought to have been specialised suction feeders. Laganemys was named in 2013 but was later determined to be synonymous with Taquetochelys.
Macrobaenidae is an extinct family of turtles, known from the Early Cretaceous to Paleogene of Laurasia. Their relationships to other turtles and whether or not they form a monophlyletic group are controversial. They are typically interpreted as stem or crown group cryptodires, but some more recent analyses have found them to lie outside crown group Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars.
Sichuanchelyidae is a family of extinct turtles in the clade Testudinata. It includes all perichelydians that are more closely related to Sichuanchelys than Meiolania, Helochelydra, or any extant turtles.
The Helochelydridae are an extinct family of stem-turtles known from fossils found in North America and Europe spanning the Early to Late Cretaceous.
Helochelydra is an extinct genus of extinct stem turtle known from the Early Cretaceous (Barremian) of the Isle of Wight, southern England.
Daspletosaurini is an extinct clade of tyrannosaurine dinosaurs that lived in Laramidia during the Late Cretaceous period. It consists of two genera: Daspletosaurus and Thanatotheristes. Four species have been described in the two genera, namely Daspletosaurus torosus, Daspletosaurus horneri, Daspletosaurus wilsoni, and Thanatotheristes degrootorum. At one point all of them were assigned as specimens of D. torosus, but several papers since 2017 have found them to represent distinct species. These studies found anagenesis in the group, whether contained in a daspletosaurin clade or paraphyletic in respect to the linage of tyrannosaurines leading up to Tyrannosaurus.
Perichelydia is a clade within Pantestudines known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and Spoochelys, known from the Mid-Cretaceous Griman Creek Formation of Australia. Kallokibotion from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia, Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually considered to be closer to the crown group than the other members of Perichelydia.
Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including Eurysternum, Plesiochelys and Thalassemys to the exclusion of Pelomedusa, Testudo and Protostega. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia.
Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus Angolachelys and Solnhofia, a definition that could potentially encompass a clade of entirely marine turtles. Angolachelonia was originally inclusive of only Solnhofia, Angolachelys and Sandownia when originally conceived by Octavio Mateus and colleagues in 2009, but later phylogenetic analyses by Serjocha Evers and Roger Benson in 2018 unites the family Sandownidae, including Angolachelys and Sandownia among other taxa, with the entirely Late Jurassic clade Thalassochelydia, where Solnhofia may be a basal member. While the placement of Solnhofia is weak and the clade that Angolachelonia represents may change with further analysis, the clade of Sandownidae and Thalassochelydia is well-supported, and does not collapse despite the uncertain evolutionary history of the group. Three alternative potential origins of Angolachelonia sensu Evers and Benson are shown below.
Jainemys is an extinct genus of bothremydid pleurodiran turtle that has been found in the Lameta Formation, India. Originally described by Sohan Lal Jain in 1977 under the name "Carteremys" pisdurensis, the species was transferred to the new genus Jainemys by Joyce & Bandyopadhyay in 2020.
Carteremys is an extinct genus of pelomedusid pleurodiran turtle from the Maastrichtian -Eocene, of India based on the type species C. leithi, which was named in 1953 by E. Williams and was originally placed in the genera Hydraspis by H. J. Carter in 1852 and Testudo, also by H. J. Carter, in 1871. A second species, C. pisdurensis, was named in 1977 by Sohan Lal Jain, but it was transferred to the separate genus Jainemys in 2020 by Joyce and Bandyopadhyay.
Sahonachelys is an extinct genus of pelomedusoid turtle from the Late Cretaceous (Maastrichtian) Maevarano Formation of Madagascar. The genus contains a single species, Sahonachelys mailakavava.
Caninemys is an extinct genus of large freshwater side-necked turtle, belonging to the family Podocnemididae. Its fossils have been found in Brazil and Colombia, in rocks dating back from the middle to late Miocene.
This list of fossil reptiles described in 2023 is a list of new taxa of fossil reptiles that were described during the year 2023, as well as other significant discoveries and events related to reptile paleontology that occurred in 2023.
